Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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csikii Jedlika, 1954, Platynus [Colpodes] 464

csikii Jedlika, 1953, Pseudotaphoxenus [Taphoxenus] 540

csikii Jedlika, 1968, Pterostichus 490

cuancao Uéno, 1991, Trechiama 322



cuatrecasasi Bolívar y Pieltain, 1940, Gomerina 521

cubicollis Hieke, 1981, Amara 552

cucujinus Reitter, 1892, Pterostichus 485

culminalis Cavazzuti, 1997, Cychrus 203



culminatus Bates, 1873, Chlaenius 353

culminicola Piochard de la Brûlerie, 1876, Bembidion 256

culminicola Monguzzi, 1982, Boldoriella 296

culminicola Cavazzuti, 1989, Carabus [Procerus] 185



culminicola Schauberger, 1921, Pterostichus [Lyperophorus] 508

cumanum Lutshnik, 1938, Bembidion 263

cumanus Fischer von Waldheim, 1823, Carabus 151

cumanus Lutshnik, 1921, Dyschirius 228

cumanus Lutshnik, 1916, Pogonus 286



cumatile Schiødte, 1841, Bembidion 266

cuneatus Régimbart, 1892, Orectochilus 29

cuniculina Dejean, 1831, Amara 563

cuniculinus Knirsch, 1929, Duvalius [Neoduvalius] 307



cuniculinus Duftschmid, 1812, Harpalus [Carabus] 377

cuniculorum Méquignon, 1921, Trechus 329

cunii Fairmaire, 1880, Ophonus 390

cuonaensis Yu, 1981, Lebia 428

cupidicornis Ishikawa & Kubota, 1994, Carabus 177

cupido Andrewes, 1935, Bembidion 263

cuprascens Bates, 1873, Apristus 431

cuprascens Andrewes, 1925, Asaphidion 242



cuprata Sturm, 1824, Sericoda [Agonum] 466

cupreata Jedlika, 1940, Pristosia [Laemostenopsis] 531

cupreatus Jedlika, 1940, Platynus [Colpodes] 464

cupreatus Schilsky, 1888, Pterostichus [Calopterus] 488

cupreoides Heer, 1837, Poecilus [Pterostichus] 484

cupreolum Solsky, 1874, Bembidion 266



cupreomicans Mandl, 1955, Carabus 145

cupreonitens Chevrolat, 1861, Carabus 139



cupreonitens Kraatz, 1878, Carabus 151

cupreonitens Viturat, 1901, Pterostichus 488

cupreothoracica Korell & Cassola, 1987, Cassolaia 101

cuprescens Motschulsky, 1860, Agonum [Agonothorax] 449

cuprescens K. W. Miller & Fery, 1995, Hydroporus 60

cuprescens Motschulsky, 1858, Myas [Trigonognatha] 475


cupreum Andrewes, 1925, Asaphidion 242

cupreum Motschulsky, 1844, Bembidion [Peryphus] 246

cupreum Dejean, 1826, Calosoma 119



cupreus Chaudoir, 1877, Carabus 168

cupreus G. Hauser, 1913, Carabus [Apotomopterus] 126

cupreus Oberthür 1935, Carabus 139

cupreus Schilsky, 1888, Carabus 160

cupreus Westhoff, 1881, Carabus 153

cupreus Krausse, 1911, Chlaenius 350

cupreus Duftschmid, 1812, Elaphrus 207

cupreus Dejean, 1829, Harpalus 374

cupreus Linné, 1758, Poecilus [Carabus] 482



cupreus Waltl, 1835, Syntomus [Dromius] 432

cupricollis Kraatz, 1886, Carabus [Eutelocarabus] 183

cupricollis Nietner, 1856, Chlaenius 351

cupricollis Pouillaude, 1914, Lesticus 471



cupricolor Nakane, 1963, Amara 556

cupriculus Reitter, 1896, Carabus 136

cuprifuscatus Deuve, 1995, Carabus 146

cuprinum Motschulsky, 1844, Agonum 452



cuprinus Fischer von Waldheim, 1820, Carabus 172

cuprinus Schilsky, 1888, Chlaenius 351

cupripenne Gemminger & Harold, 1868, Bembidion 246

cupripennis Germar, 1824, Notiobia [Poecilus] 363

cupripennis Fairmaire, 1852, Poecilus 481

curlettii Cavazzuti, 1984, Carabus 200

currax Wollaston, 1864, Nebria 85

currens Stephens, 1828, Ocys 273

currilis Normand, 1941, Anthracus 400

cursitans C. Zimmermann, 1832, Amara 556

cursitans Ledoux & Roux, 1998, Nebria 92



cursitor Stephens, 1828, Agonum 453

cursor Sturm, 1825, Amara 549

cursor Olivier, 1795, Anthia [Carabus] 347

cursor Fabricius, 1792, Bembidion [Carabus] [NO] 255

cursor O. F. Müller, 1776, Nebria [Carabus] 92

cursor Dejean, 1828, Pterostichus [Argutor] 486

cursor Uéno, 1999, Shenaphaenops 320

cursor Andrewes, 1930, Xestagonum [Anchomenus] 467

cursorius Dejean, 1828, Poecilus [Feronia] 482

curta Dejean, 1828, Amara 548



curta Andrewes, 1925, Tachyura [Tachys] 277

curtangulus Reitter, 1900, Dixus [Aristus] 365

curtatus Villers, 1789, Carabus 166

curtatus Mannerheim, 1853, Harpalus 379

curtatus Fairmaire, 1886, Pterostichus [ND] 515

curticollis Peyron, 1858, Poecilus [Feronia] 480

curticollis Fairmaire, 1866, Trechus 329

curticornis N. Ito, 1999, Trichotichnus 394

curtii J. Ochs, 1965, Carabus [Chrysocarabus] 140

curtii Giordan & Raffaldi, 1983, Duvalius 302

curtipennis Schauberger, 1936, Trichotichnus 394

curtoides Chaudoir, 1837, Zabrus 569

curtonotoides Kabak, 1997, Poecilus 484



curtula Motschulsky, 1844, Blethisa 206

curtulata Casey, 1924, Nebria 88

curtulum Jacquelin du Val, 1851, Bembidion 252

curtulus Fairmaire, 1856, Abax [Feronia] 470

curtulus Motschulsky, 1860, Bradycellus [Stenolophus] 401

curtulus Ganglbauer, 1891, Carabus 173

curtulus Motschulsky, 1850, Laemostenus [Pristonychus] 537

curtulus Ganglbauer, 1891, Molops 472

curtum Heyden, 1870, Asaphidion [Tachypus] 242

curtus Motschulsky, 1866, Gyrinus 27



curtus Régimbart, 1883, Gyrinus [HN] 27

curtus Wollaston, 1854, Orthomus [Argutor] 476

curtus Tschitschérine, 1906, Trichotichnus [Amaroschesis] 394

curtus Audinet‑Serville, 1821, Zabrus [Harpalus] 569



curtus Dejean, 1828, Zabrus 569

curvatilis Belousov & Kabak, 1998, Trechus 329



curvatus Motschulsky, 1845, Carabus 199

curvatus J. R. Sahlberg, 1880, Trechus 336

curvatus N. Ito, 1996, Trichotichnus 395



curvicrus C. G. Thomson, 1871, Amara 551

curvimana Wollaston, 1854, Tachyura 278

curvipes Tournier, 1864, Agonum [Anchomenus] 449

curvus Wrase, 1997, Psychristus 404



cuspidatus Heller, 1923, Callistoides 348

cuspidatus Kunze, 1818, Hydrovatus [Hyphydrus] 70

cuspidatus Uéno, 1985, Trechiama 322

custos Wollaston, 1854, Trechus 329

cvijici Jeannel, 1924, Duvalius [Duvalites] [DA] 307

cvrsnicensis Horvatovich, 1974, Nebria 85

cyanea Fabricius, 1787, Cicindela 108

cyanea Cristoforis, 1832, Cylindera [Cicindela] 109

cyanea Schaum, 1854, Trichisia [Isotarsus] 448



cyaneiceps Bates, 1873, Callistoides [Chlaenius] 348

cyanellus Chaudoir, 1848, Catascopus 435



cyanellus Kraatz, 1874, Harpalus 375

cyanellus Reiche & Saulcy, 1855, Poecilus [Feronia] 482

cyanellus Bates, 1889, Stenolophus 405

cyanelytrius Kraatz, 1889, Carabus [Coptolabrus] 144

cyaneofemoratus Born, 1910, Carabus [Coptolabrus] 142

cyaneolimbatus Kraatz, 1879, Carabus [Megadontus] 166

cyaneonitens Fairmaire, 1886, Chlaenius 351

cyaneosternum Mandl, 1954, Calosoma 119

cyaneoviolaceus G. Hauser, 1921, Carabus [Acoptolabrus] 122

cyaneoviolaceus Motschulsky, 1844, Carabus 170

cyaneoviridis Carret, 1897, Carabus 140

cyanescens Baliani, 1940, Amara 549

cyanescens Wesmael, 1835, Bembidion 245

cyanescens Motschulsky, 1859, Calosoma [Callisoma] 119



cyanescens Kraatz, 1882, Carabus 172

cyanescens Sturm, 1815, Carabus 168

cyanescens Poppius, 1906, Cicindela 104

cyanescens N. Ito & Wrase, 1998, Coleolissus 369

cyanescens Motschulsky, 1844, Harpalus 376

cyanescens Fairmaire, 1862, Laemostenus [Pristonychus] 539

cyanescens Hope, 1845, Loxoncus [Harpalus] 404

cyanescens Chaudoir, 1850, Pachycarus [Mystropterus] 367

cyanescens N. Ito, 1998, Trichotichnus 395

cyaneum Chaudoir, 1846, Bembidion 245

cyaneus Dejean, 1828, Anchomenus 455



cyaneus Stephens, 1828, Anisodactylus [Harpalus] 23, 361

cyaneus E. Bodemeyer, 1881, Carabus 151

cyaneus Fabricius, 1777, Carabus 138

cyaneus Brullé, 1835, Chlaenius 355



cyaneus Tschitschérine, 1893, Harpalus 383

cyaneus Ballion, 1878, Ophonus 388

cyaneus Dejean, 1830, Pachycarus [Ditomus] 367



cyaneus Gebler, 1841, Poecilus 482

cyaneus Gory, 1833, Poecilus 483

cyaneus Sciaky & Facchini, 1997, Xestopus 530



cyanicollis Gebler, 1841, Anchomenus 455

cyanicollis Dejean, 1826, Carabus 172

cyanicollis Brullé, 1837, Chlaenius 354

cyanicorne Pandellé, 1867, Asaphidion [Tachypus] 242



cyanidorsis Tschitschérine, 1890, Poecilus [Feronia] 484

cyanipennis Breuning, 1932, Carabus 124



cyanipennis Chaudoir, 1854, Catascopus 435

cyanipennis Schmidt‑Göbel, 1846, Euplynes 458



cyanipennis Eschscholtz, 1818, Laemostenus [Sphodrus] 539

cyanocephala Linné, 1758, Lebia [Carabus] 24, 427



cyanocephala Fabricius, 1798, Ophionea [Cicindela] 444

cyanocnemis C. G. Thomson, 1857, Amara 548

cyanoides Jeannel, 1942, Chlaenius 350

cyanoptera Chaudoir, 1873, Cymindis 415

cyanopterus G. Hauser, 1919, Carabus [Apotomopterus] 124

cyanopterus Tschitschérine, 1897, Harpalus [Ophonus] 375



cyanostola Lewis, 1882, Carabus [Damaster] 148

cyathigera P. Rossi, 1790, Lebia [Carabus] 429

cychricollis Lapouge, 1898, Carabus 140

cychriformis Bleuse, 1914, Carabus 168

cychrisans Lapouge, 1899, Carabus 161



cychrocephalus Fairmaire, 1858, Carabus 137

cychroides Baudi di Selve, 1864, Carabus 184

cychropalpus Peyron, 1858, Carabus 156

cycladicum Schatzmayr, 1940, Bembidion 253

cyclobregma Acciavatti & Pearson, 1989, Cylindera [Cicindela] 111

cycloderus Tschitschérine, 1903, Leistus 82

cycloderus Solsky, 1874, Liochirus [Ophonus] 366

cycloderus Chaudoir, 1880, Scarites 233

cycloderus Bates, 1873, Synuchus [Pristodactyla] 545

cycloderus Heyden, 1886, Synuchus 546

cyclodes Bates, 1892, Abacetus 347

cyclogonus Chaudoir, 1844, Harpalus 375

cycloides Régimbart, 1889, Hyphydrus 74

cyclopius Reitter, 1900, Harpalus 383

cyclopius Reitter, 1899, Scarites 233

cyclops Matsumura, 1910, Hexagonia [Calleida] 407

cyclops Crotch, 1871, Scarites 233

cycnus Busi & Rocca, 1983, Duvalius 301

cylindraceus Fairmaire, 1859, Acinopus 368

cylindraticollis Peyerimhoff, 1927, Paradromius [Dromius] 424



cylindrella Schiødte, 1837, Amara 568

cylindrica LeConte, 1878, Amara [Curtonotus] 561

cylindrica Motschulsky, 1844, Cymindis 415



cylindrica Schmidt‑Göbel, 1846, Neocollyris [Collyris] 117

cylindrica Semenov, 1889, Stenolepta 543

cylindricior Deuve, 1994, Carabus 157

cylindricollis Pini, 1871, Cychrus 203



cylindricollis Fabricius, 1798, Drypta [Cicindela] 358

cylindricus Lapouge, 1914, Carabus 157

cylindricus Sirguey, 1931, Carabus 171

cylindricus Dejean, 1825, Dyschirius [Clivina] 225

cylindricus Sciaky, 1994, Leistus 80

cylindricus Chaudoir, 1868, Percus 478

cylindricus Herbst, 1784, Pterostichus [Carabus] 488

cylindriformis W. Horn, 1912, Cylindera [Cicindela] 109

cylindriformis Uéno, 1977, Perileptus 289


cylindriformis Reitter, 1887, Pterostichus [Aphaonus] 485

cylindronotus Faldermann, 1836, Scarites 233



cymbalista Alluaud, 1923, Paussus 211

cymindiformis Reitter, 1901, Dicheirotrichus [Cardiostenus] 402

cymindoides Andrewes, 1933, Dromius 420

cymindoides Dejean, 1831, Orthotrichus [Anchomenus] 461

cymindulum Andrewes, 1930, Bembidion 20, 271

cymindulus Bates, 1892, Syntomus [Metabletus] 432

cyprensis Mandl, 1944, Cicindela 104

cyprensis Freude, 1988, Zabrus 572

cypricola Mandl, 1981, Calomera [Cicindela] 101

cypricus Roeschke, 1915, Carabus [Procrustes] 186

cyprinus Franz, 1987, Trechus 329



cyprioticus Mouchamps, 1956, Laccophilus 76

cyprium Maan, 1934, Asaphidion 242

cyprium De Monte, 1949, Bembidion 267

cyprium Jedlika, 1958, Bembidion 267

cyprius Régimbart, 1878, Hydroporus 60

cyprius Maan, 1941, Leistus 84

cyprius Wrase, 1999, Oedesis 366

cyprius Piochard de la Brûlerie, 1876, Platyderus 522

cyprius Maan, 1943, Tapinopterus [Pterostichus] 520

cyrenaica Baliani, 1928, Amara 564

cyrenaicus Koch, 1939, Harpalus 370

cyrenaicus Gridelli, 1930, Laemostenus 536

cyrenaicus Gridelli, 1930, Syntomus [Metabletus] 432

cyrili Lassalle, 1997, Laemostenus 534

cyrilli Jedlika, 1936, Speluncarius [Tapinopterus] 516

cyrtensis Reiche, 1872, Platyderus 522

cyrtophthalmus Tschitschérine, 1896, Poecilus [Feronia] 480

cyrtops Tschitschérine, 1906, Trichotichnus [Asmerinx] 395

cyrtus Glasunov, 1913, Mnuphorus 358

czadanicus Obydov, 1997, Carabus 170

czarensis Obydov, 1997, Carabus 134

czekanowskii Poppius, 1906, Pterostichus [Cryobius] 493

czerskii Zaitzev, 1953, Dytiscus 52

czipkai Heinz, 1980, Carabus 194

daanensis Deuve, 1999, Carabus 128

dabamontanus Imura, 1996, Carabus 124

dabanensis Deuve, 1998, Carabus 190

dabanensis Shilenkov, 1996, Carabus 172

dabanensis Shilenkov, 1982, Nebria 88

dabanicola Deuve, 1998, Carabus 190

dabashanensis Imura, 1995, Cychrus 203

dabashanus Imura, 1995, Carabus 129



dabieshanus Imura, 1996, Carabus 143

dabreui Andrewes, 1924, Xenodochus [Xenodus] 397

dacatrai Deuve, 1991, Carabus 133

dacatrai Deuve, 1992, Cychrus 203

dacatrai Ledoux & Roux, 1996, Nebria (as decatrai) 90

dacatrai Sciaky & Pavesi, 1994, Trechus 333

dacatraianus Deuve, 1995, Carabus 190

dacatraianus Deuve, 1996, Trechus 329

dachti Jedlika, 1968, Cymindis 412

dacicum Jeannel, 1941, Bembidion [Peryphus] 269

dacicus Csiki, 1906, Carabus 172

dacicus J. Frivaldszky, 1883, Duvalius [Trechus] 298

dacicus Sciaky, 1991, Philorhizus 425

dacoromanus Hormuzaki, 1903, Carabus 167

dadayanus Schweiger, 1964, Carabus 186

dafangensis Cavazzuti, 1994, Carabus 125

daffneri Broder, 1994, Anophthalmus 291

dagense Korge, 1971, Bembidion 264



dagestana Jedlika, 1962, Amara 555

dagestanum Jedlika, 1962, Bembidion 266

daghensis Jedlika, 1938, Pterostichus 496

daghestanicus Lapouge, 1925, Carabus 136

daghestanus Reitter, 1896, Pterostichus 504



dagliari Jedlika, 1962, Harpalus 382

dahanensis Coiffait, 1962, Laemostenus [Pristonychus] 538

dahli Preudhomme de Borre, 1879, Agonum [Anchomenus] 451

dahli Heer, 1841, Carabus 196

dahlii Dejean, 1831, Bembidion 267

dahlii Duftschmid, 1812, Nebria [Carabus] 85

dahud Morvan, 1994, Pristosia 531



dahuricus Fischer von Waldheim, 1828, Carabus 135

dahuricus Aubé, 1837, Colymbetes 45

daibosatsunis Kasahara, 1991, Trichotichnus 395

daihizanus Ishida, 1968, Pterostichus 501



daillei Pater, 1938, Ophonus 392

daimiella Bates, 1892, Pentagonica 448



daimiellus Bates, 1873, Dyschirius 224

daimio Bates, 1873, Agonum [Anchomenus] 449

daimio Bates, 1873, Cymindis 415

daisen Nakane, 1953, Carabus [Apotomopterus] 176

daisenicus Ishida, 1958, Pterostichus 507

daisenus Habu, 1973, Trichotichnus 395

daisetsuzanum Habu, 1958, Bembidion 265

daisetsuzanum Nakane, 1963, Xestagonum [Colpodes] 467



daisetsuzanus Kamiya, 1938, Agabus 36

daisetsuzanus Kôno, 1936, Carabus 134

daiyunshan Kleinfeld, 1998, Carabus 128



dalei Rylands, 1841, Amara 548

dalensis Rost, 1893, Carabus [Plectes] 132

daliangshanus Imura, 1998, Cychrus 206

dalijiashanica Hieke, 1997, Amara 554



dalila Antoine, 1943, Dyschirius 226

dallarmii Daffner, 1987, Orotrechus 316



dalmaticus Géhin, 1885, Carabus 187

dalmatina Dejean, 1828, Amara 564



dalmatina Schulz, 1909, Calomera [Cicindela] 100

dalmatinum Dejean, 1828, Agonum 452

dalmatinum Dejean, 1831, Bembidion 258

dalmatinus Duftschmid, 1812, Carabus 163

dalmatinus J. Müller, 1900, Haliplus 31



dalmatinus A. Zimmermann, 1915, Hygrotus 73

dalmatinus Dejean, 1828, Laemostenus [Pristonychus] 536



dalmatinus Chaudoir, 1843, Licinus 442

dalmatinus Dejean, 1828, Molops [Feronia] 472

dalmatinus L. Miller, 1861, Neotrechus [Anophthalmus] 315

dalmatinus L. Miller, 1881, Platyderus 522

dama P. Rossi, 1792, Carterus [Scarites] 363

damanabii Morvan, 1977, Psammodromius 426



damascena Reiche, 1876, Amara [Triaena] 568

damascenum Fairmaire, 1897, Parazuphium [Zuphium] 573

damascenus Lapouge, 1925, Carabus 157

damascenus Reiche & Saulcy, 1855, Zabrus 570



damasteroides Géhin, 1885, Carabus 144

damavandense Marggi & Huber, 1999, Bembidion 20, 269

damchungensis Deuve, 1998, Trechus 329

damhenvel Morvan, 1999, Lassalleianum 459

damone Holdhaus, 1924, Typhloreicheia [Reicheia] 223

damruz Morvan, 1999, Lagadstard 459

danae Kaláb, 1995, Carabus 175

danbaensis Kleinfeld, 1999, Carabus 162

dandongensis Kirschenhofer, 1997, Pterostichus 499

dandonis Kasahara, 1989, Pterostichus 494

dani Schweiger, 1970, Carabus 187



danicus Gmelin, 1790, Hydaticus [Dytiscus] 54

danielanus Semenov, 1904, Stomis 517



danieli Reitter, 1896, Carabus 180

danieli Reitter, 1900, Harpalus 375

danieli Reitter, 1905, Leistus 82

danieli Marcilhac, 1994, Myas [Trigonognatha] 475

danieli Reitter, 1897, Notiophilus 97



danieli Wagner, 1926, Pterostichus 504

danieli Holdhaus, 1902, Trechus 335



daniellae Bellon & Tarrier, 1972, Carabus [Eucarabus] 171

danielssoni Fery & Hosseinie, 1998, Deronectes 58



danihelkai Maan, 1936, Patrobus 285

danilevskii Obydov, 1997, Calosoma [Callisthenes] 118

danilevskii Obydov, 1993, Carabus 163

danilevskyi Kabak, 1993, Amara [Cribramara] 558

daochengensis Deuve, 1994, Carabus 175

daochengicus Deuve, 1989, Cychrus 203

daoublev Morvan, 1999, Xestagonum 467

daoulivek Morvan, 1998, Tostkar 467



daphnensis Korell, 1985, Carabus [Procerus] 186

daphnis Kurnakov, 1962, Carabus 131

darcangensis Deuve, 1995, Cychrus 204


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