Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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cordaticollis Heyden, 1884, Pterostichus 509

cordaticollis Schauberger, 1936, Trichotichnus 394

cordatissimus Straneo, 1937, Pterostichus [Feronia] 497

cordatoides Schauberger, 1926, Ophonus [Harpalus] 389

cordatulus Reitter, 1901, Microlestes 423



cordatus Lapouge, 1919, Carabus 192

cordatus Schaufuss, 1879, Dicheirotrichus [Bradycellus] 402

cordatus Chevrolat, 1854, Laemostenus [Dolichus] 539

cordatus Chaudoir, 1861, Licinus 442

cordatus Duftschmid, 1812, Ophonus [Carabus] 389

cordatus Letzner, 1842, Pterostichus 494

cordatus Uéno, 1990, Trechiama 322

cordatus Rambur, 1837, Trymosternus [Cymindis] 410

cordatus Dejean, 1825, Tschitscherinellus [Ditomus] 367

cordicervix Csiki, 1927, Carabus 169

cordicolle Jacquelin du Val, 1852, Bembidion 260



cordicollis Chaudoir, 1844, Amara [Bradytus] 552

cordicollis Ménétriés, 1832, Amara 562



cordicollis Motschulsky, 1862, Anophthalmus 292

cordicollis Chaudoir, 1846, Anthracus [Acupalpus] 399

cordicollis Chaudoir, 1843, Aptinus 212



cordicollis Wesmael, 1835, Bradycellus [Acupalpus] 401

cordicollis Chaudoir, 1842, Broscus 236

cordicollis Ganglbauer, 1886, Carabus [Tribax] 169

cordicollis Motschulsky, 1866, Carabus 187

cordicollis Motschulsky, 1865, Chlaenius 350

cordicollis Motschulsky, 1861, Clivina [Eupalamus] 220

cordicollis Chaudoir, 1835, Cychrus 203

cordicollis V. E. Jakovlev, 1887, Cymindis 415

cordicollis Fairmaire, 1868, Dicheirotrichus [Bradycellus] 402

cordicollis LaFerté‑Sénectère, 1851, Diplocheila [Rembus] 439



cordicollis Vorbringer, 1898, Dromius 420

cordicollis Audinet‑Serville, 1821, Graniger [Harpalus] 366

cordicollis Chaudoir, 1854, Laemostenus [Sphodrus] 539

cordicollis Sciaky & Wrase, 1997, Myas [Trigonognatha] 475

cordicollis Chaudoir, 1837, Nebria 93

cordicollis Dejean, 1829, Ophonus [Harpalus] 391

cordicollis H. Kolbe, 1886, Planetes 359

cordicollis Motschulsky, 1850, Pogonistes [Pogonus] 286


cordicollis Uéno, 1974, Trechiama 322

cordicollis Winkler, 1936, Trechus 339

cordicollis Morvan, 1979, Xestopus [Wittmerosphodrus] 530

cordifer Reitter, 1896, Pterostichus 504



cordiger Chaudoir, 1868, Pterostichus [Feronia] 505

cordimeatus Cavazzuti, 1997, Carabus 150

cordithoracicus Deuve, 1989, Carabus 128

cordubensis Dieck, 1869, Geocharis [Anillus] 239

cordulatus Cavazzuti, 1999, Carabus 129



coreana Baliani, 1937, Amara 560

coreana H. Kolbe, 1886, Galerita 359

coreana Jedlika, 1932, Tachyura [Tachys] 279

coreanicus Breuning, 1950, Carabus 159

coreanum Jedlika, 1946, Bembidion 266

coreanus Mlyná, 1974, Dicheirotrichus 402

coreanus Tschitschérine, 1895, Harpalus [Ophonus] 384

coreanus Tschitschérine, 1895, Myas [Feronia] 475

coreanus Nakane, 1997, Panagaeus 447

coreanus H. Kolbe, 1886, Scarites 232

coreanus Kirschenhofer, 1990, Synuchus 545

coreica Jedlika, 1936, Morphodactyla [Dolichus] 530

coreica Solsky, 1875, Nebria 94

coreicola Deuve, 1989, Carabus 183

coreicus G. Hauser, 1921, Carabus [Coptolabrus] 144

coreicus Breuning, 1950, Cychrus 205

coreicus Jedlika, 1962, Pterostichus 490

coriacea Habu, 1973, Atamuka [Kumataia] 456

coriacea Laporte, 1835, Meladema 45

coriaceipennis Chaudoir, 1863, Carabus 198



coriaceus J. R. Sahlberg, 1875, Agabus [Gaurodytes] 37

coriaceus Géhin, 1885, Carabus 192

coriaceus Linné, 1758, Carabus 187

corinna Holdhaus, 1924, Reicheia 222

cornifer Kasahara & Y. Itô, 1994, Apatrobus 280

cornilloni Giordan & Raffaldi, 1998, Duvalius 302

cornutus Dejean, 1825, Ditomus [Ditomus] 365

cornutus Uéno, 1983, Trechiama 322

coronatus Cavazzuti, 1996, Cychrus 203

corporaali Netolitzky, 1935, Bembidion 262

corporosus Motschulsky, 1861, Harpalus [Pheuginus] 374

corpulenta Putzeys, 1866, Amara [Leiocnemis] 555



corpulentior Deuve, 1990, Carabus 197

corpulentum Motschulsky, 1850, Bembidion [Peryphus] 263

corpulentus Kraatz, 1880, Carabus 196



corpulentus Motschulsky, 1860, Chlaenius 352

corpulentus Weise, 1875, Duvalius [Trechus] 306

corpulentus Schatzmayr, 1903, Gyrinus 28

corpulentus Schaum, 1864, Hygrotus [Hydroporus] 71



corpulentus Chaudoir, 1859, Molops 475

corpulentus Schaum, 1860, Neophygas [Harpalus] 387

corpulentus Gebler, 1829, Pterostichus [Omaseus] 510

corpulentus Schaum, 1864, Zabrus 570

corradoi E. Porro & G. Porro, 1968, Molops [Typhlochoromus] 475

correiroi Schatzmayr, 1943, Harpalus 375



corrosa Reiche, 1855, Cymindis 411

corrosus Bates, 1873, Platymetopus 393

corrugatius J. R. Sahlberg, 1880, Pterostichus [Feronia] 499

corrugatus Billberg, 1815, Percus [Scarites] 478

corrugis C. A. Dohrn, 1882, Carabus 145

corrugulus Mandl, 1955, Carabus 145

corsica Putzeys, 1866, Amara [Leiocnemis] 557

corsica Rivalier, 1962, Cylindera 111



corsica Gautier des Cottes, 1870, Tachyura [Tachys] 278

corsicana Roeschke, 1891, Cicindela 104

corsicanus Lapouge, 1913, Carabus 161

corsicum Csiki, 1928, Bembidion 269



corsicus Guignot, 1932, Agabus 39

corsicus Perris, 1869, Anillus 238



corsicus Perris, 1866, Anthracus [Acupalpus] 400

corsicus Born, 1906, Carabus 136

corsicus O. Schneider, 1903, Hydroglyphus [Bidessus] 56

corsicus Wehncke, 1872, Hydroporus 64

corsicus Coiffait, 1968, Laemostenus [Ceuthosphodrus] 532

corsicus Audinet‑Serville, 1821, Percus [Harpalus] 478



corsicus Tournier, 1860, Platynus [Anchomenus] 464

corticalis Motschulsky, 1844, Carabus 136

corticalis L. Dufour, 1820, Microlestes [Lebia] 422

coruhnehriensis Cavazzuti, 1990, Carabus 193



coruscus Fischer von Waldheim, 1828, Badister 441

coruscus Tschitschérine, 1895, Pterostichus [Feronia] 493

coruscus Tschitschérine, 1895, Trichotichnus [Harpalus] 395

corustes Andrewes, 1932, Licinus 442



corvinus Motschulsky, 1866, Carabus 177

corvus Duftschmid, 1812, Harpalus [Carabus] 375

cosinensis J. Müller, 1926, Typhlotrechus 344

cossyphoides Duftschmid, 1812, Licinus [Carabus] 442

costae Géhin, 1885, Carabus 130

costae Heyden, 1891, Zabrus 570



costai Emery, 1872, Cychrus 19, 203

costata Ménétriés, 1851, Pelophila 98

costata Gyllenhal, 1808, Yola [Hyphydrus] 57

costatorugosus Mandl, 1955, Carabus 145

costatulus Quensel, 1806, Brachinus 215



costatus Gerhardt, 1910, Agabus 41

costatus Morvan, 1980, Broscus 236

costatus Germar, 1824, Carabus 146

costatus Kraatz, 1878, Carabus 155

costatus Chaudoir, 1879, Oxylobus 231

costatus Ménétriés, 1851, Pterostichus [Lyperopherus] 497

costellatus Géhin, 1882, Carabus 139

costiger Chaudoir, 1856, Chlaenius 353



costiger Tschitschérine, 1900, Lesticus 471

costipennis Brancucci, 1983, Deronectes 58



costipennis Apfelbeck, 1918, Molops 473

costipennis Stöcklein, 1944, Platyderus 523

costipennis Faldermann, 1836, Zabrus 569

costulata Motschulsky, 1865, Sericoda [Batenus] 466

costulatus Motschulsky, 1859, Agabus [Colymbetes] 36



costulatus Motschulsky, 1862, Anophthalmus 291

costulatus Motschulsky, 1844, Brachinus 213

costulatus Kraatz, 1889, Carabus [Coptolabrus] 144

costulatus Petri, 1912, Carabus 199

costulatus Motschulsky, 1859, Chlaenius [Agostenus] 348



costulatus Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 542

costulatus Motschulsky, 1850, Pterostichus [Pseudomaseus] 511

costulatus Fairmaire, 1859, Scarites 233

costulifer A. Fleischer, 1898, Carabus 154

costulipennis Liebke, 1928, Brachinus 215

costulomicans Deuve & Tian, 1999, Carabus 125

costulus Géhin, 1885, Carabus 155



cosynsi Cabidoche, 1962, Aphoenops 294

cottarellii Breuning & Ruspoli, 1970, Carabus 156

cottellii Duftschmid, 1812, Molops [Carabus] 472

cottianus Jeannel, 1942, Pterostichus 511

cottyi Coquerel, 1859, Amara 555

coxalis Sharp, 1882, Agabus 36

coxalis Sharp, 1882, Bidessus 55

coxalis Fischer von Waldheim, 1844, Chlaenius 349

cracens Obydov, 1999, Carabus 174



cracoviensis Mazur, 1936, Dromius 420

crassa Stephens, 1828, Amara 553

crassecostulatus Deuve, 1991, Carabus 153



crassescens Péringuey, 1896, Tachyura [Tachys] 278

crassesculptus Kraatz, 1881, Carabus 182

crassethoracis Kleinfeld, 1999, Carabus 182


crassiceps Bates, 1874, Nebria 85

crassiceps A. Morawitz, 1862, Pterostichus 505

crassicerus N. Ito, 1985, Bradycellus 401



crassicollis Chaudoir, 1865, Neocollyris [Collyris] 117

crassicollis A. Morawitz, 1862, Pterostichus 509

crassicollis Jeannel, 1949, Trichaphaenops 344

crassicorne Putzeys, 1872, Bembidion 244

crassicornis Fairmaire, 1877, Laemostenus [Pristonychus] 536

crassicornis Motschulsky, 1864, Lebia [Lamprias] 428

crassicornis Dejean, 1825, Neocollyris [Colliuris] 117

crassicornis O. F. Müller, 1776, Noterus [Dytiscus] 34

crassicornis Fischer von Waldheim, 1842, Poecilus [Feronia] 483

crassiforma Ledoux & Roux, 1989, Nebria 94

crassilobatus Uéno, 1977, Trechiama 322

crassimargo Wollaston, 1865, Broscus 236

crassipalpis Bates, 1883, Dromius 420

crassipalpis Bates, 1873, Parabroscus [Taphria] 544

crassipes Sharp, 1882, Cybister 50

crassipes Duftschmid, 1812, Harpalus [Carabus] 375

crassipes Fischer von Waldheim, 1829, Poecilus [ND] 485



crassipes Ménétriés, 1832, Pterostichus [Feronia] 510

crassipes Uéno, 1997, Trechiama 322

crassiusculus Lewis, 1888, Omoglymmius [Rhysodes] 78

crassiusculus Fairmaire & Laboulbène, 1854, Ophonus [Harpalus] 389

crassiusculus Chaudoir, 1868, Pterostichus [Feronia] 496



crassus Lapouge, 1898, Carabus 195

crassus Sharp, 1882, Cybister 50

crassus Tschitschérine, 1895, Harpalus 379

crassus Coiffait, 1955, Hydraphaenops [Aphoenops] 311



crassus Wollaston, 1867, Hyphydrus 74

crassus C. G. Thomson, 1856, Ilybius 42

crassus Bates, 1883, Leistus 80

crassus Dejean, 1828, Zabrus 570

crates Bates, 1883, Harpalus 374

cratocephaloides Semenov, 1887, Carabus 121

cratocephalus Tschitschérine, 1897, Pterostichus [Feronia] 485

creachi Giordan & Raffaldi, 1983, Duvalius 302

crebrepunctatus Straneo, 1937, Pterostichus [Feronia] 515



crebrior Tschitschérine, 1902, Ophonus [Harpalus] 390

creda Jedlika, 1957, Amara 550

crenata Dejean, 1828, Amara 553



crenata Chaudoir, 1844, Cymindis 411

crenata Putzeys, 1873, Pristosia [Calathus] 531



crenatocostatus Lapouge, 1922, Carabus 166

crenatostriata Chaudoir, 1846, Amara [Bradytus] 553

crenatostriata Bassi, 1834, Nebria 85

crenatostriatus Chaudoir, 1846, Poecilus [Feronia] 479

crenatostriatus Fischer von Waldheim, 1829, Poecilus [HN] 480

crenatostriatus Stephens, 1828, Poecilus 483

crenatum Dejean, 1831, Bembidion 267

crenatus Dejean, 1828, Abax 469

crenatus Sturm, 1815, Carabus 166

crenatus L. Redtenbacher, 1843, Laemostenus [Pristonychus] 536

crenatus Fairmaire, 1855, Leistus 82

crenatus Grouvelle, 1903, Omoglymmius [Rhysodes] 78

crenatus Dejean, 1828, Poecilus [Feronia] 479

crenatus Duftschmid, 1812, Pterostichus [Carabus] 486

crenicollis Schaufuss, 1869, Broscus 236

crenifer Tschitschérine, 1903, Leistus 80

crenulatipennis Imura, 1996, Carabus 126

crenulatopunctatus R. F. Sahlberg, 1844, Pterostichus [Feronia] 509

crenulatum R. F. Sahlberg, 1844, Bembidion 265

crenulatus Putzeys, 1866, Dyschirius 230

crenulatus Dejean, 1828, Poecilus [Feronia] 481

crenulatus Chaudoir, 1842, Scarites 234

crenuliger Chaudoir, 1876, Poecilus 479

creper Tschitschérine, 1902, Pterostichus [Platysma] 493

crepidoderus Schaum, 1864, Zabrus 569

crepitans Linné, 1758, Brachinus [Carabus] 213

cretensis Fery, 1992, Bidessus 55

cretensis Maan, 1941, Leistus 84

creticola Antoine, 1928, Trechus 342

creticum J. Müller, 1918, Bembidion 263

creticus Maan, 1947, Acinopus 368

creticus Pic, 1903, Aptinus [Brachinus] 212

creticus Maan, 1934, Harpalus 377

creticus Holdhaus, 1912, Microlestes 422

creticus I. Frivaldszky von Frivald, 1845, Tapinopterus [Omaseus] 519

creticus Reitter, 1889, Zabrus 571

creutzeri Fabricius, 1801, Carabus 184



cribellata Chaudoir, 1860, Neocollyris [Collyris] 116

cribellatus M. F. Adams, 1812, Carabus 182



cribellum Stephens, 1828, Ophonus 391

cribellus Ménétriés, 1851, Pterostichus [Lyperopherus] 499

cribrata Putzeys, 1866, Amara [Curtonotus] 558

cribratellus Fairmaire, 1880, Methles [Hydroporus] 75

cribratellus Régimbart, 1891, Orectochilus 29



cribratus Motschulsky, 1850, Carabus 185

cribratus Quensel, 1806, Carabus 147

cribratus Reiche & Saulcy, 1855, Carterus [Ditomus] 364


cribratus Semenov, 1889, Elaphrus 207

cribratus Peyron, 1858, Ophonus [Harpalus] 390

cribratus Dejean, 1828, Pterostichus [Feronia] 503

cribrellus Reiche & Saulcy, 1855, Ophonus 390

cribricollis Chaudoir, 1846, Amara [Curtonotus] 559

cribricollis Motschulsky, 1844, Carabus 199

cribricollis Zoubkoff, 1833, Chlaenius 348

cribricollis Reiche, 1863, Cymindis 413

cribricollis A. Morawitz, 1862, Lachnolebia [Lebia] 427

cribricollis Dejean, 1829, Ophonus [Harpalus] 389

cribricollis Motschulsky, 1850, Pterostichus [Omaseus] 500

cribripennis Chaudoir, 1842, Harpalus 385

cribrithorax A. Morawitz, 1886, Carabus 147

cribrosa Chaudoir, 1865, Neocollyris [Collyris] 116

cribrosus Schaum, 1864, Clypeodytes [Hydroporus] 55



cribrulum Netolitzky, 1910, Bembidion 262

cribrum Jacquelin du Val, 1852, Bembidion 267

crimeanus Straneo, 1960, Poecilus [Pterostichus] 481

crinifer Balkenohl, 1993, Dyschirius 225



criniticoxis Larson, 1975, Hydroporus 61

crispata Germar, 1817, Yola [Hydroporus] 57

crissolensis Dodero, 1924, Doderotrechus [Duvalites] 298



cristata Lacordaire, 1835, Yola [Hydroporus] 57

cristatoides Straneo, 1955, Pterostichus 494



cristatomarginatus Goeze, 1777, Hydroporus [Dytiscus] 63

cristatus J. R. Sahlberg, 1875, Brychius 30

cristatus L. Dufour, 1820, Pterostichus [Harpalus] 512

cristianofonti Deuve & Font, 1998, Carabus 141

cristicaudis Kurnakov, 1962, Pterostichus 496

cristoforii Spence, 1833, Carabus 197

crnuljensis Absolon & Maan, 1947, Neotrechus 315



croatica Ganglbauer, 1891, Amara 555

croaticus Dejean, 1826, Carabus 163



croaticus J. Müller, 1951, Laemostenus 535

croaticus Székessy, 1938, Lymnastis 274



croaticus Kraatz, 1875, Molops 475

croaticus J. Müller, 1943, Ocys [Bembidion] 273

croaticus Dejean, 1831, Trechus 329

croaticus Hampe, 1871, Typhlotrechus [Anophthalmus] 344

crocatus Bates, 1883, Synuchus [Pristodactyla] 545

croceus Angus, Fresneda & Fery, 1993, Nebrioporus 66



crocopus Stephens, 1828, Calathus 527

croesus G. Hauser, 1912, Carabus [Coptolabrus] 143

croissandeaui Argod‑Vallon, 1888, Aphoenops [Anophthalmus] 294

crotchi Zaitzev, 1905, Agabus 41

crotchi Sharp, 1874, Hydroscapha 25

crotchi Preudhomme de Borre, 1871, Nebrioporus [Hydroporus] 66

crotchianus Wollaston, 1865, Zargus 440

crotchii Wollaston, 1864, Bembidion 243



crouzeti Puel, 1931, Anisodactylus 361

crucialis Régimbart, 1899, Sandracottus 49

cruciata Fischer von Waldheim, 1820, Cymindis [Anomoeus] 414

cruciatum Dejean, 1831, Bembidion 260



cruciatum Schiødte, 1841, Bembidion [HN] 260

cruciatus Quensel, 1806, Brachinus 214

cruciatus Dejean, 1831, Chlaenius 355

cruciatus Schilsky, 1890, Graptodytes [Hydroporus] 59

crucifer Geoffroy, 1785, Badister [Buprestis] 440

crucifer L. Redtenbacher, 1867, Macrocheilus [Planetes] 407

crucifer Lucas, 1846, Philorhizus [Dromius] 425

crucifer Piochard de la Brûlerie, 1876, Trechus 329

crucifera Reitter, 1923, Cymindis 417

crucifera Uéno, 1964, Eustra 208



cruciger Fabricius, 1798, Acupalpus [Carabus] 398

cruciger Heyden, 1889, Agatus [Glycia] 24, 408

cruciger Putzeys, 1875, Tachys 276

crudus Darge, 1992, Carabus 158

cruralis Fischer von Waldheim, 1829, Chlaenius 352

cruralis Chaudoir, 1879, Loxocrepis [Colpodes] 460

cruralis Tschitschérine, 1902, Pterostichus [Platysma] 497

cruris Csiki, 1927, Carabus 168

crux Fabricius, 1792, Rhithrodytes [Dytiscus] 68



crux Putzeys, 1875, Tachyura [Tachys] 279

cruxmajor Linné, 1758, Panagaeus [Carabus] 447

cruxminor Linné, 1758, Lebia [Carabus] 428

cruxpallidus Mandl, 1986, Callistus [Callistomimus] 347

cryobioides Chaudoir, 1868, Pterostichus [Feronia] 486

crypticola Linder, 1859, Aphoenops [Anophthalmus] 294

crypticola Leoni, 1907, Laemostenus 533

crypticola Jeanne, 1996, Platyderus 522

cryptophilus Belousov & Kabak, 1992, Trechus 329

crystallina Tschitschérine, 1903, Amara 562



csatoi Csiki, 1913, Duvalius [Anophthalmus] 300

csikensis Csiki, 1942, Carabus 135

csikii Jedlika, 1968, Amara 562

csikii Jedlika, 1932, Bembidion 250

csikii Laczo, 1912, Bradycellus 400

csikii Jedlika, 1958, Calathus 526

csikii Mallász, 1900, Carabus 151

csikii Jedlika, 1967, Cymindis 417


csikii Mihók, 1912, Duvalius [Anophthalmus] 299

csikii Jedlika, 1967, Harpalus 383


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