Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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calathoides Motschulsky, 1844, Harpalus 374

calathoides Dejean, 1828, Platyderus [Feronia] 522

calceatus Stephens, 1835, Anisodactylus 360

calceatus Duftschmid, 1812, Harpalus [Carabus] 384

calciatii Della Beffa, 1931, Harpalus 386

calcitrapus Motschulsky, 1844, Harpalus 384

californica Motschulsky, 1859, Amara [Leirus] 561

californica Motschulsky, 1844, Pelophila 98

californicus Motschulsky, 1866, Carabus 168

caligata Putzeys, 1866, Amara [Curtonotus] 558

caligata Bates, 1889, Setolebia [Lebia] 430

caligatum Jeanne & Müller‑Motzfeld, 1982, Bembidion 260

caligatus Bates, 1873, Diplous 284



caliginosa Lutshnik, 1935, Amara 550

caliginosus Curtis, 1825, Acilius 48

caliginosus LeConte, 1850, Hydroporus [HN] 62

caliginosus Stephens, 1839, Hydroporus 64

caligosus Guignot, 1946, Pseuduvarus [Amarodytes] 57

calimanensis Knirsch, 1924, Duvaliopsis [Trechus] 298

calleides Bates, 1883, Paranchodemus [Anchomenus] 462

calleyi Fischer von Waldheim, 1823, Carabus 157

callichloris Bates, 1873, Chlaenius 354

callipeplus Schaufuss, 1882, Carabus 157


callipygius Cavazzuti, 1986, Carabus 185

callispilota Bates, 1892, Tachyura [Tachys] [DA] 277

callisthenoides Semenov, 1888, Carabus 123

callistoides Reitter, 1889, Mnuphorus [Hypercosmeton] 358

callitheres Bates, 1873, Synuchus [Dolichus] 545

callitrema Bates, 1889, Lebia 428

callosum Küster, 1847, Bembidion 252

callosus C. G. Thomson, 1884, Agabus [Gaurodytes] 39

callosus Semenov, 1897, Carabus 191

calosomoides Reitter, 1896, Carabus 169

calvitarsis Breit, 1912, Pterostichus 505

calycophora Schmidt‑Göbel, 1846, Lebia 430

calydonius P. Rossi, 1790, Ditomus [Scarites] 364



camarguensis Puel, 1937, Ophonus [Metophonus] 391

cambriensis Stephens, 1835, Hydroporus 63

camelina Lutshnik, 1933, Amara 552

cameranus Arribas, 1994, Zabrus 569

cameroni Baliani, 1934, Amara 566

cameroni Andrewes, 1922, Bembidion 267



cameroni Kult, 1954, Dyschirius 225

cameroni Morvan, 1997, Lepcha 459

cameroni Andrewes, 1925, Nebria 90

cameroni G. Ochs, 1925, Orectochilus 29

cameroni Jeannel, 1923, Perileptus 288

cameroni Louwerens, 1953, Skorlagad [Colpodes] 466

cameroni Jeannel, 1923, Trechus 328

camilloi Cavazzuti & Ratti, 1998, Carabus 128



caminarius Motschulsky, 1860, Hydroporus 62

camiolicus Schauberger, 1936, Trichotichnus 396

campania Andrewes, 1937, Pterostichus 515

campanulalus Bates, 1883, Dromius 421

campestris Fischer von Waldheim, 1820, Carabus 199

campestris M. F. Adams, 1817, Carabus 136

campestris Linné, 1758, Cicindela 103



campestris Sulzer, 1761, Cicindela 105

campestris Motschulsky, 1844, Patrobus 285

campicola Reitter, 1896, Carabus 199

canadense Hayward, 1897, Bembidion 259

canadensis Putzeys, 1866, Amara [Curtonotus] 560

canadensis Lapouge, 1908, Carabus 130

canadensis Fall, 1919, Hygrotus [Coelambus] 72

canalatus Uéno, 1980, Trechiama 322

canaliculatum O. G. Costa, 1839, Clinidium [Ips] 78

canaliculatus Nicolai, 1822, Acilius [Dytiscus] 48

canaliculatus Fischer von Waldheim, 1829, Broscus [Cephalotes] 236


canaliculatus M. F. Adams, 1812, Carabus 133

canaliculatus Kabak, 1997, Leistus 84

canaliculatus Lacordaire, 1835, Nebrioporus [Hydroporus] 67

canaliculatus Chaudoir, 1843, Platyderus [Lissotarsus] 522

canaricus Machado, 1992, Zabrus 570

canariense Harald Lindberg, 1950, Amaroschema [Amara] 521

canariense Machado, 1992, Parazuphium 573

canariensis Harold, 1868, Calathus 526

canariensis Dejean, 1831, Chlaenius 354

canariensis Bedel, 1881, Nebrioporus [Hydroporus] 67

canariensis Brullé, 1839, Orthomus [Feronia] 476

canariensis Machado, 1987, Spelaeovulcania 346

canariensis Machado, 1987, Stictonectes 69



canavesanus Capra, 1940, Pterostichus 513

cancellata Dejean, 1825, Lophyra [Cicindela] 112

cancellatus Illiger, 1798, Carabus 195

cancellatus Motschulsky, 1860, Pterostichus [Lyperopherus] 497

candei Chevrolat, 184, Lophyra [Cicindela] 112

candidum Andrewes, 1935, Cymbionotum 234

candidus Andrewes, 1942, Abacetus 347

candisatus Duftschmid, 1812, Carabus 166

canellipes Motschulsky, 1844, Agonum 453

canevae Gestro, 1885, Duvalius [Anophthalmus] 302

cangshanensis Deuve, 1996, Carabus 188

cangshanicola Farka, 1999, Leistus 80



canigouensis Schaefer, 1966, Carabus 140

canigoulensis Fairmaire & Laboulbène, 1854, Pseudomasoreus [Cymindis] 418

caniguensis Jeannel, 1937, Pterostichus [Haptoderus] 490

canis Sciaky & Pavesi, 1994, Trechus 328



canjuersicus J. Ochs, 1965, Carabus [Chrysocarabus] 140

cankiriana Korell & Kleinfeld, 1985, Cephalota 102

cankirianus Blumenthal & Breuning, 1967, Carabus 158

canoi Franz, 1981, Trechus 342

canonicus Casey, 1884, Harpalus 371

canovasae Toribio & Beltrán, 1993, Microtyphlus 239



cansiglianus Bernau, 1914, Carabus 187

cantaber Chaudoir, 1868, Pterostichus [Feronia] 512



cantaber Ganglbauer, 1915, Zabrus 569

cantabrica Bruneau de Miré, 1964, Nebria 86

cantabricus Piochard de la Brûlerie, 1868, Acupalpus [Stenolophus] 397

cantabricus Chevrolat, 1840, Carabus 167

cantabricus Sharp, 1882, Hydroporus 60

cantabricus Carabajal, García & Rodríguez, 1999, Hydrotrechus 312



cantabricus Jeanne, 1970, Platyderus 523

cantabricus Schaufuss, 1862, Pterostichus [Haptoderes] 490



cantabricus Schaufuss, 1862, Pterostichus 512

cantabricus Jeannel, 1927, Trechus 330

cantalicum Fauvel, 1882, Bembidion 252



cantalicus Jeanne, 1971, Carabus 130

cantalicus P. Machard, 1974, Carabus 130

cantalicus Pic, 1912, Platambus 44

cantalicus Chaudoir, 1868, Pterostichus [Feronia] 511

cantalicus Fauvel, 1888, Trechus 328

canteti Cabidoche, 1967, Geotrechus 310



cantonensis Born, 1922, Carabus [Apotomopterus] 126

cantonensis G. Hauser, 1918, Carabus [Apotomopterus] 124

caoduroi Pace, 1986, Duvalius 302

caoduroi Vigna Taglianti, 1982, Lessinodytes 314



capellae Depoli, 1929, Carabus 187

capensis Régimbart, 1894, Hydroglyphus [Bidessus] 56

capensis Sulzer, 1776, Lophyra [Cicindela] 112

capicola Péringuey, 1896, Acupalpus 398

capicola Péringuey, 1896, Drypta 358

capillatus Joseph, 1871, Anophthalmus [DA] 291

capillosa Ledoux & Roux, 1992, Nebria 94

capitalis Fairmaire, 1858, Paradromius [Dromius] 425

capitata Putzeys, 1866, Clivina 220

capitatus Chaudoir, 1854, Laemostenus [Pristonychus] 537

capitatus Chaudoir, 1850, Pterostichus [Feronia] 485

capitatus Geoffroy, 1785, Trechus [Buprestis] 339

capito J. Müller, 1918, Bembidion 270

capito Lewis, 1881, Carabus [Damaster] 148

capito Kryzhanovskij & Emetz, 1973, Cymindis 415

capito Audinet‑Serville, 1821, Dixus [Aristus] 365

capito A. Morawitz, 1862, Harpalus 384

capito Vereschagina & Kabak, 1997, Laemostenus 537

capito Tschitschérine, 1900, Pterostichus [Pseudoderus] 486



capito Chaudoir, 1855, Scarites 234

capitolinus Kurnakov, 1962, Pterostichus 504

caporiaccoi Netolitzky, 1935, Bembidion 244

cappadocica Franzen, 1996, Cephalota 102

cappadocicus Pawowski, 1976, Trechus 328

caprai Binaghi, 1939, Sphodropsis 543

caprai Jeannel, 1927, Trechus 328

capricornis Herbst, 1784, Noterus [Dytiscus] 34

captivorum Netolitzky, 1943, Bembidion 260



captus Blackburn, 1888, Polyderis [Tachys] 274

capucinus Géhin, 1885, Carabus 169

caraboides Sharp, 1882, Agabus 39

caraboides Schrank, 1781, Asaphidion [Cicindela] 242


caraboides Waltl, 1838, Carabus 187

caraboides Linné, 1758, Cychrus [Tenebrio] 19, 203

caraboides Motschulsky, 1839, Elaphropus 273

caraffai Sainte‑Claire Deville, 1906, Bembidion 255



caramaniae Peyron, 1858, Zabrus 571

caramanica J. R. Sahlberg, 1913, Tachyura [Tachys] [DA] 279

caramanus Fairmaire, 1886, Carabus 130

carantanus Depoli, 1925, Carabus 184

carantii Sella, 1874, Duvalius [Anophthalmus] 302



carbo Poppius, 1906, Pterostichus [Cryobius] 490

carbonaria Eschscholtz, 1829, Nebria 94

carbonarium Dejean, 1828, Agonum 449

carbonarius Fabricius, 1801, Agabus [Dytiscus] 39

carbonarius Ballion, 1878, Carabus 192

carbonarius Motschulsky, 1850, Carabus 199

carbonarius P. Rossi, 1790, Chlaenius [Carabus] 350

carbonarius Clark, 1863, Hydrovatus [Hydroporus] 71

carbonatus Schaufuss, 1882, Carabus 165

carbonicolor A. Morawitz, 1886, Carabus 192

carbonicolor Solsky, 1874, Poecilus 484

carceli Gory, 1833, Carabus 182

carchinii Vigna Taglianti, Magrini & Vanni, 1993, Duvalius 302

cardiaderum Solsky, 1874, Bembidion 244

cardiaderus Putzeys, 1872, Harpalus 376

cardioderus Fairmaire, 1889, Platynus [Colpodes] 464

cardioderus Chaudoir, 1850, Polyderis [Tachys] 274

cardioderus Chaudoir, 1846, Pterostichus [Feronia] 499

cardioderus Putzeys, 1870, Trechus 328

cardionota Putzeys, 1878, Amara 564

cardionotoides Hieke, 1988, Amara 564

cardionotum Putzeys, 1875, Bembidion 256

cardoni Fleutiaux, 1890, Calomera [Cicindela] 99

cardoni Severin, 1890, Cybister 50

cardoni Antoine, 1922, Harpalus 374

cardui Dejean, 1831, Amara 563

careniger Chaudoir, 1863, Carabus 133

caricum J. R. Sahlberg, 1908, Bembidion 257

caricus Kataev & Wrase, 1997, Harpalus 372

carinatus Duftschmid, 1812, Abax [Carabus] 469

carinatus Brullé, 1839, Calathus 526

carinatus Charpentier, 1825, Carabus 195

carinatus Motschulsky, 1839, Carabus 164

carinatus Chaudoir, 1859, Laemostenus [Sphodrus] 532

carinatus Aubé, 1838, Nebrioporus [Hydroporus] 66

cariniceps Putzeys, 1866, Coryza 221


cariniceps Baudi di Selve, 1864, Dyschirius 225

carinifer Andrewes, 1942, Abacetus 346

carinifrons Reitter, 1900, Coryza [Clivina] 221

carinifrons W. Horn, 1901, Neocollyris [Collyris] 116

carinigerum Korge, 1971, Bembidion 268

carinigerum Kasahara & Satô, 1997, Ischnagonum 458

carinthiacus Sturm, 1815, Carabus 179

carinthiacus J. Müller, 1916, Laemostenus 535

carinthiacus Mandl, 1940, Orotrechus 316

carinthiacus Meschnigg, 1946, Pterostichus 490

carinthianus Born, 1915, Carabus 179

carinulatus Chaudoir, 1869, Carabus 174

carinulatus Schaufuss, 1862, Laemostenus [Pristonychus] 532

carinulatus Fairmaire, 1859, Platytarus [Cymindis] 410

carlhlindrothi Kangas, 1980, Bembidion 262

carli Wewalka, 1992, Hydroporus 60



carlittensis Régimbart, 1901, Haliplus 32

carlittensis Jeanne, 1971, Trechus 339

carmelita Lapouge, 1907, Carabus 187

carmelitanus Mateu, 1982, Metadromius 421

carmenae Zaballos & Ruiz‑Tapiador, 1995, Typhlocharis 241

carminatii Dodero, 1917, Boldoriella [Trechus] 296

carnica J. Müller, 1924, Cymindis 412

carnicus Ganglbauer, 1902, Abax 469



carnicus Schweiger, 1969, Bradycellus 24, 400

carnicus Csiki, 1927, Carabus 163

carnicus Depoli, 1926, Carabus 138

carnifex Netolitzky, 1922, Bembidion 266

carnioliae J. Müller, 1921, Trechus 328

carniolicus Crotch, 1871, Carabus 163

carniolicus Géhin, 1885, Carabus 163

carniolicus Motschulsky, 1866, Cychrus 202

carniolicus Ganglbauer, 1891, Pterostichus 504

carniolicus Ganglbauer, 1897 (as 1896), Trechus 330

carolensis Aubry, 1974, Platyderus 522

caroli Roeschke, 1907, Carabus [Imaibius] 155

carolinus Fabricius, 1792, Carabus 140

carouxensis Schaefer, 1969, Carabus [Autocarabus] 195

carpathica Bielz, 1850, Nebria 85

carpathicum J. Müller, 1918, Bembidion 259

carpathicus Gistel, 1857, Abax 469

carpathicus Palliardi, 1825, Carabus 151

carpathicus L. Miller, 1868, Deltomerus [Patrobus] 282

carpathicus Kult, 1944, Pterostichus 503



carpathicus Rybiski, 1902, Trechus 328

carpathicus Schauberger, 1936, Trichotichnus 396

carpathus Born, 1902, Carabus 135

carpaticus Rybiski, 1902, Trechus 328

carpetana Zaballos, 1989, Typhlocharis 241

carpetanum Sharp, 1901, Bembidion 269

carradei Gautier des Cottes, 1866, Pterostichus [Feronia] [ND] 515

carrarae Jeannel, 1928, Duvalius 302



carrarai Sanfilippo, 1963, Hyphydrus 74

carrerei Coiffait, 1953, Aphoenops 294



carriana Casey, 1924, Amara 550

carrilloi Toribio & Rodríguez, 1997, Trechus 328



carsicola Depoli, 1929, Cychrus 202

carsticus Kasahara & Y. Itô, 1989, Pterostichus 507

cartereti Deuve, 1982, Carabus 183

carteroides Fairmaire, 1868, Graniger [Ophonus] 366

carteroides Zurcher, 1911, Ophonus [Harpalus] 389

carthusiana Jeannel, 1941, Oreonebria 96

carthusianus Sainte‑Claire Deville, 1902, Pterostichus [Platysma] 508

cartolarii Pomini, 1936, Duvalius 301

carvalhoi Serrano & Borges, 1986, Calathus 528

casalei Perrault, 1985, Amerizus 241

casalei F. Battoni, 1986, Calathus 524

casalei Cavazzuti, 1986, Carabus 163

casalei Cavazzuti, 1996, Cychrus 203

casalei Vigna Taglianti, 1969, Doderotrechus 298

casalei Sciaky, 1992, Duvalius 302

casalei Dubault & Lassalle, 1991, Laemostenus 534

casalei Straneo, 1984, Pterostichus 488

casaleianus Deuve, 1994, Carabus 124

casaleianus Kirschenhofer, 1991, Pterostichus 504



cascadiensis Hatch, 1949, Harpalus 378

caschmirensis L. Redtenbacher, 1844, Carabus 155

casellii Gestro, 1898, Duvalius [Anophthalmus] 302

caspia Ménétriés, 1832, Cicindela 107

caspia Fischer von Waldheim, 1821, Lophyra [Cicindela] 113

caspica J. R. Sahlberg, 1904, Megacephala 118

caspicus Dejean, 1831, Brachinus 214

caspicus Motschulsky, 1850, Chlaenius 351

caspicus Semenov, 1909, Eremosphodrus 532

caspiricus Deuve, 1982, Trechus 328



caspium Dejean, 1826, Calosoma 121

caspius Putzeys, 1866, Dyschirius 228

caspius Ménétriés, 1832, Gyrinus 27

caspius Steven, 1806, Harpalus [Carabus] 374

caspius Wehncke, 1875, Hygrotus [Hydroporus] 71


caspius Ménétriés, 1832, Laemostenus [Pristonychus] 536

caspius Casale, 1988, Pseudotaphoxenus 541

caspius Ménétriés, 1832, Pterostichus [Feronia] 498



caspius Kolenati, 1845, Tachys 21, 276

cassideus Fabricius, 1792, Licinus [Carabus] 442

cassolai Bonavita & Vigna Taglianti, 1993, Bembidion [Ocydromus] 245

casta Tschitschérine, 1898, Aristochroa 470



castalium Casey, 1918, Bembidion 262

castanea Putzeys, 1866, Amara [Curtonotus] 559



castanea Putzeys, 1861, Clivina 220

castanea Westwood, 1837, Clivina 219

castanea Klug, 1832, Merizomena [Cymindis] 409

castanea Bonelli, 1810, Oreonebria [Alpaeus] 95

castanea Andrewes, 1925, Tachyura [Tachys] 277

castaneipenne Motschulsky, 1844, Agonum 453

castaneipenne Jacquelin du Val, 1852, Bembidion 258



castaneipennis Ménétriés, 1832, Carabus 165

castaneipennis O. G. Costa, 1839, Carabus 130

castaneipennis Sciaky, 1987, Ophonus 389

castaneipennis Bates, 1873, Stenolophus 405

castaneipes Fairmaire, 1889, Pogonus 287



castaneus Gyllenhal, 1808, Agabus [Dytiscus] 39

castaneus Sharp, 1882, Agabus 39

castaneus Motschulsky, 1858, Apotomus 237

castaneus Heer, 1837, Deronectes [Hydroporus] 58

castaneus Aubé, 1838, Hydroporus 62

castanipes Gebler, 1847, Pterostichus [Platysma] 505

castanopterum Stephens, 1828, Bembidion [Notaphus] 248

castanopterus A. Villa & G. B. Villa, 1833, Carabus 180



castanopterus G. Hauser, 1913, Carabus [Isiocarabus] 156

castanopterus Heer, 1837, Trechus 337

casterini Capra, 1940, Pterostichus 513

castificus P. Moravec & Wrase, 1998, Trechus [Epaphius] 325

castiliana Hieke, 1983, Amara 554

castilianus Dejean, 1826, Carabus 167

castilianus Fery, 1995, Graptodytes 59

castilicum Netolitzky, 1918, Bembidion 264



castiliensis Born, 1924, Carabus 146

castiliensis Jedlika, 1950, Chlaenius 352



castillanus Coiffait, 1956, Laemostenus [Pristonychus] 539

castillianus Vuillefroy, 1866, Harpalus 376

castroi Martínez y Saez, 1873, Zabrus 569



catakensis P. Machard, 1985, Carabus 157

catalaunicus Géhin, 1885, Carabus 194

catalonicus Escolà & Cancio, 1983, Aphoenops 293



catalonicus Jeanne, 1988, Brachinus 213

catalonicus Ganglbauer, 1886, Carabus 168

catalonicus Jeanne, 1974, Licinus 442

catalonicus Jeanne, 1981, Poecilus 483

catalonicus K. Daniel & J. Daniel, 1906, Pterostichus 497



catalonicus Ganglbauer, 1915, Zabrus 569

cataphractus Cavazzuti, 1998, Carabus 188

catei Sciaky & Wrase, 1997, Pterostichus 501

catena Fabricius, 1775, Lophyra [Cicindela] 112

catenaria Brown, 1944, Blethisa 206

catenatus Panzer, 1803, Carabus 151

cateniger A. Morawitz, 1886, Carabus 190

catenulata Fischer von Waldheim, 1820, Nebria 89

catenulatus Scopoli, 1763, Carabus 151

cathaica Bates, 1874, Cylindera [Cicindela] 111

cathaica Sciaky & Pavesi, 1994, Nebria 89

cathaicus Sciaky, 1994, Pterostichus 514

cathaicus Bates, 1873, Synuchus [Pristodactyla] 545

cathaicus Sciaky & Pavesi, 1995, Trechus 328

catharinae Netolitzky, 1943, Bembidion 245



catharinae Schauberger, 1933, Ophonus [Harpalus] 391

catoirei Dejean, 1825, Pheropsophus [Brachinus] 217



caucasica Motschulsky, 1844, Amara [Leirus] 559

caucasica M. F. Adams, 1817, Calomera [Cicindela] 99



caucasica Motschulsky, 1844, Cicindela 105

caucasica Motschulsky, 1844, Lebia 428

caucasica Ménétriés, 1832, Nebria 85

caucasica A. Fleischer, 1921, Orientoreicheia [Reicheia] 222

caucasicola Lutshnik, 1935, Amara 556

caucasicola Netolitzky, 1918, Bembidion 261



caucasicola Lutshnik, 1915, Myriochila [Cicindela] 114

caucasicola Tschitschérine, 1893, Pterostichus [Feronia] 496


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