Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana
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alluaudi Antoine, 1957, Zabrus 572 almasyi Fassati, 1990, Bembidion 245 almeriensis Mateu, 1954, Acinopus 367
almonius Reitter, 1903, Trechus 326 almorae Andrewes, 1920, Chlaenius 353 almorensis Champion, 1923, Protopaussus 211 almosi Bokor, 1921, Duvalius [Trechus] 300 almum J. R. Sahlberg, 1900, Bembidion 247 almushensis Mandl, 1975, Carabus 194
alonsoi Cortés, 1988, Chlaenius 354 alotrus Lassalle & Remond, 1998, Carabus 154 alpestre Heer, 1838, Agonum [Anchomenus] 449 alpestris Heer, 1837, Agabus 38 alpestris A. Villa & G. B. Villa, 1833, Amara 563 alpestris Daffner, 1996, Anophthalmus 292 alpestris Sturm, 1815, Carabus 180 alpestris Beuthin, 1888, Cicindela 105 alpestris Heer, 1837, Cicindela 105 alpestris L. Redtenbacher, 1849, Harpalus 381 alpestris Falkenström, 1939, Hydroporus 60 alpestris Dejean, 1828, Molops [Feronia] 472 alpestris Gistel, 1857, Molops [HN] 472 alpestris Heer, 1837, Pterostichus [Argutor] 493 alpestris Heer, 1837, Trichotichnus [Harpalus] 396 alpherakii Semenov, 1898, Carabus 146 alphonsi J. Müller, 1914, Anophthalmus [Trechus] 290
alpicola Dejean, 1828, Amara 563 alpicola Heer, 1837, Amara 548 alpicola Jeannel, 1941, Bembidion [Peryphus] 269 alpicola Heer, 1837, Carabus 136 alpicola Fuss, 1860, Leistus 82 alpicola Motschulsky, 1866, Nebria [Alpaeus] 85
alpicola Ganglbauer, 1891, Reicheiodes [Dyschirius] 230 alpicola Sturm, 1825, Trechus 326
alpigradus Schauberger, 1927, Abax 470 alpigradus Sermet, 1985, Carabus 171 alpigradus Reitter, 1888, Trechus 326 alpina Paykull, 1790, Amara [Carabus] 558
alpinus Motschulsky, 1860, Agabus [Colymbetes] 35 alpinus Dejean, 1829, Aptinus 212
alpinus Dejean, 1828, Laemostenus [Pristonychus] 536 alpinus Paykull, 1798, Oreodytes [Dytiscus] 67
alpinus Curtis, 1827, Patrobus [NO] 22, 285 alpinus Meschnigg, 1934, Philorhizus [Dromius] 425 alpinus Baudi di Selve, 1871, Scotodipnus 240
alpinus Morita, 1997, Trichotichnus 395 alpivagus Tschitschérine, 1899, Harpalus 372
altaianus Jedlika, 1958, Pterostichus 505 altaica Jedlika, 1957, Amara [Zabrus] 565 altaica Eschscholtz, 1829, Cicindela 103 altaica Gebler, 1833, Cymindis 416
altaica Gebler, 1847, Nebria 94 altaica Jedlika, 1954, Rhopalostyla [Lebia] 430 altaicum Gebler, 1833, Bembidion 265 altaicus Motschulsky, 1844, Amara [Leirus] 559 altaicus Motschulsky, 1864, Brachinus 213 altaicus Gebler, 1847, Carabus 149 altaicus Jedlika, 1968, Harpalus 383 altaicus Gebler, 1848, Ilybius 43 altaicus Jedlika, 1968, Pseudotaphoxenus [Taphoxenus] 542 altaicus Germar, 1824, Pterostichus [Poecilus] 505 altaiensis Poppius, 1906, Pterostichus [Cryobius] 489
alter Semenov & Znojko, 1929, Martyr 443 alternans Motschulsky, 1845, Bembidion [Notaphus] 254 alternans Faldermann, 1836, Calathus 524 alternans Kraatz, 1887, Carabus 179 alternans Palliardi, 1825, Carabus 161 alternans Chaudoir, 1856, Chlaenius 348 alternans Rambur, 1837, Cymindis 411 alternans Gravenhorst, 1807, Hygrotus [Dytiscus] 72 alternans Dejean, 1828, Licinopsis [Pristonychus] 540 alternans Carret, 1897, Pterostichus [Omaseus] 499 alternans Bedel, 1905, Singilis 438 alternans Balkenohl, 1999, Trilophus 222
altestriatum Netolitzky, 1934, Bembidion 243 alticola Bruneau de Miré & Legros, 1963, Agabus 39 alticola A. Fiori, 1903, Bembidion 260 alticola Britton, 1948, Bradycellus 400 alticola Bellier, 1880, Carabus 171 alticola Andrewes, 1926, Cymindis 412 alticola Bates, 1878, Dicheirotrichus 402 alticola Laneyrie, 1952, Duvalius 304 alticola Uéno, 1989, Epaphiopsis 309 alticola Jeannel, 1941, Geotrechus 309 alticola Sharp, 1882, Hydroporus 63 alticola Uéno & Saito, 1991, Jujiroa 459 alticola Leoni, 1907, Laemostenus 533 alticola Wollaston, 1864, Masoreus 357 alticola Bates, 1891, Morphodactyla [Pristodactyla] 530 alticola Grouvelle, 1913, Omoglymmius [Rhysodes] 78 alticola Jeannel, 1928, Orotrechus 317 alticola Wollaston, 1864, Platyderus 521 alticola Antoine, 1956, Pogonus 287 alticola Löbl, 1995, Sphaerius [Microsporus] 26 alticola Iablokoff‑Khnzorian, 1971, Trechus [Altaiotrechus] 334 alticola Wollaston, 1854, Trechus 326 alticola J. Schmidt, 1998, Xestagonum 467 alticola Fairmaire, 1889, Xestopus [Pristonychus] 530 altimirai Lagar Mascaró, 1964, Chlaenius 353 altiphila Hieke, 1995, Amara 552 altissima Hieke, 1981, Amara 552 altitudinis Ledoux & Roux, 1996, Archastes 79 altitudinorum Breuning, 1943, Carabus [Megodontus] 164 altitudinum Breuning, 1934, Carabus 164 altitudinum Deuve & Mourzine, 1998, Cychrus 205 altomontensis Branes, 1987, Cychrus 205 altyntuanus Shilenkov, 1996, Carabus 172 alutacea Lesne, 1896, Clivina 219 alutaceus Gebler, 1829, Chlaenius 348 alutaceus Sturm, 1826, Cychrus 203 alutaceus Wollaston, 1857, Dromius 419 alutaceus Tschitschérine, 1893, Poecilus [Feronia] 481 alutensis Savulescu, 1972, Carabus 172 alveolata Bates, 1892, Mastax 217 alveolatus Semenov, 1898, Carabus 145 alveolatus Chaudoir, 1879, Oxylobus 231 alyshensis Deuve & Quéinnec, 1992, Trechus 337 alysidotus Illiger, 1798, Carabus 191 alzonai Deuve, 1990, Carabus 156 alzonai Deuve & Ledoux, 1989, Nebria 90 alzonai Binaghi, 1935, Pterostichus 511
amabilis Hampe, 1871, Amara 567 amabilis Reitter, 1919, Brachinus 212 amabilis L. Redtenbacher, 1867, Callistomimus 348 amabilis Dejean, 1831, Cylindera [Cicindela] 110 amabilis Chaudoir, 1852, Drypta 358 amabilis Ledoux, Roux & Sawada, 1991, Nebria 88 amabilis Schaufuss, 1863, Neotrechus [Anophthalmus] 315 amabilis Tschitschérine, 1893, Poecilus [Feronia] 481 amaenus Chaudoir, 1859, Metacolpodes [Colpodes] 460 amagisanum Morita, 1994, Negreum [Colpodes] 460 amagisanus Tanaka & Ishida, 1972, Pterostichus 498 amamiensis Satô, 1961, Hydroglyphus [Guignotus] 56 amamiooshimensis Habu, 1964, Parena 433 amamioshimae Habu, 1978, Synuchus 544 amamioshimense Habu, 1955, Habragonum [Agonum] 458
amani Breit, 1933, Tapinopterus 519 amanoi Nakane, 1968, Pterostichus 497 amarae Andrewes, 1920, Chlaenius 353 amarellus Bates, 1891, Harpalus 372 amariformis Motschulsky, 1844, Harpalus 372 amariformis Bates, 1892, Prakasha [Platymetopus] 393
amaroides Piochard de la Brûlerie, 1875, Amphimasoreus 521 amaroides Faldermann, 1836, Harpalus 383 amaroides Putzeys, 1877, Pristosia [Calathus] 530 amaroides Dejean, 1828, Pterostichus [Feronia] 489 amasiae Semenov, 1891, Laemostenus 537 amasicus Csiki, 1927, Carabus 185
amaurum Bates, 1883, Bembidion 243 amauryi Haury, 1898, Carabus 201
ambiellensis Jeannel, 1948, Hydraphaenops [Aphoenops] 311 ambigena Bates, 1878, Amara 557 ambigenus Reiche, 1853, Harpalus 373 ambigenus Bates, 1883, Pterostichus 510 ambigenus Desbrochers des Loges, 1906, Pterostichus [HN] 489 ambigua Glasunov, 1902, Nebria 90 ambigua Mäklin, 1857, Sericoda [Anchomenus] 466 ambiguum Dejean, 1831, Bembidion 252 ambiguum Fairmaire, 1891, Xestagonum [Colpodes] 467 ambiguus Dejean, 1829, Acinopus 367 ambiguus Bates, 1883, Apatrobus [Penetretus] 280 ambiguus Paykull, 1790, Calathus [Carabus] 526 ambiguus Lapouge, 1914, Carabus 157 ambiguus Csiki, 1931, Chlaenius 352 ambiguus Aubé, 1838, Hydroporus 65 ambiguus Dejean, 1825, Pheropsophus [Brachinus] 217 ambiguus Fairmaire, 1858, Pterostichus [Feronia] 485 ambiguus Rambur, 1838, Zabrus 569
amblygonellus Jeannel, 1964, Trechus 326 amblygonus Jeannel, 1926, Chaetoduvalius [Duvalius] 297
amblyops Bolívar y Pieltain, 1940, Pseudoplatyderus 524 amblypterus Chaudoir, 1868, Pterostichus [Feronia] 489 ambroggii Antoine, 1961, Laemostenus [Pristonychus] 533 ambrolauricus Belousov, 1989, Trechus 326 ambulans C. Zimmermann, 1832, Amara 555 ambulans Deuve, 1992, Cychrus 205 ambulator Andrewes, 1930, Xestagonum [Anchomenus] 467 amdoensis Hieke, 1996, Amara 552
amethystinum Semenov & Redikorzev, 1928, Calosoma 118 amethystinus Dejean, 1828, Laemostenus [Pristonychus] 536 amicorum Wrase, 1993, Eocarterus 365 amicorum Zaballos, 1998, Geocharis 239 amicorum Uéno, 1999, Kusumia 314 amicorum J. Schmidt, 1996, Platynus 463 amicorum Jaeger, 1997, Psychristus 404 amicorum P. Moravec & Wrase, 1998, Trechus 326 amicta Wollaston, 1864, Cymindis [Tarus] 417 amiculus Guignot, 1948, Canthydrus 34 amicus Kryzhanovskij, 1995, Andrewesius 456 amicus L. Dufour, 1843, Harpalus 373 amicus Reitter, 1900, Harpalus 380 ammophila Tschitschérine, 1898, Bleusea 369 ammophilus Dejean, 1829, Acinopus 369 amnicola J. R. Sahlberg, 1880, Agabus [Gaurodytes] 37 amnicola J. R. Sahlberg, 1900, Bembidion 243 amoena Faldermann, 1835, Diacheila [Blethisa] 206 amoena Dejean, 1829, Dioryche [Platymetopus] 369 amoenulum Péringuey, 1896, Bembidion 254 amoenulus Semenov, 1889, Agatus [Glycia] 408 amoenulus Jedlika, 1934, Metacolpodes [Colpodes] 460
amoenum R. F. Sahlberg, 1844, Bembidion 266 amoenus Solsky, 1874, Agabus 35
amoenus Wollaston, 1864, Paradromius [Dromius] 424 amoenus Dejean, 1828, Pterostichus [Feronia] 489
amorei Ganglbauer, 1907, Pterostichus 503 amphibolus J. Müller, 1922, Dyschirius 227 amphinomum Bates, 1883, Nipponagonum [Colpodes] 460 ampliata Bates, 1873, Amara [Bradytus] 553
ampliatus Zaitzev, 1927, Hydroporus 60 ampliceps J. R. Sahlberg, 1921, Hydroporus 65 amplicollis Mannerheim, 1853, Amara [Celia] 556 amplicollis A. Villa & G. B. Villa, 1844, Carabus 180 amplicollis Bleuse, 1914, Carabus 168 amplicollis Belousov, 1999, Caucasorites 297 amplicollis Ménétriés, 1848, Harpalus [NP] 372 amplicollis Toledo, 1994, Hydronebrius 41 amplicollis Fairmaire, 1880, Leistus 82 amplicollis Fairmaire & Laboulbène, 1854, Pterostichus [Feronia] 514 amplicollis Fairmaire, 1859, Trechus 326 amplicollis Uéno, 1955, Trephionus 546 amplior Deuve, 1994, Carabus 121 amplior Machado, 1992, Paradromius 424 amplipennis Baliani, 1943, Amara 553 amplipennis Lapouge, 1925, Carabus 179 amplipennis Bates, 1873, Dicheirotrichus 403 amplipennis J. Müller, 1911, Neotrechus [Trechus] 315
amplius Escalera, 1921, Calathus 526 amplum J. R. Sahlberg, 1908, Bembidion 20, 249 amplus Joseph, 1871, Anophthalmus 291 amplus J. Müller, 1916, Molops 472 amputatoides Mlyná, 1979, Harpalus 372 amrishi Makhan, 1999, Haliplus 32 amurense Motschulsky, 1859, Bembidion [Peryphus] 271 amurense Tschitschérine, 1895, Bembidion 250 amurensis Lutshnik, 1935, Amara 549 amurensis Géhin, 1885, Carabus 135 amurensis A. Morawitz, 1862, Cylindera [Cicindela] 110 amurensis Motschulsky, 1860, Demetrias 418 amurensis Fedorenko, 1991, Dyschirius 228 amurensis J. Balfour‑Browne, 1944, Dytiscus 52 amurensis Andrewes, 1924, Lebidia 426 amurensis Sharp, 1882, Nebrioporus [Deronectes] 65 amurensis Reitter, 1887, Philorhizus [Dromius] 426 amurensis Poppius, 1906, Pterostichus [Cryobius] 489 amurlandicus Shilenkov, 1994, Carabus 171 ana Uéno, 1955, Ja 458 anachoreta Ménétriés, 1832, Pterostichus [Feronia] 485 anadoluensis Kataev, 1993, Harpalus 381 anadyricus Csiki, 1930, Pterostichus 492 anae Morvan, 1982, Trechus 326 anagae Medina & Oromí, 1991, Orthomus [Wolltinerfia] 477
analis Aubé, 1838, Hydroporus 60 analis Jedlika, 1963, Pterostichus 514
anami Ledoux, 1977, Carabus 154 anamurensis Jedlika, 1965, Metadromius 421
anatolica Jedlika, 1965, Tachyura [Tachys] [DA] 278 anatolicum Jedlika, 1968, Bembidion 252 anatolicum Korge, 1964, Bembidion 267 anatolicus Schweiger, 1968, Badister 440 anatolicus Wewalka, 1972, Bidessus 54 anatolicus Jedlika, 1969, Calathus 528 anatolicus Chaudoir, 1857, Carabus [Procrustes] 186 anatolicus Jedlika, 1962, Carterus 363 anatolicus Motschulsky, 1865, Cychrus 202 anatolicus K. Daniel, 1904, Harpalus [Ophonus] 374 anatolicus Tschitschérine, 1898, Harpalus 372 anatolicus J. Balfour‑Browne, 1963, Hydroporus 60 anatolicus Guignot, 1957, Hyphydrus 74 anatolicus Casale, 1988, Laemostenus 533
anatolicus Mlyná, 1977, Molops 473 anatolicus Coiffait, 1956, Parvocaecus [Winklerites] 240 anatolicus Jedlika, 1968, Parvocaecus [Scotodipnus] [DA] [HN!] 240 anatolicus Jedlika, 1963, Platyderus 522 anatolicus J. Schmidt, 1996, Platynus 462 anatolicus Chaudoir, 1850, Poecilus [Feronia] 481 anatolicus Jedlika, 1963, Pterostichus 489
anatolicus Ganglbauer, 1915, Zabrus 572 anaulax Uéno, 1978, Yamautidius 345
ancestralis Belousov, 1998, Nannotrechus 314 anchocephalus Reitter, 1896, Carabus 147 anchomenoides Bates, 1873, Bradycellus [Tachycellus] 401 anchomenoides Bates, 1889, Chlaenius 353 anchoralis Chevrolat, 1845, Abroscelis [Cicindela] 99 anchoralis Sharp, 1884, Nebrioporus [Deronectes] 66
ancorifer Uéno, 1995, Trechiama 321 andalusia Rambur, 1837, Nebria 92 andalusiacus Rambur, 1837, Brachinus 212 andalusiacus Géhin, 1885, Carabus 161 andalusiacus Jedlika, 1950, Chlaenius 352 andalusiacus J. Vives & E. Vives, 1982, Laemostenus [Ceuthosthenes] 533 andalusiacus Mateu, 1957, Orthomus 477 andalusiacus Jeannel, 1927, Trechus 331
andalusicus Wehncke, 1872, Haliplus 31 andalusicus Reitter, 1897, Penetretus [Deltomerus] 285
andamanicus Régimbart, 1899, Copelatus 47 andarensis Bolívar y Pieltain, 1923, Nebria 85
andoi Uéno, 1959, Rakantrechus 319 andorranus Barthe, 1909, Carabus 168 andorranus Español & Comas, 1984, Hypotyphlus 239 andorranus Jeannel, 1937, Pterostichus [Haptoderus] [HN!] 489
andorranus Puel, 1924, Pterostichus 498 andorranus Jeanne, 1972, Trechus 339 andreae Tschitschérine, 1898, Amara 559
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