Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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rivosa L. Miller, 1868, Nebria 86

rivulare Dejean, 1831, Bembidion 248

rivularis Motschulsky, 1844, Cymindis 411



rivularis Motschulsky, 1860, Harpalus 376

rivularis Gyllenhal, 1810, Trechus [Bembidium] 325

rizeanus Imura & Su, 1998, Carabus 133

rizensis Ledoux, 1975, Carabus 193

rjabuchini Lafer, 1989, Synuchus 546



roberti Coiffait & A. Gaudin, 1950, Aphoenops 294

roberti Deuve & Mourzine, 1998, Carabus 189

roberti Fery & Hosseinie, 1998, Deronectes 59

roberti Abeille de Perrin, 1903, Duvalius [Trechus] 305

robertorum Fery, 1999, Hydroporus 63

robichei Auvray, 1994, Carabus 194

robinii Laboulbène, 1849, Aepopsis [Trechus] 288

roboretanus J. Müller, 1933, Orotrechus 317

roborovskii Mikhailov, 1988, Bembidion 258

roborovskii Zaitzev, 1953, Hygrotus [Coelambus] 72

roborowskii Semenov, 1887, Carabus 191

roborowskii Tschitschérine, 1899, Dicheirotrichus [Trichocellus] 403

roborowskii Semenov, 1889, Nebria 88

robusta Baliani, 1932, Amara 551

robusta Semenov, 1891, Bronislavia 363



robusta Mateu, 1952, Tachyura 278

robustapicalis Belousov & Kabak, 1998, Trechus 339

robustior Breuning, 1966, Carabus 126

robustior Uéno, 1997, Ishikawatrechus 312

robustoelegans Deuve & Vigna Taglianti, 1992, Carabus 126

robustoides Jeanne, 1996, Platyderus 523

robustus Coiffait, 1958, Aphoenops 294

robustus E. Deyrolle, 1869, Carabus 157



robustus Kraatz, 1878, Carabus 151

robustus J. Ochs, 1949, Laemostenus [Antisphodrus] [HN] 532

robustus Schaufuss, 1865, Laemostenus [Pristonychus] 534

robustus Kurnakov, 1961, Lindrothius 529



robustus Faldermann, 1836, Microderes 387

robustus Dejean, 1828, Molops [Feronia] 474

robustus Fairmaire, 1894, Myas [Aurisma] 476

robustus Dejean, 1831, Odotoncarus [Ditomus] 366

robustus Jeannel, 1928, Orotrechus 317

robustus Wollaston, 1854, Orthomus [Argutor] 476

robustus A. Morawitz, 1862, Panagaeus 447



robustus Straneo, 1941, Percus 478

robustus Jeannel, 1923, Perileptus 289

robustus Mateu, 1952, Platyderus 523

robustus Lassalle, 1993, Pseudotaphoxenus 542

robustus Uéno, 1958, Rakantrechus 319

robustus Sciaky, 1998, Stomis 518

robustus Habu, 1978, Synuchus 546

robustus N. Ito, 1999, Trichotichnus 394



robustus Motschulsky, 1862, Typhlotrechus [Anophthalmus] 344

robustus C. Zimmermann, 1831, Zabrus 571



roccae Born, 1923, Carabus 129

roccae Schatzmayr, 1925, Stomis 518

roccai F. Battoni, 1984, Calathus 526

roccai Schatzmayr, 1929, Pterostichus 503

roccai Jeannel, 1927, Trechus 334

rochasi Perroud & Montrouzier, 1864, Rhantaticus [Hydaticus] 49

rochetii Chaudoir, 1855, Scarites 233

rodriguezi Breuning & Ruspoli, 1977, Carabus 186

rodriguezi Pierre Meyer & Mollard, 1998, Carabus 167

roeri Blumenthal, 1976, Carabus 187

roeschkeanus Semenov, 1906, Carabus 138

roeschkei Csiki, 1901, Carabus 170

roeschkei Heller, 1923, Cychrus 205

roeselii Füessly, 1775, Cybister [Dytiscus] 51

rolex Morvan, 1995, Broscosoma 235

rolex Morvan, 1992, Deltocolpodes 457

rolex Morvan, 1991, Leistus 81

rolex Morvan, 1995, Pheropsophus 218

rolex Morvan, 1995, Pterostichus 516



rollei A. Fleischer, 1907, Diachromus 362

rolphi Kraatz, 1890, Calomera [Cicindela] 100

rolphii Fairmaire, 1867, Graphipterus 359

romanellus Imura, 1999, Carabus 159

romani Imura, 1998, Carabus 159

romani Raynaud, 1968, Carabus [Chrysocarabus] 140

romani Sciaky, 1994, Straneostichus 518

romanowi Semenov, 1897, Carabus 133

romanus Born, 1905, Carabus 166

romanus A. Fiori, 1904, Haliplus 30

romanus K. Daniel, 1903, Leistus 18, 84

romualdi J. Müller, 1905, Laemostenus 534

ronayi Csiki, 1906, Carabus 173

ronchettii Born, 1900, Carabus 151

ronchettii Reitter, 1909, Pterostichus 496

ronchettii Reitter, 1911, Trechus 339


ronganensis Deuve, 1998, Carabus 156

roni Fery, 1999, Hydroporus 63

roninus Bates, 1873, Harpalus 385

ronkayi Deuve, 1992, Carabus 183

roparzhemoni Morvan, 1982, Trechus 339

rosaceus Geoffroy, 1785, Platyderus [Buprestis] 522

rosai Bucciarelli, 1978, Boldoriella 296



rosai Bucciarelli, 1962, Duvalius 305

rosalesi Reiche, 1863, Carabus 167

rosaliae Deuve, 1997, Carabus 175



rosayanus A. Dufour, 1977, Carabus 171

rosea Naviaux, 1995, Neocollyris 116



rosenbergeri Seidlitz, 1887, Graphoderus 49

roseni Reitter, 1897, Carabus 169

roseni Jeannel, 1929, Duvalius 300

roseni A. Zimmermann, 1919, Hydroporus 64

roseni Brancsik, 1899, Microderes [Pangus] 387

roseri Faldermann, 1836, Carabus 181



rossicus Semenov, 1898, Brychius 30

rossii Schaum, 1857, Asaphidion [Tachypus] 243



rossii Depoli, 1929, Calathus 525

rossii Dejean, 1826, Carabus 130

rossii Magrini & Vanni, 1991, Duvalius 305

rossii Ponza, 1805, Gynandromorphus [Carabus] 362

rostagnoi Luigioni, 1904, Carabus 151

rostandi Deuve, 1991, Carabus 142

rosti Beuthin, 1885, Carabus [Melancarabus] 181

rosti Roeschke, 1907, Cychrus 202

rosti Shilenkov, 1999, Leistus 82



rostianus Roeschke, 1907, Carabus [Imaibius] [HN] 155

rostianus Semenov, 1907, Carabus 155



rostislavi Semenov, 1907, Calosoma 118

rostowtzowi Tschitschérine, 1896, Cymindis 416

rostowtzowi Tschitschérine, 1899, Poecilus [Feronia] 485

rostratus Motschulsky, 1862, Anophthalmus 292

rostratus Linné, 1761, Cychrus [Tenebrio] 203

rostratus Duftschmid, 1812, Stomis [Carabus] 518

rothi Dejean, 1829, Carabus 173

rothi Jeannel, 1926, Duvalius 299



rothschildi Born, 1889, Carabus [Coptolabrus] 141

rotroui Antoine, 1941, Amara 555

rotroui Antoine, 1938, Carabus 146

rotroui Pic, 1924, Neptosternus [Laccophilus] 77

rotroui Antoine, 1957, Poecilus 481

rotroui Antoine, 1934, Trechus 339



rottenmannicus Sokolá, 1907, Carabus 180

rotudangulus Jedlika, 1957, Harpalus 376

rotundangulus A. Morawitz, 1862, Pterostichus 511

rotundata Dejean, 1828, Amara 555

rotundata Schmidt‑Göbel, 1846, Dolichoctis [Mochtherus] 436

rotundatulus Lorenz, 1998, Chlaenius 355

rotundatum Chaudoir, 1852, Omophron 208

rotundatus Wehncke, 1872, Agabus 40

rotundatus Ledoux, 1989, Archileistobrius 79

rotundatus Jacquelin du Val, 1857, Calathus 524

rotundatus Born, 1895, Carabus 194

rotundatus Klug, 1832, Graphipterus 359

rotundatus Chaudoir, 1844, Harpalus 372

rotundatus Gschwendtner, 1931, Herophydrus [Hyphoporus] 71

rotundatus Paykull, 1790, Olisthopus [Carabus] 461

rotundatus Dejean, 1829, Ophonus [Harpalus] 389

rotundatus Aubé, 1836, Peltodytes [Cnemidotus] 33

rotundatus Chaudoir, 1866, Platyderus 523

rotundatus Yu, 1992, Straneostichus [Aristochroa] 518

rotundatus Dejean, 1831, Trechus 339

rotundatus Rambur, 1838, Zabrus 569



rotundicollis Carret, 1898, Acinopus 369

rotundicollis Motschulsky, 1865, Agonum [Agonocyrtes] 451

rotundicollis Schaufuss, 1862, Amara [Leiocnemis] 563

rotundicollis Solsky, 1875, Amara [Leiocnemis] 558

rotundicollis Dejean, 1828, Calathus 524

rotundicollis Mandl, 1955, Carabus 194

rotundicollis Rambur, 1837, Carterus 364



rotundicollis Dejean, 1826, Chlaenius 352

rotundicollis Mandl, 1978, Chlaenius 355

rotundicollis Faldermann, 1835, Dyschirius [Clivina] 229

rotundicollis Reitter, 1894, Eremosphodrus [Sphodrus] 532



rotundicollis Kolenati, 1845, Harpalus 374

rotundicollis Motschulsky, 1848, Harpalus [Bioderus] 374

rotundicollis Motschulsky, 1860, Leistus 82

rotundicollis Heinz & Ledoux, 1990, Nebria 90



rotundicollis Marsham, 1802, Olisthopus [Carabus] 461

rotundicollis Fairmaire & Laboulbène, 1854, Ophonus [Harpalus] 391

rotundicollis Duftschmid, 1812, Pterostichus [Carabus] 507

rotundicollis Stephens, 1835, Pterostichus [Omaseus] [HN] 511

rotundicollis Sturm, 1824, Pterostichus [Platysma] [HN] 486

rotundicollis Ménétriés, 1836, Zabrus 571



rotundicollis Rambur, 1838, Zabrus 569

rotundipennis Reiche, 1862, Masoreus 357

rotundipennis Chaudoir, 1843, Reicheiodes [Dyschirius] 230



rotundipennis Uéno, 1980, Trechiama 323

rotundipennis Duftschmid, 1812, Trechus [Carabus] 340



rotundipennis Fairmaire, 1859, Zabrus 568

rotundithorax Sciaky & Wrase, 1997, Pterostichus 501

rotundus Sciaky, 1994, Pterostichus 514

roubali Makolski, 1928, Amara 557

roubali Jeannel, 1926, Duvalius 307

roubali Maan, 1938, Dyschirius 229

roubali Schauberger, 1928, Harpalus 374

roubali Jedlika, 1951, Lebia 429



roubali Maan, 1941, Leistus 82

roubali Maan, 1933, Patrobus 285

roubali Schauberger, 1927, Pterostichus 488

roubaliana Lutshnik, 1934, Amara 562

roubalianus Lutshnik, 1928, Pterostichus [Platysma] 500



roubalianus Kult, 1946, Reicheiodes [Dyschirius] 230

rougemonti Morvan, 1997, Andrewesius 456

rougemonti Morvan, 1999, Aparupa 456

rougemonti Ledoux & Roux, 1988, Nebria 93



rougemonti Luna de Carvalho, 1989, Paussus [Cochliopaussus] 210

rougemonti Morvan, 1996, Platynus [Batenus] 463

rougemonti Mateu, 1990, Trechus 324

rougemontianus Lorenz, 1998, Paussus 210



rousi Vysoký, 1983, Bembidion 256

rousi Pulpán & Reška, 1977, Leistus 84

rousi Bulirsch & Hrka, 1994, Orientoreicheia 222

rousi Kirschenhofer, 1982, Pterostichus 496

rousianus Gottwald, 1985, Carabus 132

rousianus Kirschenhofer, 1987, Pterostichus 505

rousorum Fassati, 1992, Bembidion 268

rousseleti Ledoux & Roux, 1988, Nebria 87



roussellei Colas, 1969, Carabus 140

rousselli Colas, 1963, Pterostichus 498

rouxi Barthélemy, 1835, Calomera [Cicindela] 100

rouxi Deuve, 1995, Trechus 340

rouziei Breuning & Ruspoli, 1970, Carabus 158

rovirai Lagar Mascaró, 1975, Oscadytes 477

rovnicensis Daffner, 1996, Anophthalmus 292

rubefactus Bates, 1873, Harpalus 384

rubens Tschitschérine, 1898, Amara 552

rubens I. Frivaldszky von Frivald, 1835, Cicindela 104

rubens Fairmaire, 1872, Oedesis [Ditomus] 366

rubens Fabricius, 1792, Trechus [Carabus] 340

rubescens Jeanne, 1967, Cicindela 107

rubicunda Quensel, 1806, Nebria [Carabus] 93



rubicundus Coquerel, 1859, Orthomus [Feronia] 477

rubicundus Chaudoir, 1850, Tachys 275

rubidicollis Verhoeff, 1891, Calathus 527

rubidofemoratus Csiki, 1930, Pterostichus 503

rubidum Andrewes, 1924, Bembidion 262

rubidus Perris, 1857, Haliplus 32

rubizzaniorum Cavazzuti, 1994, Carabus 156

rubricata Motschulsky, 1864, Calleida 408

rubriceps Andrewes, 1934, Cymindis 414



rubricollis Bedel, 1910, Mimocolliuris [Ophionea] 444

rubricrus Géhin, 1885, Carabus 171

rubricrus Andrewes, 1934, Pristosia 531

rubriola Bates, 1883, Loxocrepis [Colpodes] 460

rubripalpis Csiki, 1930, Pterostichus 505

rubripennis Casey, 1918, Amara [Curtonotus] 559

rubripennis C. G. Thomson, 1857, Patrobus 285

rubripes Dejean, 1828, Abacetus [Feronia] 346

rubripes Dejean, 1831, Calathus 526



rubripes A. Nicolas, 1898, Carabus 139

rubripes Géhin, 1876, Carabus 136

rubripes Duftschmid, 1812, Harpalus [Carabus] 381



rubripes J. R. Sahlberg, 1875, Hydroporus 65

rubripes Ganglbauer, 1889, Molops 472

rubripes Audinet‑Serville, 1821, Nebria 87



rubripes A. Morawitz, 1862, Panagaeus 447

rubripes A. Fiori, 1896, Pterostichus 503

rubripes Motschulsky, 1860, Stereocerus [Steroderus] 517

rubripicta Andrewes, 1928, Parena 434

rubroangulata Baliani, 1938, Amara 557

rubrofemorata Shilenkov, 1975, Nebria 88



rubrotestacea A. Fleischer, 1892, Cymindis 413

rude De Monte, 1947, Bembidion 255

rudestriatus Reitter, 1883, Pterostichus [Feronia] 501

rudimentalis Chaudoir, 1868, Pterostichus [Feronia] 490

rudnicaensis Meggiolaro, 1961, Orotrechus 317



rudnikus Jedlika, 1938, Pterostichus 506

rudolphi Ganglbauer, 1891, Trechus 340

rueckbeili Breuning, 1932, Carabus 123

rueckbeilianus Deuve, 1994, Carabus 150

rueppelii Guérin‑Méneville, 1847, Salpingophora [Cicindela] 114

rufa Chaudoir, 1843, Siagona 218

rufangulus Marsham, 1802, Calathus [Carabus] 526

rufescens Dejean, 1829, Amara [Harpalus] 552



rufescens Breuning, 1932, Carabus 150

rufescens Gebler, 1845, Cymindis 416



rufescens Hope, 1838, Dromius 419

rufescens Jedlika, 1961, Jujiroa [Taphoxenus] 459



rufescens Fabricius, 1775, Leistus [Carabus] 83

rufescens Strøm, 1768, Nebria [Carabus] 88



rufescens Guérin‑Menéville, 1823, Ocys [Tachys] 272

rufescens Pic, 1908, Polistichus 574

ruffoanus Casale, Giachino, Vailati & Vigna Taglianti, 1996, Duvalius 306

ruffoi Magistretti, 1956, Duvalius 305

ruffoi Tamanini, 1954, Orotrechus 317

ruffoi Sciaky, 1984, Pterostichus 513

ruffoi Franciscolo, 1961, Scarodytes 69

rufibarbis Fabricius, 1792, Ophonus [Carabus] 391

rufibasis Reitter, 1893, Cymindis 415

ruficapillus Matits, 1910, Acupalpus 399

ruficapillus Mannerheim, 1852, Hydroporus 65

ruficeps Ménétriés, 1832, Agabus [Colymbetes] 37

ruficeps Chaudoir, 1873, Cymindis 412



ruficeps Schaum, 1857, Demetrias 418

ruficeps Chaudoir, 1861, Galerita 359

ruficeps Chevrolat, 1861, Haliplus 33

ruficeps Curtis, 1833, Harpalus 378

ruficeps Boheman, 1848, Herophydrus [Hydroporus] [add, syn of guineensis] 71

ruficeps Gozis, 1873, Lebia 429

ruficeps Motschulsky, 1845, Mastax 217

ruficeps Chaudoir, 1876, Styphlomerus 218

ruficolle Panzer, 1796, Bembidion [Elaphrus] 258

ruficollis Gautier des Cottes, 1858, Atranus [Anchomenus] 457

ruficollis Stephens, 1828, Bradycellus [Trechus] 401

ruficollis Gautier des Cottes, 1868, Calathus 527

ruficollis Schaller, 1783, Copelatus [Dytiscus] 47

ruficollis Gebler, 1845, Cymindis 417



ruficollis Graëlls, 1858, Cymindis 412

ruficollis Jeannel, 1937, Dolichus 530

ruficollis Kolenati, 1845, Dyschirius 229

ruficollis DeGeer, 1774, Haliplus [Dytiscus] 31



ruficollis Fabricius, 1787, Hydaticus [Dytiscus] 54

ruficollis Motschulsky, 1844, Paradromius [Dromius] 424



ruficollis Marsham, 1802, Platyderus [Carabus] 522

ruficollis Schilsky, 1908, Rhantus 47

ruficorne Fischer von Waldheim, 1829, Agonum 455

ruficorne Sturm, 1825, Bembidion 268

ruficornis Chaudoir, 1850, Aephnidius 357



ruficornis Dejean, 1828, Amara 556

ruficornis J. R. Sahlberg, 1880, Amara 561

ruficornis Schaufuss, 1882, Amblystomus 360

ruficornis Kraatz, 1882, Carabus 170

ruficornis Lucas, 1842, Carterus [Ditomus] 364

ruficornis Putzeys, 1846, Dyschirius 230



ruficornis Fabricius, 1775, Harpalus [Carabus] 385

ruficornis Zetterstedt, 1838, Hydroporus 62

ruficornis Audouin & Brullé, 1839, Neocollyris [Collyris] 117

ruficornis Goeze, 1777, Paranchus [Carabus] 462

ruficrus Kirby, 1837, Dicheirotrichus [Trechus] 403

ruficrus Ménétriés, 1832, Ophonus [Harpalus] 388

rufifemoratus Stephens, 1828, Poecilus 482

rufifemoratus A. Fiori, 1896, Pterostichus 503

rufifemoratus Stephens, 1828, Pterostichus [Omaseus] 511

rufifrons Duftschmid, 1805, Hydroporus [Dytiscus] 64

rufifrons O. F. Müller, 1776, Hydroporus [Dytiscus] 64



rufifrons Fabricius, 1792, Suphrodytes [Dytiscus] 70

rufifrons Chaudoir, 1862, Zuphium 574

rufilabris Fischer von Waldheim, 1828, Dromius 421



rufilabris Motschulsky, 1864, Dyschirius 225

rufilabris Motschulsky, 1845, Loricera 98

rufimacula J. Müller, 1918, Bembidion 253



rufimana Kirby, 1837, Amara [Curtonotus] 561

rufimana Motschulsky, 1844, Amara [Leirus] 561

rufimanus A. Fleischer, 1898, Dyschirius 224



rufimanus LeConte, 1848, Harpalus 378

rufimanus Marsham, 1802, Harpalus [Carabus] 383

rufimanus Chaudoir, 1846, Pterostichus [Feronia] 495

rufimanus White, 1846, Rhantus [Colymbetes] 47

rufino J. R. Sahlberg, 1913, Nebrioporus 67

rufinus Beheim & Breuning, 1943, Carabus [Eucarabus] 133

rufipalpis Sturm, 1818, Harpalus 381

rufipalpis Chaudoir, 1865, Neocollyris [Collyris] 117



rufipalpis Chaudoir, 1846, Pterostichus [Feronia] 505

rufipalpis Schaum, 1864, Zabrus 570

rufipennis Gschwendtner, 1933, Agabus [Gaurodytes] 36



rufipennis Beuthin, 1885, Cicindela 104

rufipennis Dejean, 1828, Penetretus [Patrobus] 285



rufipennis Baudi di Selve, 1890, Pterostichus 503

rufipennis Tanaka, 1958, Pterostichus [HN] 501

rufipes Dejean, 1828, Amara 568



rufipes Dejean, 1831, Bembidion 259

rufipes Gyllenhal, 1810, Bembidion 259

rufipes Illiger, 1801, Bembidion [Elaphrus] 268

rufipes Paykull, 1790, Bembidion [Carabus] 251

rufipes Guérin‑Méneville, 1829, Broscus [Cephalotes] 236

rufipes Boheman, 1858, Caelostomus [Drimostoma] 471

rufipes Fabricius, 1792, Calathus [Carabus] 526

rufipes A. Nicolas, 1898, Carabus 197

rufipes Kraatz, 1879, Carabus 196

rufipes Semenov, 1887, Carabus 176

rufipes Chaudoir, 1843, Carterus [Odogenius] 364



rufipes Dejean, 1826, Chlaenius 352

rufipes Štrba, 1910, Chlaenius 351

rufipes Jedlika, 1940, Chydaeus 362



rufipes Motschulsky, 1861, Clivina [Eupalamus] 220

rufipes Gebler, 1825, Cymindis 416


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