Estimating survival and exploitation rates from a tagging study on juvenile Atlantic halibut



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Table 7. Summary of recovery of high and low value

tags for releases in 2006 and 20077




Released

Recovery Year

2006

2007

Not

recoveried

2006

$0

423

.

2

421

$100

1052

51

8

993

2007

$0

7

.

.

7

$100

915

.

.

915



Table 8. Estimates of survival and tag-recovery from the Hoenig et al (1988) model denoted by M(*) F(t) assuming

Instantaneous natural survival rate (M)




Estimate

SE

All years

0.084

0.026










Instantaneous Tag-recovery rates ()

Year

Estimate

SE

1995

0.036

0.008

1996

0.042

0.009

1997

0.048

0.009

1998

0.033

0.005

1999

0.036

0.004

2000

0.033

0.004

2001

0.026

0.004

2002

0.022

0.004

2003

0.024

0.004

2004

0.013

0.003

2005

0.006

0.002

2006

0.007

0.003

2007

0.004

0.002


Table 9. Goodness-of-fit (observed vs expected) test from Hoenig et al (1998) model M(*) F(t) as computed by Surviv

Release

Cohort


df

Chi-square

Goodness-of-fit



p-value

1995

11

18.6

0.068

1996

2

4.0

0.133

1997

8

8.4

0.393

1998

9

16.6

0.055

1999

8

19.4

0.012

2000

6

15.6

0.016

2001

6

17.6

0.007

Total

56

142.0

.000

Details at the individual cell level are available from the authors.

Appendix A

Surviv code used for Hoenig et al (1998) model.
Halibut tagging experiment
Hoenig et al (1998) model with constant survival rate and year varying recovery rates.

See Hoenig et al (1988) CJFAS 55, 1466-1476


Parameterized in term of M (instantaneous natural mortality rate) and

f(i) - instantaneous fishing mortality rate


It is assumed that reporting rate is 100% as no reward bands applied to estimate

this rate.

It is assumed that tag shedding is zero as no double tags were applied to estimate

this rate.

This code can be used directly with the online surviv program at Patuxent.

Visit: http://www.mbr-pwrc.usgs.gov/software/surviv.html


Releases took place from 1999 to 2005 (7 years).

Recoveries took place from 1999 to 2007 (13 years).


/* Number of parameters is 14

Parms 1 to 13 are the F

parm 14 is M */

------ Cut and paste code below into the code box at ---------

http://www.mbr-pwrc.usgs.gov/software/surviv.html
-----------------------------------------------------------------------

proc title 'Halibut model - Hoenig et al (1998) model M(*) F(t) .1';


proc model npar=14 addcell;
inline declare f(i)=s(i);
inline do i=1,7 /* loop for each band-year */;

inline prd=1;

inline sump=0;

inline do j=i,13 /* loop for each recovery year */;

inline tmp=f(j)+s(14);

inline srv=exp(-tmp);

inline p(j-i+1,i)=prd*(1-srv)*f(j)/tmp;

inline sump=sump+p(j-i+1,i);

inline prd=prd*srv;

inline end do;

inline p(14-i+1,i)=1-sump /* compute last cell (not recovered) */;

inline end do;

cohort=648 ; 22 :; 24 :; 29 :; 21 :; 20 :; 9 :; 7 :; 1 :;

3 :; 0 :; 0 :; 1 :; 0 :;

cohort=34 ; 0 :; 0 :; 0 :; 2 :; 0 :; 0 :; 2 :; 0 :; 0 :; 1 :; 1 :; 0 :;

cohort=200 ; 4 :; 3 :; 7 :; 6 :; 3 :; 2 :; 0 :; 1 :; 0 :; 0 :; 0 :;

cohort=887 ; 24 :; 36 :; 35 :; 15 :; 10 :; 7 :; 2 :; 4 :; 3 :; 1 :;

cohort=892 ; 12 :; 23 :; 27 :; 11 :; 12 :; 9 :; 3 :; 2 :; 0 :;

cohort=349 ; 1 :; 9 :; 9 :; 8 :; 2 :; 0 :; 0 :; 0 :;

cohort=799 ; 11 :; 19 :; 22 :; 12 :; 3 :; 5 :; 5 :;


labels;

s(1)=f(1);

s(2)=f(2);

s(3)=f(3);

s(4)=f(4);

s(5)=f(5);

s(6)=f(6);

s(7)=f(7);

s(8)=f(8);

s(9)=f(9);

s(10)=f(10);

s(11)=f(11);

s(12)=f(12);

s(13)=f(13);

s(14)=m;
/* Now to estimate the model */

proc estimate novar maxfn=32001 name=base;



initial; all=.1; s(14)=.2;
proc stop;



1 This can be run online at: http://www.mbr-pwrc.usgs.gov/software/surviv.html.





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