Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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caucasicola Tschitschérine, 1904, Trechus 339

caucasicum Motschulsky, 1844, Bembidion [Leja] 252



caucasicum Motschulsky, 1850, Bembidion [Peryphus] 261

caucasicus Chaudoir, 1846, Bradycellus [Acupalpus] 24, 400



caucasicus Géhin, 1884, Carabus [Macrogenius] 186

caucasicus M. F. Adams, 1817, Carabus 185



caucasicus Motschulsky, 1850, Carabus 131

caucasicus Motschulsky, 1859, Derostichus 441



caucasicus Semenov, 1900, Dromius 419

caucasicus Fedorenko, 1996, Dyschirius [Dyschiriodes] 230

caucasicus Zaitzev, 1927, Hydroporus 62

caucasicus Chaudoir, 1846, Laemostenus [Pristonychus] 538

caucasicus Chaudoir, 1867, Leistus 81



caucasicus Chaudoir, 1846, Lindrothius [Calathus] 529

caucasicus Motschulsky, 1845, Morion 443

caucasicus Dejean, 1831, Oedesis [Ditomus] 366



caucasicus Kolenati, 1845, Ophonus 390

caucasicus Zaitzev, 1927, Platambus 44

caucasicus Kryzhanovskij, 1968, Platyderus 522



caucasicus Motschulsky, 1850, Poecilus 484

caucasicus Ménétriés, 1832, Pterostichus [Feronia] 496



caucasicus Jedlika, 1952, Taphoxenus 544

caucasicus Chaudoir, 1846, Trechus 328



caucasicus C. Zimmermann, 1831, Zabrus 572

cauchoisi Colas, 1953, Laemostenus [Ceuthosphodrus] 533

caudangula Kabak, l997, Cymindis 418

caudicornis Kurnakov, 1963, Parapenetretus [Penetretus] 284

cauracum Schuler, 1969, Bembidion [Peryphanes] 258

caurina Andrewes, 1935, Hexagonia 407

caussicola Jeannel, 1919, Speotrechus [Trechus] 320

caussolensis J. Ochs, 1937, Laemostenus 532

caussolsicus Bellon, 1971, Carabus [Dysmictocarabus] 140

causticus Audinet‑Serville, 1821, Brachinus 215

caustomarginatus Imura & Mizusawa, 1994, Carabus 149



cautus Peyerimhoff, 1904, Duvalius [Trechus] 303

cautus Wollaston, 1854, Trechus 324



cauvini Sirguey, 1931, Carabus 139

cavalieri Colas, 1965, Carabus [Dysmictocarabus] 140

cavallensis Bernau, 1914, Carabus 180

cavallensis Jeannel, 1928, Orotrechus 316

cavallensis Jeannel, 1927, Trechus 335

cavazzutianus Casale, 1973, Carabus 193

cavazzutii Morvan, 1998, Andrewesius 456

cavazzutii Deuve, 1995, Epaphiopsis [Pseudepaphius] 309

cavazzutii Casale, 1983, Laemostenus 534

cavazzutii Farka & Sciaky, 1998, Leistus 82

cavazzutii Casale & Sciaky, 1994, Myas [Trigonognatha] 475

cavazzutii Casale & Giachino, 1989, Pontodytes 318

cavazzutii Casale & Sciaky, 1999, Pseudotaphoxenus 540

cavazzutii Morvan, 1998, Xestagonum 467

caveirensis Borges & Serrano, 1993, Cedrorum 471



cavernicola Kraatz, 1880, Carabus 182

cavernicola J. Müller, 1916, Molops 475

cavernicola J. Frivaldszky, 1881, Trechus 328

cavernosus I. Frivaldszky von Frivald, 1838, Carabus 181



caviceps Klug, 1834, Collyris 116

caviceps Jeannel, 1935, Kozlovites 313

cavicola Schaum, 1858, Laemostenus [Sphodrus] 534


cavicollis Straneo, 1985, Lesticus 471

cavicollis Sciaky & Pavesi, 1997, Pseudotaphoxenus 540

cavifrons Mandl, 1974, Carabus 155

cavifrons Grouvelle, 1914, Omoglymmius [Rhysodes] 78

cavifuga Ganglbauer, 1900, Duvalius [Trechus] 298

cavillatum De Monte, 1947, Bembidion 245

cavipennis Bates, 1873, Parena [Crossoglossa] 433

cayrosensis Giordan, 1985, Duvalius 305

cazorlensis Mateu, 1953, Laemostenus [Ceuthosphodrus] 534

cazorlensis Lagar Mascaró, Fresneda & Hernando, 1987, Nebrioporus [Potamonectes] 66

ceballosi Mateu, 1953, Trechus 328

cebennicus Balazuc & Bruneau de Miré, 1964, Anillus 238

cebennicus Guignot, 1932, Graptodytes 59

cebennicus Jeannel, 1937, Laemostenus [Ceuthosphodrus] 533

cecchiniae Jakobson, 1907, Pterostichus [Platysma] 504

cechenoides Reitter, 1885, Nebria 85

cecileae Beheim & Breuning, 1943, Carabus [Eucarabus] 128

cekirgensis Schweiger, 1964, Carabus 131

celata Ledoux & Roux, 1999, Nebria 89



celatus Clark, 1862, Hydroporus 62

celebensis J. Thomson, 1857, Catascopus 435

celebensis Schaufuss, 1885, Miscelus 437

celer Olivier, 1795, Acupalpus [Carabus] 399

celer Fabricius, 1792, Bembidion [Carabus] 251

celinae Genest, 1991, Arctaphaenops 295

celioides Baliani, 1934, Amara 552



celioides Ménétriés, 1848, Harpalus 376

cellarum M. F. Adams, 1817, Taphoxenus [Carabus] 544

cellinae J. Müller, 1963, Orotrechus 317

celox Schiødte, 1841, Gyrinus 27

celsus Semenov, 1898, Carabus 150

celtibericus Germar, 1824, Carabus 161

celtibericus Jeanne, 1969, Pterostichus [Haptoderus] 491

celtibericus Jeanne, 1970, Zabrus 569

celticus Lapouge, 1898, Carabus 195

cenisius Kraatz, 1878, Carabus 180

cenobitus Ledoux & Roux, 1999, Archastes 79

centenarius Knirsch, 1925, Duvalius [Trechus] 306

centralis Schauberger, 1929, Harpalus 370

centralis Mlyná, 1977, Molops 472

centralis Antoine, 1963, Singilis 438

centralis J. R. Sahlberg, 1900, Tachys 276

centralis Nonveiller, Pavievi & Popovi, 1994, Trechus 328

centrasiaticum Semenov & Znojko, 1929, Colpostoma 441

centriustatus Reitter, 1894, Tachys 275


centromaculatus Wollaston, 1864, Tachys [DA] 276

cephalaeus Descourtilz, 1827, Molops 474

cephalicus Csiki, 1927, Carabus 156

cephallenicus J. Müller, 1936, Molops 475



cephallonicus Maan, 1941, Leistus 83

cephalonica Winkler, 1911, Reicheadella [Reicheia] 222

cephalonicus Winkler, 1914, Trechus 328

cephalotellus Belousov, 1995, Trechus 328

cephalotes Reiche, 1861, Agabus 39

cephalotes Reitter, 1896, Amblystomus 360

cephalotes Knirsch, 1926, Aphaenopidius 293

cephalotes Dejean, 1826, Badister 440

cephalotes Linné, 1758, Broscus [Carabus] 236



cephalotes A. Nicolas, 1919, Carabus [Iniopachys] 156

cephalotes Sokolá, 1909, Carabus 184

cephalotes Sharp, 1882, Cybister 50

cephalotes Fairmaire & Laboulbène, 1854, Harpalus 370

cephalotes Motschulsky, 1861, Harpalus [Pseudoophonus] 384

cephalotes Dejean, 1826, Odotoncarus [Ditomus] 366

cephalotes Raffray, 1885, Paussus 210

cephalotes Coiffait, 1969, Pseudanillus 238



cephalotes Gautier des Cottes, 1870, Tapinopterus [Feronia] 519

cephalotes Jeannel, 1960, Trechus 328

cephalotes Putzeys, 1870, Trechus 331

cerberus Dieck, 1869, Aphoenops [Anophthalmus] 294



cerberus Dieck, 1870, Aphoenops [Anophthalmus] 294

cerberus Ishikawa, 1962, Carabus 159

cerberus Uéno, 1957, Ishikawatrechus 312

cerberus Ganglbauer, 1905, Taphoxenus [Sphodrus] 544



cercurus Schiødte, 1841, Gyrinus 28

cerdanica Aubry, 1970, Amara [Percosia] 556

cerdanus Lapouge, 1925, Carabus 155

cerdonicus Abeille de Perrin, 1903, Trichaphaenops [Trechus] 344

ceresae Schatzmayr, 1925, Stomis 518

ceresai Binaghi, 1938, Trechus 328

ceresiacus Born, 1896, Carabus [Orinocarabus] 180

ceresyi Aubé, 1838, Nebrioporus [Hydroporus] 67

cereus Lapouge, 1925, Carabus 169

cerisyi Dejean, 1826, Carabus 186



cerkes Basquin & Darge, 1986, Carabus 185

cernagorensis Apfelbeck, 1904, Carabus 198

cernisorensis Decou, 1962, Duvalius 300

cerovensis V. B. Guéorguiev, 1964, Pheggomisetes 318

cerrutii Sbordoni & Di Domenico, 1967, Duvalius 302

cerrutii Focarile, 1950, Trechus 341


certans Netolitzky, 1930, Bembidion 255

certus Reitter, 1896, Carabus [Tribax] 200

cervenkai Sciaky, 1994, Pterostichus 500

cervicalis Casey, 1924, Amara [Celia] 564

cervicis Motschulsky, 1844, Harpalus 378

cervicis Csiki, 1930, Pterostichus 491

cervini Dupuis, 1911, Agonum [Platynus] 451

cervini Stierlin, 1879, Amara 566

cespitis Kraatz, 1881, Cicindela 104

cevennicus E. Jacquet, 1886, Carabus 141

ceylanica Nietner, 1858, Tachyura [Bembidium] 278

ceylanicus Nietner, 1856, Hololeius [Chlaenius] 356

ceylonica Motschulsky, 1861, Batoscelis 400

ceylonica Motschulsky, 1859, Sericoda [Agonothorax] 466



ceylonicus Guignot, 1935, Hydaticus 54

ceylonicus Chaudoir, 1861, Miscelus 437

chabo Habu, 1955, Synuchus [Calathus] 545

chachalgensis Zamotajlov, 1994, Deltomerus 282

chadianus Cavazzuti & Ratti, 1999, Carabus 124



chaffanjoni Lapouge, 1930, Calosoma [Charmosta] 121

chaffanjoni Lesne, 1898, Carabus 172



chakaensis Mazzi, 1995, Carabus 183

chakassicus Obydov, 1995, Carabus 164



chakouri Reitter, 1909, Microlestes 424

chakrata Andrewes, 1935, Bembidion 271

chalcea Andrewes, 1925, Tachyura [Tachys] 278

chalcentus Bates, 1873, Harpalus 374

chalceola Bates, 1883, Nippononebria [Nebria] 95

chalceola Bates, 1873, Trigonotoma 520



chalceolus Chaudoir, 1869, Abacetus 347

chalceum Stephens, 1828, Bembidion [Tachypus] 251

chalceus Chaudoir, 1857, Brachyodes 444

chalceus Erichson, 1837, Dyschirius 225

chalceus G. Ochs, 1936, Orectochilus 29

chalceus Marsham, 1802, Pogonus [Carabus] 287



chalceus Bates, 1873, Stenolophus 405

chalceus Faldermann, 1836, Zabrus 569

chalcicola Franz, 1949, Nebria 86

chalciope Bates, 1891, Amara [Leiocnemis] 553

chalcites Dejean, 1828, Amara 548

chalcites Andrewes, 1936, Leistus 80

chalcocephalus Wiedemann, 1823, Pristomachaerus [Panagaeus] 356

chalcochlorus Chaudoir, 1852, Carabus 157

chalcodes Andrewes, 1935, Bembidion 250

chalcomum Bates, 1873, Agonum [Anchomenus] 454


chalconatus Mannerheim, 1830, Carabus 157

chalconatus Panzer, 1796, Ilybius [Dytiscus] 42

chalconotum Ménétriés, 1832, Agonum 449

chalcophaea Bates, 1873, Amara 567



chalcopterum Dejean, 1831, Bembidion 247

chalcostola J. Ochs, 1965, Carabus [Chrysocarabus] 140

chalcothorax Chaudoir, 1868, Lesticus [Triplogenius] 471



chalibaeum Gradl, 1881, Agonum [Platynus] 451

chalingensis Kleinfeld, 1997, Carabus 126

chalingxian Kleinfeld, 1998, Carabus 141

chalusi Jedlika, 1968, Zabrus [Harpalus] 572

chalybaeum J. R. Sahlberg, 1913, Bembidion 270

chalybaeus J. R. Sahlberg, 1903, Agabus 39

chalybaeus Motschulsky, 1864, Brachinus 214

chalybaeus Faldermann, 1836, Chilotomus [Ditomus] 364



chalybeatus C. G. Thomson, 1860, Ilybius 43

chalybeus Sturm, 1824, Badister 441

chalybeus Voet, 1778, Carabus [Buprestis] 136

chalybeus Putzeys, 1846, Dyschirius 225



chalybeus Dejean, 1828, Laemostenus [Pristonychus] 536

chalybeus Palliardi, 1825, Myas [Abax] 475

chambae Andrewes, 1934, Pristosia [Calathus] 530

chambouqensis Lassalle, 1998, Carabus 158

chambruni Breuning & Ruspoli, 1970, Carabus 193

chameleon Motschulsky, 1866, Pterostichus [Logarus] 486

champagnati Salgado Costas, 1991, Apoduvalius 295

champenoisi Croissandeau, 1893, Pterostichus 490

championi Andrewes, 1924, Asaphidion 242

championi Andrewes, 1923, Axonya 235



championi Bedel, 1907, Calodromius [Dromius] 419

championi Kult, 1951, Clivina 219

championi Andrewes, 1928, Cymindis 412

championi Morvan, 1992, Deltocolpodes 457

championi Andrewes, 1929, Dyschirius 228

championi Andrewes, 1920, Leistus 80

championi J. Balfour‑Browne, 1946, Microdytes 75

championi Jeannel, 1923, Neoblemus 288



championi Sharp, 1912, Ophonus 390

championi Andrewes, 1934, Pristosia [Calathus] 530

championi Andrewes, 1926, Pterostichus 510

championi Andrewes, 1925, Tachyura [Tachys] 278

championi Jeannel, 1920, Trechus 328

chamunensis Fedorenko, 1992, Dyschirius 226

chamzaabadensis Casale, 1988, Pseudotaphoxenus 541

chan Breuning, 1932, Carabus 145



changaica Jedlika, 1966, Cymindis 417

changaica Horvatovich, 1973, Nebria 18, 88

changaicus Lapouge, 1908, Carabus 199

changaicus Jedlika, 1968, Harpalus 383

changdensis Deuve, 1997, Carabus 125

changeonleei Ishikawa & Kim, 1983, Carabus [Damaster] 121

changi G. Hauser, 1913, Carabus [Apotomopterus] 126

changi Habu, 1978, Oxycentrus 392

changlingensis Li, 1992, Dyschirius 230

changyangensis Kleinfeld, 1998, Carabus 123

chaos Imura, 1993, Carabus 191



chaoyangensis Kanô, 1936, Cymindis 411

chappuisi Coiffait, 1955, Aphoenops 294

chappuisi Jeannel, 1926, Duvalius 299

characta Andrewes, 1925, Tachyura [Tachys] 278



charbinensis Breuning, 1961, Carabus 183

charbonieri Sirguey, 1931, Carabus 171

charchirensis Hieke, 1993, Amara 559

chareti Deuve, 1993, Carabus 146

chareti Deuve, 1994, Cychrus 203

chareti Lassalle, 1995, Omphreus 446

charetianus Auvray, 1992, Carabus 153

charillus Bates, 1883, Agonum [Anchomenus] 452



charini Lafer, 1988, Agabus [Eriglenus] 37

charis Andrewes, 1930, Amara 552

charis Bates, 1892, Stenolophus 405

charon Meggiolaro, 1964, Anophthalmus 22, 292

charon Dieck, 1869, Aphoenops [Anophthalmus] 294

charon Dieck, 1870, Aphoenops [Anophthalmus] 294

charon Andrewes, 1926, Bembidion 20, 271



charpentieri Sirguey, 1931, Carabus 139

chasanensis Lafer, 1989, Harpalus 374

chasautianus Vysoký, 1984, Pterostichus 496

chaslii Fairmaire, 1886, Nebria 93

chaudoiri Schaum, 1858, Amara 567

chaudoiri Gebler, 1847, Carabus 170



chaudoiri Ballion, 1871, Cicindela 105

chaudoiri Andrewes, 1919, Coptodera 435



chaudoiri Motschulsky, 1850, Harpalus 370

chaudoiri Alluaud, 1932, Lebia [Nematopeza] 429

chaudoiri Perrault, 1986, Leistus 82

chaudoiri Boheman, 1858, Mimocolliuris [Ophionea] 444

chaudoiri Arrow, 1901, Pheropsophus 217

chaudoiri Fairmaire, 1869, Pseudomasoreus [Cymindis] 418

chaudoiri Baillon, 1870, Scarites 234


chaudoiri Levrat, 1859, Trechus 340

chaudoirianus Jakobson, 1908, Brachinus 215



chaudoirianus Lapouge, 1909, Carabus 175

chaudoirianus Lutshnik, 1922, Harpalus 383

chaudoirianus Puel, 1938, Scarites 233

chaudoiriellus Deuve, 1995, Carabus 157

chaudoirii Wollaston, 1860, Apotomus 237

chaudoirii Chaudoir, 1850, Bembidion 266



chaudoirii Motschulsky, 1850, Bembidion [Peryphus] 261

chaudoirii Hochhuth, 1846, Cybister 51

chaudoirii Brisout de Barneville, 1867, Hydraphaenops [Anophthalmus] 311



chaudoirii Reiche & Saulcy, 1855, Pachycarus 367

chauveloti Breuning & Ruspoli, 1977, Carabus 186

chavesi Alluaud, 1919, Bradycellus 400

chechcirensis Lafer, 1979, Pterostichus 495

chefsuricus Reitter, 1896, Pterostichus 504

cheikensis Deuve, 1992, Carabus 158

cheikhermonensis Deuve, 1992, Carabus 157

chejuensis Deuve, 1991, Carabus 154

chelmosensis Maan, 1940, Molops 475

chelmosensis Maan, 1944, Nebria 87

cheloscelis Bates, 1873, Dyschirius 225

chelys Andrewes, 1921, Omophron 208

cheminorum Deuve, 1988, Carabus 161

chendae Schatzmayr, 1922, Orotrechus [Trechus] 316

chengjiangensis Huang, 1993, Harpalus 384

chengkouensis Imura, 1995, Carabus 159

cheni Deuve, 1992, Carabus 124

chenianus Deuve & Tian, 1999, Carabus 124

chenpengi Li, 1992, Pterostichus 515

chensicieni Kryzhanovskij, 1994, Xestagonum [Polychaetagonum] 467

chevrieri Fairmaire, 1888, Pseudotaphoxenus [Pristonychus] 540

chevrierii Heer, 1837, Harpalus 382

chevrierii Heer, 1837, Oreonebria [Nebria] 95

chevrolati Gaubil, 1846, Acupalpus [Stenolophus] 398

chevrolati Cristoforis & Jan, 1837, Carabus 186



chevrolatii Gautier des Cottes, 1867, Calathus 526

chevrolatii Laporte, 1833, Parazuphium [Zuphium] 573



chevsura Lutshnik, 1934, Amara 562

chiappai Sciaky, 1987, Anophthalmus 291

chiarae Monguzzi, 1982, Boldoriella 296

chiarae Monguzzi, 1993, Dalmataphaenops 23, 297

chiarae Magrini, Vanni & Degiovanni, 1997, Duvalius 304

chibi Habu, 1958, Agonum 452



chibi Satô, 1972, Laccophilus 77

chicioarae Jeannel, 1931, Duvalius 299

chikaichii Uéno, 1957, Trechiama 322

chikongshanensis G. Hauser, 1914, Carabus [Coptolabrus] 141

chilensis Gory, 1833, Laemostenus [Pristonychus] 536

chiloleuca Fischer von Waldheim, 1820, Cephalota [Cicindela] 102

chiloschizontus Huang, 1995, Harpalus 384

chinada Andrewes, 1921, Dioryche 369

chinadense Jedlika, 1940, Agonum [Colpodes] 452

chinadensis Jedlika, 1932, Harpalus 383

chinchiro Uéno, 1974, Kurasawatrechus 313

chindoensis Kwon & Park, 1989, Carabus [Damaster] 142

chinense Boheman, 1858, Agonum [Anchomenus] 454



chinense Jedlika, 1935, Agonum [Colpodes] 452

chinense Csiki, 1901, Bembidion 258

chinense Kirby, 1819, Calosoma 120

chinensis A. Zimmermann, 1919, Agabus [Gaurodytes] 36

chinensis Tschitschérine, 1894, Amara 553

chinensis Jedlika, 1933, Apristus 431

chinensis Chaudoir, 1850, Brachinus 215

chinensis Jedlika, 1953, Bradycellus [Tachycellus] 401

chinensis Jedlika, 1934, Calleida 408

chinensis Breuning, 1927, Carabus 156

chinensis Csiki, 1927, Carabus 124

chinensis Li, 1992, Carabus [Tomocarabus] 149

chinensis DeGeer, 1774, Cicindela 108



chinensis Régimbart, 1899, Copelatus 47

chinensis Jedlika, 1965, Craspedophorus 446

chinensis Jedlika, 1935, Cymindoidea 410

chinensis Jedlika, 1939, Daptus 369

chinensis Semenov, 1889, Dolichus 530

chinensis Jedlika, 1965, Dromius 421



chinensis Kult, 1949, Dyschirius 225

chinensis Bänninger, 1949, Eustra 208

chinensis Schauberger, 1932, Gnathaphanus 362

chinensis Falkenström, 1932, Haliplus 31

chinensis Sharp, 1882, Hygrotus [Coelambus] 71

chinensis Hlisnikovský, 1955, Hyphydrus 74

chinensis Csiki, 1901, Ilybius 42

chinensis Boheman, 1858, Laccophilus 76

chinensis Boheman, 1858, Lebia 428

chinensis Bates, 1872, Nebria 94

chinensis Jedlika, 1963, Odacantha [Colliuris] 444

chinensis Tschitschérine, 1895, Ophonus 391

chinensis Régimbart, 1892, Orectochilus 29

chinensis Jedlika, 1960, Orthotrichus 461


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