Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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thoracica Hoppe & Hornschuh, 1825, Lebia 428

thoracica W. Horn, 1892, Neocollyris [Collyris] 117

thoracica Kolenati, 1845, Tachyura [Tachys] 280

thoraciculus Cavazzuti, 1998, Carabus 127

thoracicum Fischer von Waldheim, 1829, Agonum [HN] 455



thoracicum Geoffroy, 1785, Agonum [Buprestis] 452

thoracicus Schilsky, 1889, Acupalpus [Stenolophus] 398

thoracicus Fischer von Waldheim, 1842, Calathus 526

thoracicus Germar, 1824, Carabus 182

thoracicus Duftschmid, 1812, Dyschirius [Scarites] [HN] 226

thoracicus P. Rossi, 1790, Dyschirius [Scarites] 227

thoracicus Motschulsky, 1844, Harpalus 380

thoracicus Stephens, 1828, Harpalus 375

thoracicus Machado, 1992, Lymnastis 274

thoracicus Gebler, 1843, Pseudotaphoxenus [Sphodrus] 542

thorectes Bates, 1873, Pterostichus 513

thorectes Uéno, 1983, Trechus 342

thorectoides Jedlika, 1958, Pterostichus 487



thoreii Clark, 1862, Porhydrus [Hydroporus] 68

thorencicus J. Ochs, 1967, Carabus [Megodontus] 165

thoreyi Dejean, 1828, Agonum 453

thorongiensis Hieke, 1990, Amara 564

thorungiensis J. Schmidt, 1994, Trechus 342

thracicus V. B. Guéorguiev, 1966, Hydroporus 65

thracicus J. Müller, 1932, Laemostenus 533

thracicus Pawowski, 1973, Trechus 337

thulensis J. R. Sahlberg, 1887, Pterostichus [Feronia] 491

thunbergi Gmelin, 1790, Anchomenus [Carabus] 455

thunbergi Quensel, 1806, Platymetopus [Carabus] 393

thunbergi A. Morawitz, 1862, Pterostichus 494



thuringianus Born, 1911, Carabus [HN] 196

thuringianus Schilsky, 1888, Carabus 181

thurtharus Ali & Jasim, 1989, Gyrinus 28

tianbaoshan Kleinfeld, 1998, Carabus 129

tianchi Perreau, 1992, Trechus 342

tianpingensis Deuve & Yu, 1992, Carabus 127

tianshanicus Mikhailov, 1976, Brachinus 213



tianshanicus Uéno & Pawowski, 1981, Oroblemites 316

tianshanicus Komarov & Kabak, 1995, Philorhizus 426



tianshanicus Semenov, 1908, Pseudotaphoxenus [Taphoxenus] 542

tianshanivagus Deuve, 1993, Trechus 342



tianshanskyi Lutshnik, 1934, Amara 562

tianshuicus Imura, 1995, Carabus 149

tiantai Kleinfeld, 1997, Carabus 143

tianzhuicus Imura, 1996, Carabus 156

tibetana Andrewes, 1934, Pristosia [Calathus] 531

tibetanophilus Deuve, 1991, Carabus 189

tibetanus Zaitzev, 1908, Agabus 37

tibetanus Grundmann, 1956, Chlaenius 352

tibetanus Semenov, 1904, Elaphrus 207

tibetanus Zaitzev, 1953, Hydroporus 65

tibeticus Andrewes, 1930, Harpalus 383



tibiale Dejean, 1828, Agonum 451

tibiale Duftschmid, 1812, Bembidion [Elaphrus] 246



tibialis Motschulsky, 1844, Amara [Leirus] 561

tibialis Paykull, 1798, Amara [Carabus] 551



tibialis Motschulsky, 1850, Brachinus 214

tibialis Bleuse, 1896, Carabus 168

tibialis Fischer von Waldheim, 1842, Carabus 199

tibialis Dejean, 1822, Cephalota [Cicindela] 103

tibialis Dejean, 1826, Chlaenius 350

tibialis Schaum, 1863, Craspedonotus 236

tibialis Reitter, 1887, Deltomerus 283

tibialis Andrewes, 1933, Dischissus 447

tibialis Audinet‑Serville, 1821, Harpalus 373

tibialis Motschulsky, 1860, Leistus 82

tibialis Bonelli, 1810, Nebria [Alpaeus] 87



tibialis G. R Waterhouse, 1833, Notiophilus 97

tibialis Westwood, 1842, Paussus 211



tibialis Motschulsky, 1844, Syntomus [Dromius] 433

tibiscinus Csiki, 1906, Carabus 197

tichomirovi Erjiomin, 1990, Pterostichus 492

tichonis Jakobson, 1907, Harpalus 383



tichonis Semenov, 1899, Pterostichus [Feronia] 485

tichyi Kult, 1946, Laemostenus [Pristonychus] 539



ticinense Paul Meyer, 1936, Bembidion 255

ticinensis Born, 1920, Carabus 166

ticinensis Stierlin, 1902, Molops [Tanythrix] 475

ticinensis Bänninger, 1951, Nebria 93



ticinus Born, 1920, Carabus 196

tiedemanni Breuning, 1935, Carabus 165



tieffenbachii Schaum, 1862, Tapinopterus [Pterostichus] 518

tieguanzi Imura, 1990, Carabus 129



tienmushaniense Kirschenhofer, 1984, Bembidion 263

tienmushanus Sciaky, 1997, Pterostichus 498

tientei J. Thomson, 1857, Carabus 127



tigancanus Born, 1908, Carabus 152

tigridis Andrewes, 1927, Anchomenus [Anchomenus] 455

tigridis Ali, 1967, Brachinus 216

tilesii Fischer von Waldheim, 1823, Pseudotaphoxenus [Sphodrus] 540

tilitshoensis J. Schmidt, 1994, Trechus 342

tillyi O. Berlov, 1997, Cychrus 19, 202

tim Kleinfeld, 1999, Carabus 156



timhaditensis Antoine, 1939, Cymindis 413

timida Motschulsky, 1844, Amara [Percosia] 565

timuri Kabak, 1994, Poecilus 485

tinctulus Bates, 1873, Harpalus 386

tinctum Zetterstedt, 1828, Bembidion 250



tinctus Clark, 1862, Hydroporus 64

tinctus Sharp, 1882, Hydrovatus 70

tinerfensis Mateu, 1961, Paradromius [Dromius] 424

tingens Reitter, 1900, Harpalus 376

tingitana Alluaud, 1932, Aepopsis 288



tingitana Putzeys, 1870, Amara [Leiocnemis] 554

tingitana Antoine, 1963, Cymindis 413

tingitanus Fairmaire, 1876, Harpalus 379

tingitanus Lucas, 1846, Pterostichus [Omaseus] 499

tingitanus Putzeys, 1870, Trechus 342



tippmanni Breuning, 1936, Carabus 151

tippmannianus Breuning, 1943, Carabus [Eucarabus] 151

tiragalloi Sanfilippo, 1978, Bidessus 55

tiresias Piochard de la Brûlerie, 1872, Aphoenops [Anophthalmus] 294



tiresias Piochard de la Brûlerie, 1873, Aphoenops [Anophthalmus] 294

tiridates Reitter, 1900, Harpalus 383

tiro Semenov, 1898, Carabus 189

tirolensis Schulz, 1908, Cicindela 104

tiruka Andrewes, 1924, Amara 557

tishetshkini Belousov & Kabak, 1994, Trechus 342

tisiphone Jeannel, 1917, Aphoenops 294

tismanae Winkler, 1936, Dicropterus [Scotodipnus] 239

tismanae Jeannel, 1928, Duvalius 300

tisselitensis Antoine, 1963, Cymindis 411

titan Zolotarev, 1913, Carabus [Tribax] 201

titanus Breuning, 1933, Carabus 197

titanus Solsky, 1874, Carenochyrus 363

titanus Sciaky, 1998, Stomis 518

tithonus Reitter, 1900, Harpalus 383

tiupicus Belousov & Kabak, 1994, Trechus 343

tizianoi Marggi & Huber, 1999, Bembidion 269


tjanschanicum Tschitschérine, 1895, Bembidion 269

tjanschanicus Semenov, 1889, Harpalus 383



tkatschukowi B. Bodemeyer, 1928, Carabus [Plectes] 132

tobaensis Lassalle, 1999, Acalathus 529



tockoensis Jedlika, 1936, Amara 549

todai Morita & Kanie, 1997, Pterostichus 498

toelgi Breit, 1933, Tapinopterus 519

toelgi Breit, 1926, Zabrus 572

tofaceus Uéno, 1970, Rakantrechus 319

togyusanus Imura, 1989, Carabus [Tomocarabus] 149

togyusanus Park & Kwon, 1996, Pterostichus 497

tohokuensis Ishikawa, 1984, Carabus 176

tokarae Habu, 1974, Agonum [Platynus] 454

tokaraensis Nakane, 1963, Copelatus 47

tokararum Lindroth, 1956, Synuchus 546



tokarensis Satô, 1972, Limbodessus [Uvarus] 57

tokatensis Roeschke, 1898, Carabus 194

tokatensis Chaudoir, 1863, Cicindela 106

tokatensis Vigna Taglianti, 1976, Parvocaecus [Winklerites] 240

tokejii Yoshida & Tanaka, 1960, Pterostichus 502

toksanbaicus Belousov & Kabak, 1993, Trechus 342

tokui Satô, 1985, Hydroporus 65

tokui Uéno, 1989, Trechiama 324



tokunagai Kamiya, 1932, Liodessus [Bidessus] 57

tokunoshimanus Nakane, 1956, Cillenus [Bembidion] 272

tokunoshimanus Satô, 1962, Dineutus 26

tokyoensis Nakane, 1953, Dyschirius 229

tokyoensis Habu, 1956, Oodes 445

tolbonuri Müller‑Motzfeld, 1984, Bembidion 256

toledana Schulz, 1909, Cylindera [Cicindela] 110

toledanoi Farka, 1999, Leistus 81



toletanum Perris, 1864, Bembidion 264

tolli Poppius, 1910, Carabus 172

tolli Zaitzev, 1907, Colymbetes 45

tolli Kataev & Shilenkov, 1996, Dicheirotrichus 403

tolminensis J. Müller, 1922, Anophthalmus [Trechus] 293

tombeanus Ganglbauer, 1904, Duvalius [Trechus] 301

tombesii F. Battoni, 1976, Calathus 528

tomensis Gebler, 1847, Pterostichus [Omaseus] 506

tomentosus Motschulsky, 1845, Acilius 49

tomentosus Vigors, 1825, Craspedophorus [Panagaeus] 446

tomentosus Poppius, 1905, Hydroporus 63

tomentosus Dejean, 1831, Oedesis [Ditomus] 366

tomokunii Satô, 1985, Copelatus 48

tomokunii Satô, 1981, Microdytes 75



tomoricensis J. Müller, 1923, Amara 566

tomoricensis J. Müller, 1923, Molops 473



tomoricensis Horvatovich, 1974, Nebria 84

tomoricensis J. Müller, 1935, Tapinopterus 520

tomskoensis Jedlika, 1960, Pseudotaphoxenus [Taphoxenus] 542

tongchuanensis Imura, 1994, Carabus 198

tongdosanus Kwon & Lee, 1984, Carabus 183

tongensis Breuning, 1950, Carabus 146

tonggulensis Zamotajlov & Sciaky, 1996, Diplous 284

tonkinensis Wasmann, 1921, Ceratoderus 209

tonkinensis Janssens, 1948, Platyrhopalus 211

tonkinensis Balkenohl, 1999, Trilophus 222

tonklii Drovenik, 1989, Aphaenopidius 293

tonklii Kirschenhofer, 1992, Trichotichnus 397

topaz Belousov & Kabak, 1998, Trechus 342

torgos Morvan, 1998, Xestagonum 469

toribioi Ortuño, 1988, Typhlocharis 241

torigaii Uéno, 1973, Kurasawatrechus 313

toropovi Obydov, 1997, Carabus 145

torosa Ledoux, Roux & Sciaky, 1994, Nebria 91

torosensis Jedlika, 1961, Harpalus 383

torosiense Jedlika, 1961, Bembidion 272

torosum Marggi & Huber, 1999, Bembidion 252

torosus I. Frivaldszky von Frivald, 1835, Carabus 158

toroticus Belousov & Kabak, 1996, Trechus 342

torpidus Cavazzuti, 1997, Carabus 190

torquatus Cavazzuti, 1995, Carabus 127

torquatus Fischer von Waldheim, 1829, Cybister [Dytiscus] 51

torrentialis Apfelbeck, 1908, Trechus 342

torrentialis Apfelbeck, 1912, Trechus 342

torressalai Coiffait, 1956, Laemostenus [Pristonychus] 539

torressalai Coiffait, 1958, Microtyphlus 240

torressalai Jeanne, 1996, Platyderus 523

torretassoi Schatzmayr, 1928, Cychrus 202

torretassoi Schatzmayr & Koch, 1934, Helenaea 273

torretassoi Schatzmayr, 1937, Licinus 442

torretassoi Schatzmayr, 1932, Ophonus [Harpalus] 390

torretassoi J. Müller, 1928, Orotrechus 317

torretassoi Schatzmayr & Koch, 1934, Tachys [DA] 277

torretassoi Jeannel, 1937, Trechus 342

torrida Panzer, 1796, Amara [Carabus] 561

torridiformis Poppius, 1906, Harpalus 381

torridoides Reitter, 1900, Harpalus 383



torridus Motschulsky, 1844, Harpalus 381

torta W. S. MacLeay, 1825, Dioryche [Harpalus] 369



torulosus Fischer von Waldheim, 1823, Cychrus 203

torva Lutshnik, 1933, Amara 556

tosana Oike, 1936, Cicindela 106

tosanus Nakane, Iga & Uéno, 1953, Carabus [Apotomopterus] 177

tosanus Kasahara & Y. Itô, 1997, Pterostichus 513

toshioi Habu, 1981, Ja 458

tosioi Uéno, 1972, Trechus 325

tosta Andrewes, 1925, Tachyura [Tachys] 280

totomarginalis DeGeer, 1774, Dytiscus 52

toulgoeti Deuve, 1989, Carabus 127



tournieri Putzeys, 1866, Dyschirius 229

tournieri Gautier des Cottes, 1860, Molops 474

tourteani Sirguey, 1931, Carabus 139

touzalini Lapouge, 1924, Calosoma [Callipara] 119

touzalini Lapouge, 1911, Carabus [Coptolabrus] 183

touzalini Andrewes, 1920, Chlaenius 349



touzalini Andrewes, 1923, Pterostichus 502

towadensis Habu, 1955, Synuchus [Calathus] 545

toxanbaicus Kabak, 1990, Poecilus 481

toyodai Morita & Kurosa, 1998, Pterostichus 498

toyohamaensis Shimizu, 1996, Carabus [Damaster] 122

trabecula Neboiss, 1984, Tenomerga 25

trabuccarius Fairmaire, 1858, Carabus 168

trabzonicum Belousov & Sokolov, 1994, Bembidion 246

trachydermon Andrewes, 1936, Scarites 234

trachynodes Bates, 1891, Carabus 176

trachypachys Sciaky & Pavesi, 1995, Trechus 342

trachys Andrewes, 1923, Chlaenius 349

traessarti Feng, 1936, Hydroglyphus [Bidessus] 56

tragbaliensis Hieke, 1990, Amara 566

trajectum Netolitzky, 1939, Bembidion 272

tranquebarica Bonelli, 1813, Clivina 220

tranquillus Habu, 1954, Trichotichnus 397

transalpinus Schauberger, 1922, Bradycellus 400

transalpinus Breit, 1914, Platyderus 523



transalpinus Heer, 1837, Poecilus [Pterostichus] 483

transbaicalica Motschulsky, 1844, Cicindela 107



transbaicalicum Motschulsky, 1844, Bembidion [Peryphus] 243

transbaicalicus Motschulsky, 1844, Pterostichus [Molops] 505

transbaikalica Jedlika, 1963, Cymindis 417

transbaikalicus A. Fleischer, 1914, Paradromius [Dromius] 424

transbenacanus Ganglbauer, 1908, Molops 475

transcarpathicus Shilenkov & Rizun, 1989, Duvalius 300



transcaspia Brancsik, 1899, Amara [Percosia] 566

transcaspica Semenov, 1895, Cephalota [Cicindela] 102

transcaspica Semenov, 1891, Cymindis 415

transcaspica Semenov, 1891, Diplocheila [Rembus] 439



transcaspica Glasunov, 1902, Nebria 91

transcaspica Semenov, 1889, Stenolepta 543

transcaspicum Semenov, 1889, Asaphidion [Tachypus] 243

transcaspicum Semenov, 1891, Cymbionotum [Coscinia] 235

transcaspicus Mateu, 1991, Apristus 431

transcaspicus Tschitschérine, 1899, Dicheirotrichus [Trichocellus] 403



transcaspicus Semenov, 1907, Dyschirius 225

transcaspicus Tschitschérine, 1898, Harpalus 370

transcaspicus Glasunov, 1909, Poecilus [Platysma] 480

transcaucasica Lutshnik, 1935, Amara 557

transcaucasica Putzeys, 1866, Clivina 219

transcaucasicum Lutshnik, 1938, Bembidion 246



transcaucasicum Chaudoir, 1876, Cymbionotum [Coscinia] 235

transcaucasicus Chaudoir, 1876, Abacetus 347



transcaucasicus Zaitzev, 1927, Agabus 37

transcaucasicus Csiki, 1927, Carabus 199

transcaucasicus Casale, 1988, Laemostenus 538

transcaucasiens Hieke, 1970, Amara 567

transcurrens Csiki, 1927, Carabus 165

transdanubialis Kenyery, 1983, Carabus 179

transdanubialis Csiki, 1946, Cicindela 105

transdanubiensis Nonveiller, Pavievi & Popovi, 1994, Trechus 328



transeuns Lapouge, 1925, Carabus 130

transfuga Semenov, 1898, Carabus 165

transfuga Chaudoir, 1871, Pogonus 288

transfuga Schaum, 1864, Zabrus 572

transfugus Apfelbeck, 1907, Carabus 173

transgrediens Gschwendtner, 1923, Hydroporus 65



transiens K. Daniel, 1902, Bembidion 262

transiens Mateu, 1957, Orthomus 477

transiliensis Kryzhanovskij, 1974, Amara [Curtonotus] 561

transiliensis Semenov, 1897, Carabus 192

transiliensis Glasunov, 1909, Poecilus [Platysma] 480

transilvanicus A. Fleischer, 1905, Dyschirius 225

translucida Ballion, 1871, Cymindis 411

transmontanus Semenov, 1908, Taphoxenus 544



transmutans Bates, 1886, Stenolophus 405

transparens Gebler, 1829, Bembidion [Peryphus] 248



transparens Motschulsky, 1844, Harpalus 372

transparens Motschulsky, 1850, Tachys 275

transpyrenaeus Breit, 1933, Poecilus [Pterostichus] 484

transsylvanica Csiki, 1902, Duvaliopsis [Anophthalmus] 298

transsylvanica Germar, 1824, Nebria 87



transsylvanicum Bielz, 1852, Bembidion 256

transsylvanicum K. Daniel & J. Daniel, 1898, Parazuphium [Zuphium] 573

transsylvanicus Dejean, 1826, Carabus 181



transsylvanicus Chaudoir, 1861, Pterostichus [Feronia] 495

transsylvanicus Csiki, 1928, Tachys 276

transsylvanicus Mandl, 1936, Tapinopterus 520



transversa Schmidt‑Göbel, 1846, Coptodera 435

transversa Morvan, 1972, Cymindis 414

transversalis Schaum, 1862, Anthracus [Stenolophus] 400

transversalis Csiki, 1927, Carabus 186

transversalis Dejean, 1822, Cicindela 108

transversalis Gautier des Cottes, 1861, Haliplus 33

transversalis Pontoppidan, 1763, Hydaticus [Dytiscus] 54

transversalis Duftschmid, 1812, Pterostichus [Carabus] 488

transversalis Motschulsky, 1844, Trachypachus 33

transverserugosus Mandl, 1955, Carabus 145

transversestriatus DeGeer, 1774, Colymbetes [Dytiscus] 45

transversicollis Baliani, 1943, Amara 557

transversicollis Putzeys, 1866, Amara [Curtonotus] 559

transversithorax Reitter, 1923, Cymindis 416

transversum J. Müller, 1918, Bembidion 249



transversus Eidam, 1941, Carabus 196

transversus C. G. Thomson, 1870, Haliplus 31

transversus Motschulsky, 1865, Microschemus [Epicosmus] 447

transversus Motschulsky, 1844, Ophonus 391



transversus Chaudoir, 1868, Pterostichus [Feronia] 504

transvolgensis Semenov, 1895, Cymindis 415

transvolgensis Semenov, 1904, Haliplus 32

transylvanicus Jeannel, 1921, Trechus 339

tranteevi V. B. Guéorguiev, 1964, Pheggomisetes 318



trapesunticus Eichler, 1924, Carabus 158

trapeti Bleuse, 1885, Carabus 168

trapezicollis Apfelbeck, 1904, Badister 440

trapezicollis Chaudoir, 1846, Cicindela 105

trapezicollis Chaudoir, 1859, Orthomus [Feronia] 476

trapezicollis Bates, 1889, Syleter [Clivina] 221

tratnikovi Lutshnik, 1928, Amara 551

travnikanus Apfelbeck, 1904, Cychrus 205



trawasangboi Morvan, 1982, Carabus 162

trebicianus J. Müller, 1914, Anophthalmus [Trechus] 292

trebinjense Apfelbeck, 1899, Bembidion 270

trebinjensis Apfelbeck, 1904, Neotrechus [Trechus] 316



trechiformis Jordan, 1894, Tachys 275

trechioides Motschulsky, 1862, Anophthalmus 292

trechoides Reitter, 1895, Ocys 273



trechoides Hope, 1845, Stenolophus [Harpalus] 405

trechomorphum Uéno, 1971, Caecidium 272



tremebunda W. S. MacLeay, 1825, Cylindera [Cicindela] 111

tremebundum Lutshnik, 1933, Poecilus [Platysma] 479

tremula Audouin & Brullé, 1839, Cylindera [Cicindela] 110

trentinus Kraatz, 1877, Carabus 196

trentinus Ganglbauer, 1903, Laemostenus 532

trentinus Csiki, 1932, Ophonus [Harpalus] 390

trepidum LeConte, 1848, Bembidion [Ochthedromus] 248

trescavicensis Ganglbauer, 1891, Duvalius [Trechus] 300

tresnjevikensis Horvatovich, 1974, Nebria 85


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