Estimating survival and exploitation rates from a tagging study on juvenile Atlantic halibut

Summary and recommendations 5. References

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4. Summary and recommendations

5. References.

Brownie C., Anderson D.R., Burnham K.P. and Robson D.S. (1985). Statistical inference from band recovery data: a handbook. United States Fish and Wildlife Service Resource Publication Number 156.
Cooch E.G. and White G.C. (2007). Program MARK: A gentle introduction. http://www.phidot.org/software/mark/docs/book/.
Hoenig J. M., Barrowman N.J., Hearn W. S. and Pollock K. H. (1998). Multiyear tagging studies incorporating fishing effort data. Canadian Journal of Fisheries and Aquatic Sciences 55, 1466-1476.
Hoenig J. M., Barrowman N. J., Pollock K. H., Brooks E. N., Hearn W. S. and Polachek T. (1998b). Models for tagging data that allow for incomplete mixing of newly tagged animals. Canadian Journal of Fisheries and Aquatic Sciences 55, 1477-1483.
Nichols J. D., Stokes S. L., Hines J. E. and Conroy M. J. (1982). Additional comments on the analysis of homogeneous survival rates in modern bird banding estimation models. Journal of Wildlife Management 46, 953-962.
Pollock K. H. and Raveling D. G. (1982). Assumptions of modern band-recovery models with emphasis on heterogeneous survival rates. Journal of Wildlife Management 46, 88-98.
Lebreton J.-D., Burnham K. P., Clobert J. and Anderson D. R. (1992). Modelling survival and testing biological hypotheses using marked animals. A unified approach with case studies.. Ecological Monographs 62, 67-118.
White G. C. and Burnham K. P. (1999). Program MARK: Survival estimation from populations of marked animals. Bird Study 46 (suppl), s120-s139.
White, G.C. (1992). PC SURVIV Users Manual. Dept. of Fishery and Wildlife Biology, Colorado State University, Fort Collins, CO. Available from: http://www.mbr-pwrc.usgs.gov/software.html.

Figure 1. Fates of animals in the Brownie model.

The parameter S is the probability that the fish survives the year; K is the probability that the fish is harvested; c is the probability that the tag is retrieved from the fish; and is the probability that the tag is read properly and reported to the tagging agency.

Figure 2. A simplified fate diagram for the Brownie et al (1985) model.

The parameter S is the probability that the fish survives the year;

Figure 3. Standardized residual

plot from Hoenig et al. (1998) model M(*) F(t) assuming that

Table 1. Raw data from the Brownie et al (1985) model arranged in tabular format.
		Recovery Years
Year	Number Tagged	1	2	3
1
2
3

Table 2. Expected number of recoveries in the Brownie et al. (1985) model arranged in tabular format.
		Recovery Years
Year	Number Tagged	1	2	3
1
2
3

Table 3. Expected number of recoveries in the Hoenig et al (1998) model when fishing is continuous over the year. This is a reparameterization ot Table 2 assuming that and are constant over time.
		Recovery Years
Year	Number Tagged	1	2	3
1
2
3

Table 4. Number of halibut released by month and year. Only those fish with a known release date are reported
Year of Release	Month of release												Total Releases
Year of Release	1	2	3	4	5	6	7	8	9	10	11	12	Total Releases
1995	.	.	62	122	136	150	73	53	13	35	.	.	644
1996	.	.	.	.	.	.	.	.	5	8	1	20	34
1997	.	5	24	10	32	62	46	.	21	.	.	.	200
1998	.	.	225	105	145	260	131	6	6	4	2	.	884
1999	.	90	59	69	111	207	279	68	4	4	.	.	891
2000	.	.	.	5	55	198	61	30	.	.	.	.	349
2001	.	.	.	122	58	302	290	21	.	2	.	.	795
2002	.	.	.	.	.	6	42	.	.	.	.	.	48
2003	.	.	.	.	.	5	4	17	.	.	.	.	26
2004	.	.	.	.	.	85	221	37	21	.	.	.	364
2005	15	35	8	.	.	237	49	3	3	.	.	.	350
2006	31	22	1	.	42	691	602	86	.	.	.	.	1475
2007	.	.	.	.	326	.	596	.	.	.	.	.	922
All	46	152	379	433	905	2203	2394	321	73	53	3	20	6982

Table 5. Number of halibut recovered by year and month of recovery. Only those fish with known recovery times are displayed.
Year	Month of recaptures												All
Year	1	2	3	4	5	6	7	8	9	10	11	12	All
1995	.	.	.	.	1	1	1	2	3	1	1	.	10
1996	1	.	.	11	2	.	8	7	1	1	1	1	33
1997	.	2	1	.	4	4	5	1	2	.	3	2	24
1998	.	8	3	2	8	10	6	2	2	4	1	1	47
1999	1	2	3	11	11	18	9	6	4	.	1	.	66
2000	3	15	18	.	4	16	15	8	2	.	4	1	86
2001	14	15	7	3	3	4	9	6	7	9	4	1	82
2002	16	15	7	8	3	8	13	6	4	4	1	1	86
2003	7	5	8	6	9	7	15	6	6	3	.	.	72
2004	10	6	7	.	.	2	8	3	5	3	.	4	48
2005	6	4	4	3	.	1	5	1	3	.	.	1	28
2006	2	3	2	1	.	5	7	8	12	7	5	.	52
2007	5	13	15	2	4	7	11	.	.	.	.	.	57
All	65	88	75	47	49	83	112	56	51	32	21	12	691

Table 6. Summary statistics for the halibut program for the Hoenig et al (1998) model. Only those fish with known dates of release and recovery are used. The data may not be consistent with Tables 4 and 5 because of problems in the database, e.g. a recovered fish may not have a legible tag number; the tag number of a released fish may not be known. The “recovery year” extends from 1 May of the year to 30 April of the next year (inclusive).
Release Year	Recovery year
Release Year	Releases	1995	1996	1997	1998	1999	2000	2001	2002	2003	2004	2005	2006	2007	Never recovered
1995	648	22	24	29	21	20	9	7	1	3	.	.	1	.	511
1996	34	.	.	.	.	2	.	.	2	.	.	1	1	.	28
1997	200	.	.	4	3	7	6	3	2	.	1	.	.	.	174
1998	887	.	.	.	24	36	35	15	10	7	2	4	3	1	750
1999	892	.	.	.	.	12	23	27	11	12	9	3	2	.	793
2000	349	.	.	.	.	.	1	9	9	8	2	.	.	.	320
2001	799	.	.	.	.	.	.	11	19	22	12	3	5	5	722
2002	48	.	.	.	.	.	.	.	.	.	.	.	.	.	48
2003	26	.	.	.	.	.	.	.	.	.	1	.	.	.	25
2004	364	.	.	.	.	.	.	.	.	.	5	2	5	1	351
2005	350	.	.	.	.	.	.	.	.	.	.	1	2	.	347
2006	1475	.	.	.	.	.	.	.	.	.	.	.	51	10	1414
2007	922	.	.	.	.	.	.	.	.	.	.	.	.	.	922

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Halibut -> Atlantic Halibut fishing mortality estimated from tagging on the Scotian Shelf and the southern Grand Banks

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