Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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darcangicus Deuve, 1995, Carabus 135

dardanellicus Kraatz‑Koschlau, 1886, Carabus [Procerus] 185

dardazicus Deuve, 1990, Carabus 149

dardiellus Bates, 1889, Carabus 155

dardum Bates, 1889, Bembidion 268

darfurensis J. Balfour‑Browne, 1947, Yola 57

dargei Deuve, 1988, Carabus 156



darghariensis Dostal, 1986, Notiophilus 97

dariae Zamotajlov, 1994, Deltomerus 282

darjelingensis Putzeys, 1877, Amara 548

darjensis Jedlika, 1952, Taphoxenus 544

darlingtoni Uéno, 1955, Minypatrobus 284



darlingtoni Lindroth, 1956, Synuchus 546

darlingtonicum Jedlika, 1951, Bembidion 265

darvasicum Mikhailov, 1984, Bembidion 256

darvazensis Deuve, 1990, Carabus 148

darvazica Kabak, 1999, Cymindis 416

darvazicum Mikhailov, 1976, Colpostoma 441

darvazicus Kryzhanovskij, 1968, Pterostichus 486



dashdorzhi Jedlika, 1967, Neophygas [Harpalus] 387

dashtensis Heinz, 1970, Carabus 182

datianshanicus Kleinfeld, 1997, Carabus 124

daugavense Barševskis, 1993, Bembidion 261

daurica Motschulsky, 1844, Amara [Leirus] 559



daurica Motschulsky, 1844, Cylindera [Cicindela] 109

dauricum Motschulsky, 1844, Bembidion [Leja] 261

dauricum Motschulsky, 1844, Calosoma 121

dauricus Solsky, 1875, Carabus 136

dauricus Fedorenko, 1996, Dyschirius [Dyschiriodes] 227

dauricus Gebler, 1832, Dytiscus 52


dauricus A. Morawitz, 1862, Elaphrus 207

dauricus Motschulsky, 1860, Notiophilus 96

dauricus Motschulsky, 1860, Orectochilus 28

dauricus Motschulsky, 1860, Oreodytes [Hydrocoptus] 67

dauricus A. Zimmermann, 1924, Peltodytes 33

dauricus Fischer von Waldheim, 1823, Pseudotaphoxenus [Sphodrus] 540

dauricus Gebler, 1832, Pterostichus [Molops] 505

dauti Zamotajlov, 1989, Carabus 131

davaai Jedlika, 1968, Bembidion 246



davaicum Jedlika, 1968, Bembidion 262

davanensis Sciaky & Pavesi, 1994, Trechus 329

davatchii Morvan, 1975, Amara [Leiocnemis] 562

davatchii Morvan, 1971, Bembidion [Peryphus] 256



davatchii Morvan, 1971, Bembidion [Peryphidium] 20, 269

davatchii Morvan, 1982, Carabus 162

davatchii Morvan, 1975, Cymindis 412

davatchii Morvan, 1970, Deltomerus 282

davatchii Morvan, 1974, Nebria 90

davatchii Ledoux, 1975, Ocys 272

davatchii Morvan, 1981, Ophonus 390

davatchii Morvan, 1970, Platyderus 522

davatchii Morvan, 1973, Platyderus 523

davidi Tschitschérine, 1897, Amara 567

davidi Schuler, 1956, Bembidion [Actedium] 266

davidi Fairmaire, 1887, Cylindera [Cicindela] 109

davidi Vondel, 1991, Haliplus 31

davidi Tschitschérine, 1897, Harpalus 375

davidi Tschitschérine, 1897, Harpalus [Ophonus] 384

davidi Bedel, 1883, Hygrobia 35

davidi Naviaux, 1994, Neocollyris 116

davidi Chaudoir, 1878, Orthogonius 446

davidi Fairmaire, 1887, Panagaeus 447

davidi Tschitschérine, 1897, Pterostichus [Feronia] 515

davidi Tschitschérine, 1897, Trichotichnus [Amaroschesis] 394

davidiani Belousov, 1990, Trechus 329

davidianus Fairmaire, 1888, Broscus 236

davidianus Tschitschérine, 1903, Harpalus 375

davidioides Deuve & Yu, 1992, Carabus 124

davidis Lucas, 1882, Amphizoa 35

davidis Géhin, 1885, Calosoma 120

davidis H. Deyrolle, 1878, Carabus 124

davidis Fairmaire, 1886, Cychrus 204

davidis Fairmaire, 1891, Diplous [Patrobus] 283

davidis Fairmaire, 1889, Dolichus [Pristonychus] 530

davidis Liebherr, 1989, Paranchodemus 462



davidis Fairmaire, 1887, Paropisthius [Elaphrus] 98

davidis Fairmaire, 1887, Physodera 438

davidis Fairmaire, 1889, Platynus [Colpodes] 464

davidis Fairmaire, 1886, Platyrhopalus 211

davidsoni Mandl, 1986, Callistomimus 348

davidsoni Mandl, 1978, Carabus 162

davidsoni Mandl, 1978, Chlaenius 354

davidsoni Deuve, 1989, Perileptus 289

davisii G. R. Waterhouse, 1833, Notiophilus 97

davisii Curtis, 1831, Oreodytes [Hydroporus] 67

daweiensis Deuve, 1992, Carabus 125

dawuensis Deuve, 1992, Carabus 175

daxuense Toledano, 1998, Bembidion 251



dayaoshan Kleinfeld, 1996, Carabus 127

dayaoshanicus Imura, 1995, Carabus 126

dayongensis Kleinfeld, 1996, Carabus 156

dayoshan Kleinfeld, 1996, Carabus 124

dayremi Saulcy, 1906, Trechus 327

debarrosmachadoi Franciscolo, 1968, Hydaticus 53

debdouensis Obenberger, 1918, Harpalus 370

debduensis Antoine, 1941, Cymindis 413

debduensis Antoine, 1962, Poecilus 480

debilicostis Kraatz, 1878, Carabus 136

debilicostis Kraatz, 1897, Carabus [Chrysocarabus] 140

debilipes Régimbart, 1899, Agabus 39

debilis Semenov, 1897, Carabus 182

debilis Antoine, 1959, Carterus 364

debilis Coiffait, 1958, Geotrechus 310

debilis LeConte, 1878, Micromalthus 25

debilis Wollaston, 1864, Paranchus [Anchomenus] 462

debilis Kryzhanovskij & Mikhailov, 1971, Sirdenus 288

debilis Wollaston, 1871, Trechus 329

debilis Uéno, 1982, Yamautidius 345

debrunieri Briel, 1964, Harpalus [Ophonus] 386

decastriensis Lafer, 1979, Pterostichus 495



decebali Mallász, 1929, Carabus 139

decemlineatus Mannerheim, 1853, Hygrotus [Hydroporus] 72

decemmaculata Dokhtouroff, 1885, Cicindela 105

decemmaculata Baudrimont, 1923, Eurynebria 79

decempunctata Dejean, 1825, Cylindera [Cicindela] 111



decempustulata Ménétriés, 1848, Cicindela 104

decemstriata Dejean, 1831, Tachyura [Bembidium] 279

decemstriatus Roubal, 1948, Notiophilus 97

deceptrix Tschitschérine, 1898, Pterostichus [Feronia] 515



decipiens Fischer von Waldheim, 1828, Cephalota [Cicindela] 102

decipiens L. Dufour, 1820, Chlaenius [Harpalus] 352

decipiens Gautier des Cottes, 1872, Cicindela 107

decipiens Dejean, 1829, Harpalus 375

decipiens Sharp, 1878, Hydroporus 60

decipiens Waltl, 1835, Poecilus 481

declive C. A. Dohrn, 1884, Calosoma [Callisthenes] 118

declivis Semenov, 1889, Broscus 236

decolor Apfelbeck, 1911, Bembidion 263

decolor A. Morawitz, 1886, Calosoma [Callisthenes] 119

decolor Fischer von Waldheim, 1823, Carabus 198



decolor Fairmaire & Laboulbène, 1854, Harpalus 370

decolor Andrewes, 1925, Polyderis [Tachys] 274

decolor Jeannel, 1938, Trechus 329

decolorata W. Horn, 1907, Cylindera [Cicindela] 111

decolorata Chaudoir, 1850, Tachyura [Tachys] 279



decoloratus Kraatz, 1886, Carabus 171

decoloratus Reitter, 1884, Carabus 194

decoloratus Reitter, 1900, Harpalus 379

decora Fischer von Waldheim, 1829, Cymindis 412



decora Chaudoir, 1871, Lebia [Nematopeza] 429

decorata Guignot, 1956, Yola 57

decoratum Duftschmid, 1812, Bembidion [Elaphrus] 267

decoratus Gyllenhal, 1810, Hygrotus [Hyphydrus] 73

decorum Brullé, 1839, Bembidion 254

decorum Panzer, 1799, Bembidion [Carabus] 255



decorus Seidlitz, 1888, Carabus 198

decorus Klug, 1834, Hydaticus 53

decorus Bedel, 1913, Syntomus [Metabletus] 432

decorus Grimmer, 1841, Trechus 334

decoui Casale & Laneyrie, 1982, Duvalius 300



decurtatus Lapouge, 1922, Carabus 137

defanus Zamotajlov, 1988, Deltomerus 282



defecta Casey, 1918, Amara [Celia] 564

deficiens Casey, 1918, Amara [Curtonotus] 558

definita Casey, 1918, Amara [Celia] 564

deflexicollis Fairmaire, 1880, Zabrus 572

defossus Bates, 1883, Pterostichus 497

defreinai Kirschenhofer, 1981, Nebria 91

degeicola Deuve, 1996, Carabus 176



degener Semenov, 1899, Acinopus 368

degenerata Schaufuss, 1862, Nebria 93

degeneratus Géhin, 1885, Carabus 134

degeneratus Westhoff, 1882, Hydaticus 54

degeneratus Mäklin, 1881, Pterostichus [Feronia] 491

degensis Deuve, 1989, Carabus 190



degiovannii Magrini & Vanni, 1985, Duvalius 302

degorsi Croissandeau, 1893, Pterostichus [Lagarus] 486

degouvei Ganglbauer, 1909, Percus 478

dehaanii Chaudoir, 1848, Carabus 177

deharvengi Deuve, 1987, Stevensius 320

dehesicola Garcia‑Paris & Paris, 1996, Carabus [Rhabdotocarabus] 146

dehraduni Vazirani, 1969, Hyphoporus 73



dejeani Putzeys, 1866, Amara [Curtonotus] 560

dejeani Buquet, 1840, Anisodactylus 361

dejeani Heyden, 1891, Bembidion 267

dejeani Putzeys, 1846, Bembidion 248

dejeani Ganglbauer, 1891, Calathus 525



dejeani Fischer von Waldheim, 1822, Carabus 199

dejeani Géhin, 1885, Carabus [Procerus] [HN] 185

dejeani Brullé, 1832, Gyrinus 27



dejeani W. Horn, 1895, Neocollyris [Collyris] 117

dejeani Stichel, 1923, Pachycarus 367

dejeani Csiki, 1932, Parophonus [Trichotichnus] 393

dejeani Eschscholtz, 1829, Physodera 438

dejeani Jeanne, 1996, Platyderus 522

dejeani Rambur, 1837, Siagona 218



dejeani Rouget, 1882, Syntomus [Metabletus] 432

dejeani Putzeys, 1870, Trechus 327

dejeani Jakobson, 1907, Zabrus 572

dejeanii Fischer von Waldheim, 1823, Carabus [Procerus] 164

dejeanii Fischer von Waldheim, 1832, Cephalota [Cicindela] 102

dejeanii Dejean, 1831, Chlaenius [Epomis] 353



dejeanii Hope, 1831, Cicindela 108

dejeanii Dejean, 1826, Nebria 85



dejeanii Dejean, 1826, Pelophila 98

dejeanii Dejean, 1831, Percus [Feronia] 478



dejeanii Gistel, 1857, Pterostichus [Abaxodes] 489

dejeanii Aubé, 1838, Sandracottus [Hydaticus] 49



dekraatzi Kraatz, 1881, Carabus 174

dekraatzi Oberthür, 1883, Nebria 95

delagei Colas, 1949, Laemostenus [Ceuthosphodrus] 533

delamarei Decou, 1967, Duvalius 306



delamarei Dajoz, 1960, Laemostenus [Ceuthosphodrus] 533

delaroueriei Morvan, 1994, Pterostichus 510

delarouzeei Pandellé, 1867, Trechus 329

delarouzei Jacquelin du Val, 1857, Deronectes [Hydroporus] 58

delatorrei Uyttenboogaart, 1928, Asaphidion 21, 242

delavayi Morvan, 1997, Andrewesius 456

delavayi Fairmaire, 1886, Carabus 124

delavayi Fairmaire, 1886, Cylindera [Cicindela] 109



delavayi Fairmaire, 1888, Myas [Aurisma] 475

delavayi Fairmaire, 1887, Pristosia [Calathus] 531

delavayi Tschitschérine, 1897, Trichotichnus [Amaroschesis] 394

delbreili Genest, 1983, Aphoenops 294

delecolleae Giordan, 1989, Duvalius 304

delectabilis Ledoux & Roux, 1995, Nebria 89

delectus Wollaston, 1864, Graptodytes [Hydroporus] 59

delerei Fassati, 1957, Bembidion 268



delerei Mandl, 1955, Carabus 201

deletum Audinet‑Serville, 1821, Bembidion 259

deletus Schmidt‑Göbel, 1846, Orthogonius 446

deletus Régimbart, 1899, Platynectes 44



delhermi Mazéret, 1923, Peltodytes [Cnemidotus] 33

delhermi Saulcy, 1906, Trechus 23, 327

deliae Morvan, 1973, Bembidion [Nepha] 252

deliae Morvan, 1972, Carabus 161

deliae Morvan, 1999, Deliaesianum 457

deliae Morvan, 1970, Deltomerus 282

deliae Morvan, 1981, Laemostenus [Pristonychus] 538

deliae Morvan, 1977, Leistus [Leistophorus] 82

deliae Morvan, 1997, Lepcha 459

deliae Morvan, 1996, Platynus [Colpodes] 464

deliae Morvan, 1995, Skoeda 466

deliae Morvan, 1971, Trechus 329

delicatula Sciaky & Pavesi, 1994, Nebria 88

delicatulus Schaum, 1844, Bidessus [Hydroporus] 55



delicatulus LaFerté‑Sénectère, 1851, Chlaenius 352

delicatulus Coiffait, 1962, Hydraphaenops 311

delicatus Casey, 1918, Pterostichus [Cryobius] 490

deliciola Bates, 1889, Tachyura [Tachys] 279

deliciolus Bates, 1873, Callistoides [Chlaenius] 348

delictus Zaitzev, 1906, Dytiscus [Macrodytes] 52

delineata Ledoux & Roux, 1998, Nebria 92



dellabeffai Capra, 1940, Pterostichus 513

delphinensis Abeille de Perrin, 1869, Duvalius [Anophthalmus] 302



delphinensis K. Daniel & J. Daniel, 1890, Nebria 93

delprati Deuve, 1996, Cathaiaphaenops 297

deltshevi V. B. Guéorguiev, 1965, Duvalius 308

demangei Fleutiaux, 1902, Cylindera [Cicindela] 112

demauxi Coiffait, 1958, Geotrechus 310



demelti Heinz, 1977, Carabus 131

demelti Korge, 1962, Harpalus 375

demessus Andrewes, 1923, Microlestes 422

demetrii Semenov, 1903, Carabus 169

demidenkoae Dudko, 1999, Bembidion 264



demircapicus P. Moravec, 1986, Trechus 329

demissus Cavazzuti & Ratti, 1998, Carabus 141

demissus Jeannel, 1962, Trechus 329

demoflysi Normand, 1938, Laccophilus 76

demoflysi Normand, 1938, Laemostenus 534



demontei Ravizza, 1968, Bembidion 252

demulensis Zamotajlov & Sawada, 1996, Apatrobus 281

dendrobates Bedel, 1900, Paradromius [Dromius] 424

deneveui Fairmaire, 1859, Laemostenus [Sphodrus] 539

densaticollis Fairmaire, 1889, Cymindis 412

densatus Peyerimhoff, 1931, Stictonectes 69

densegranulatum Mandl, 1954, Calosoma 121

densepunctulatus Mandl, 1955, Carabus 201



denserugatum Géhin, 1885, Calosoma 119

densestriatus Csiki, 1927, Carabus 170

densestriolatus Mandl, 1965, Carabus 162

densicornis Fischhuber, 1977, Trechus [Epaphius] 325

dentata P. Rossi, 1790, Drypta [Carabus] 358

dentata Uéno, 1999, Kusumia 314

dentatus Zamotajlov & Sawada, 1996, Apatrobus 281

dentatus J. Schmidt & Morvan, 1999, Calathus 528

dentatus Stephens, 1828, Harpalus 23, 372

dentatus Belousov, 1998, Porocimmerites 318

dentatus Jeannel, 1947, Pterostichus [Haptoderus] 492

dentatus Andrewes, 1929, Scarites 234

denteletus B. V. Guéorguiev, 1997, Molops 474

dentelloides Netolitzky, 1943, Bembidion 250

dentellum Dejean, 1831, Bembidion 267

dentellum Thunberg, 1787, Bembidion [Carabus] 250



dentex Gistel, 1838, Cicindela 107

dentibasis Hieke, 1988, Amara 563

denticaudis Kurnakov, 1962, Pterostichus 496

denticolle Gebler, 1833, Calosoma 120

denticollis Reitter, 1887, Leistus 82

denticollis Jeannel, 1923, Perileptus 289

denticollis Schauberger, 1936, Trichotichnus 394

denticornis Donovan, 1804, Platyrhopalus [Paussus] 211

denticulata Holdhaus, 1924, Typhloreicheia [Reicheia] 223

denticulatus Wrase, 1995, Badister 440



denticulatus Lapouge, 1919, Carabus 192

denticulatus Chaudoir, 1880, Scarites 233

dentifer Uéno, 1978, Allotrechiama 290

dentifera Antoine, 1939, Cymindis 413

dentilabris Chaudoir, 1844, Myriochila [Cicindela] 114

dentipennis Motschulsky, 1850, Brachinus 215


dentipes Putzeys, 1866, Dyschirius 225

dentipes C. Zimmermann, 1831, Zabrus 569

deorum Jeanne, 1972, Carabus [Procrustes] 187

depilatus Bates, 1889, Pterostichus [Molops] 494

deplanatum A. Morawitz, 1862, Bembidion 270

deplanatus Guignot, 1952, Agabus 36

deplanatus Chaudoir, 1843, Calathus 527



deplanatus Fischer von Waldheim, 1823, Carabus [Plectes] 169

deplanatus Motschulsky, 1859, Carabus 187

deplanatus Gyllenhal, 1826, Hydroporus [Hyphydrus] 61

deplanatus Ménétriés, 1832, Poecilus [Feronia] 480

deplanatus Chaudoir, 1842, Pterostichus [Pseudomaseus] 511

deplanatus Motschulsky, 1850, Pterostichus [Pseudocryobius] [HN] 491

depolianus Bernau, 1913, Carabus 184

depressa C. Zimmermann, 1832, Amara 551

depressa Kult, 1951, Clivina 219

depressa Brullé, 1834, Heteracantha 387

depressa Heer, 1841, Oreonebria [Nebria] 95

depressangula Poppius, 1908, Amara 548

depressicollis Rosenhauer, 1856, Deronectes [Hydroporus] 58

depressidorsis Reitter, 1896, Pterostichus 504

depressipenis Sciaky & Pavesi, 1995, Trechus 329



depressipennis Mandl, 1955, Carabus 130

depressipennis Jedlika, 1953, Reflexisphodrus [Taphoxenus] 543

depressiusculum Motschulsky, 1850, Bembidion [Peryphus] 265

depressiusculus G. Ochs, 1940, Orectochilus 29

depressiusculus Chaudoir, 1844, Pterostichus [Pseudomaseus] 511

depressum Ménétriés, 1832, Bembidion 245



depressum Morvan, 1972, Bembidion [Peryphus] 271

depressum Jedlika, 1936, Xestagonum [Colpodes] 469

depressus Olivier, 1795, Abax [Carabus] 470

depressus Brullé, 1836, Calathus 526



depressus Gautier des Cottes, 1866, Calathus 527

depressus Bonelli, 1810, Carabus 184

depressus Brullé, 1832, Carterus [Ditomus] 364

depressus Eidam, 1925, Cychrus 203

depressus A. Fiori, 1896, Deltomerus 282

depressus Gebler, 1829, Diplous [Patrobus] 284

depressus Boheman, 1848, Distichus [Scarites] 231

depressus Duftschmid, 1812, Harpalus [Carabus] 375

depressus Breit, 1914, Leistus 83

depressus Paykull, 1790, Licinus [Carabus] 442

depressus Sturm, 1815, Licinus [HN] 442

depressus Fabricius, 1775, Nebrioporus [Dytiscus] 66



depressus Motschulsky, 1866, Percus 478

depressus Stephens, 1828, Perileptus [Lymnaeum] 288

depressus Audinet‑Serville, 1821, Platyderus [Harpalus] 522

depressus Dejean, 1831, Platynus 462

depressus Motschulsky, 1844, Pogonistes [Pogonus] 286

depressus Dejean, 1828, Pterostichus [Feronia] 495

depressus Meixner, 1922, Trechus 335

depressus N. Ito, 1996, Trichotichnus 395

deqinensis Xie & Yu, 1993, Aristochroa 471

dequaei Dupré, 1988, Geotrechus 310

dequensis Hieke, 1999, Amara 567

derasus Sharp, 1882, Laccophilus 77

derbendensis Jedlika, 1967, Tachyura [Tachys] 279

derelictum Alluaud, 1926, Bembidion 255

derelictus Dawson, 1854, Acupalpus [Stenolophus] 399

derelictus Clark, 1862, Hydroporus 61

dermatodes Fairmaire, 1868, Graniger [Ophonus] 366

derogatus Andrewes, 1929, Scarites 233

derongensis Deuve, 1994, Carabus 175

dertosensis Lagar Mascaró, 1964, Platyderus 522



desbordesi Maindron, 1899, Pheropsophus 218

desbordesi A. Gaudin, 1935, Trechus 340



desbrochersi Jeannel, 1942, Poecilus 482

descarpentriesi Morvan, 1975, Chlaenius 351

descendens Fischer von Waldheim, 1828, Cylindera [Cicindela] 109

descendens Bedel, 1902, Platyderus 521

descensus Schaufuss, 1871, Carabus 167

deserta Krynicky, 1832, Amara [Curtonotus] 559


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