Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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dives Lucas, 1846, Bembidion 270

dives Dejean, 1826, Chlaenius 352

dives Gory, 1833, Plutacia [Cicindela] 114

divisa Porta, 1923, Cicindela 105

divleo Morvan, 1995, Skoeda 466



divus Roeschke, 1908, Carabus [Coptolabrus] 141

djakonovi Semenov & Znojko, 1928, Nebria 89



djalrezi Coiffait, 1962, Laemostenus [Pristonychus] 538

djanaschvilii Ljovuschkin, 1970, Troglocimmerites 344

djavelidzei Retezár, 1992, Carabus 132

dlabolai Jedlika, 1968, Neophygas [Harpalus] 387


dobbertini J. Schmidt, 1998, Xestagonum 467

dobiaschi Beuthin, 1894, Carabus 151

dobretsbergeri Reitter, 1890, Calathus [Thermoscelis] 527

dobrogeanus Ienitea, 1962, Hydroporus 61



dobrudjensis Born, 1902, Carabus 187

dobzhanskii Semenov & Znojko, 1932, Carabus 145

dobzhanskyi Lutshnik, 1935, Amara 548

dochii Apfelbeck, 1906, Tapinopterus [Pterostichus] 519



docilis Casey, 1918, Amara [Celia] 564

dodensis Deuve, Lassalle & Quéinnec, 1985, Pristosia 531

doderoana Casale, 1985, Typhloreicheia 223

doderoi Gridelli, 1925, Acinopus 368

doderoi Baliani, 1926, Amara 563

doderoi Ganglbauer, 1891, Bembidion 267

doderoi Gestro, 1885, Duvalius [Anophthalmus] 303

doderoi Jeannel, 1920, Geotrechus 310

doderoi Schatzmayr, 1909, Molops 472

doderoi Bänninger, 1924, Nebria 87

doderoi Jeannel, 1947, Pterostichus [Haptoderus] 490

doderoi Balbi, 1892, Rhegmatobius [Microtyphlus] 240

doderoi Jeannel, 1927, Trechus 330

doderoi Holdhaus, 1924, Typhloreicheia [Reicheia] 223

doenitzi Harold, 1881, Broscosoma [Miscodera] 235



dofleini Csiki, 1927, Carabus 179

dohertyi Jordan, 1894, Tachys 275

dohertyi W. Horn, 1905, Therates 115

dohrni Gebler, 1847, Carabus 172

dohrni Kraatz, 1887, Carabus [Coptolabrus] 144

dohrnii Fairmaire, 1866, Anchomenus 455

dohrnii Fairmaire, 1863, Licinus 442

dohrnii Chaudoir, 1852, Trigonotoma 520

dokhtouroffi Ganglbauer, 1887, Carabus 148

dokhtourowi V. E. Jakovlev, 1884, Cylindera [Cicindela] 109

dolabratus Paykull, 1798, Colymbetes [Dytiscus] 45

dolens C. R. Sahlberg, 1827, Agonum [Harpalus] 450



dolens Kraatz, 1879, Carabus 195

dolens Putzeys, 1873, Clivina 219



dolens Fleutiaux, 1886, Cylindera [Cicindela] 109

dolens Tschitschérine, 1900, Pterostichus [Platysma] 510

dolicapax Ledoux & Roux, 1992, Nebria 88

dolichognathus Deuve, 1995, Carabus 144

dolichognathus Deuve, 1990, Cychrus 204

dolini Deuve, 1992, Carabus 159

dolini Kabak, 1992, Laemostenus 537

dolomitana K. Daniel & J. Daniel, 1898, Amara 563



dolomitanus Scholz, 1935, Agabus 39

dolomitanus Mandl, 1958, Carabus 180



dolomitanus Meggiolaro, 1964, Laemostenus [Antisphodrus] 535

dolomitanus Ganglbauer, 1900, Pterostichus 507

dolomitanus Jeannel, 1931, Trechus 330



dolomiticola Depoli, 1939, Abax 470

dolonicus Obydov, 1996, Carabus 159

dolonicus Belousov & Kabak, 1992, Trechus 337

dolops Jeannel, 1937, Duvalius 303



dolorosum Lindroth, 1963, Bembidion 262

dolorosum Motschulsky, 1850, Bembidion [Peryphus] 255



dolosum Puel, 1938, Agonum 449

dolosus Csiki, 1927, Carabus 195

dolpoensis J. Schmidt, 1999, Calathus 528



dominicae Ruspoli, 1976, Carabus [Eupachycechenus] 168

dominici Thomé, 1987, Carabus 158

dominici Brandmayr & Zetto Brandmayr, 1984, Oxypselaphus [Anchus] 462



dominus Reitter, 1885, Carabus 173

domonense Minowa, 1932, Asaphidion 242

donazzoloi Daffner, 1988, Orotrechus 317

donckieri Boileau, 1896, Carabus 124

donckieri Born, 1898, Carabus [Coptolabrus] 143

dongalensis Klug, 1832, Lophyridia [Cicindela] 113

dongba Zamotajlov, 1999, Naxipenetretus 284

dongchuanicus Deuve, 1994, Carabus 128

dongdaensis Deuve, 1996, Carabus 150

dongjugensis Deuve, 1997, Carabus 175

dongolaensis Hieke, 1997, Amara 563

doramasensis Uyttenboogaart, 1929, Pseudomyas 524

doriae Sharp, 1882, Deronectes 58

doriae Fairmaire, 1859, Duvalius [Anophthalmus] 303

doris Duftschmid, 1812, Bembidion [Elaphrus] 249

doris Panzer, 1796, Bembidion [Carabus] 270



doris Stephens, 1828, Bembidion [Lopha] 248

doris Jedlika, 1962, Pterostichus 486

dorjulensis Morvan, 1978, Pterostichus 510

dormeri W. Horn, 1898, Therates 116

dormeyeri Reitter, 1897, Bembidion 266

dorogostaiskianus Deuve & Imura, 1992, Carabus 134

dorogostaiskii Shilenkov, 1983, Carabus 149

dorsalis Fabricius, 1787, Acupalpus [Carabus] 399

dorsalis Pontoppidan, 1763, Anchomenus [Carabus] 455

dorsalis Dejean, 1831, Brachinus 215

dorsalis Fischer von Waldheim, 1820, Cymindis [Anomoeus] 413

dorsalis Gebler, 1829, Cymindis 412


dorsalis Dejean, 1829, Dicheirotrichus [Harpalus] 402

dorsalis Dejean, 1831, Drypta 358

dorsalis Gyllenhal, 1808, Gyrinus 27

dorsalis Dejean, 1826, Myriochila [Cicindela] 113

dorsalis Motschulsky, 1864, Stenolophus 406

dorsalis Fabricius, 1787, Suphrodytes [Dytiscus] 69

dorsata Brullé, 1834, Myriochila [Cicindela] 113



dorsatus Reitter, 1900, Daptus 369

dorsiger Duftschmid, 1812, Badister [Carabus] 441

dorsiger Aubé, 1838, Hydaticus 53

dorsiger Reitter, 1903, Trechus 339

dorsigera Schaum, 1863, Parena [Bothynoptera] 434

dorsistriatus A. Morawitz, 1862, Trechus 325

dorsolineata Schaum, 1860, Lophyra [Cicindela] 113

dorsolineolata Chevrolat, 1845, Lophyra [Cicindela] 113

dorsoplagiatus Fairmaire, 1880, Rhithrodytes [Hydroporus] 68

dorsostriatum Fairmaire, 1888, Agonum 452



dorsuarium Bedel, 1879, Bembidion 261

dostali Kirschenhofer, 1984, Bembidion 20, 259

dostali Donabauer, 1995, Neoblemus 288

dostali Casale, 1988, Pseudotaphoxenus 540

dostali Kirschenhofer, 1981, Pterostichus 507

dostojevskji Tschitschérine, 1895, Chlaenius 355

dostojewskii Tschitschérine, 1896, Cymindis 415

douei Peyron, 1858, Chlaenius 351

draco Bezina, 1999, Carabus 175

draco R. T. Bell, 1977, Yamatosa [Yamatoa] 78

draconis Deuve, 1990, Cychropsis 201

drannadaghensis Breuning, 1964, Carabus 193

drannadaghensis Breuning, 1967, Cychrus 202

dregei Aubé, 1838, Hydaticus 53

dremmi Morvan, 1995, Platynus [Colpodes] 464

drenskyi Breuning, 1928, Carabus 195

drescheri Fischer von Waldheim, 1817, Pterostichus [Harpalus] 509

drescoi Jeannel, 1953, Apoduvalius 295

drescoi Nègre, 1957, Pterostichus 497



dressleri A. Zimmermann, 1920, Rhantus 46

dreuxi Jedlika, 1968, Agonum 454



dreuxi Jedlika, 1968, Amara 548

dreuxi Deuve, 1990, Broscodera [Sinobrosculus] 235

dreuxi Deuve, 1994, Carabus 156

dreuxioides Deuve, 1998, Carabus 128



drewseni LeConte, 1862, Colymbetes 45

driolii Bognolo & M. Etonti, 1996, Anophthalmus 22, 291

drolmae Lassalle, 1999, Acalathus 529


dromicoides Chaudoir, 1852, Cylindera [Cicindela] 109

dromioides Jacquelin du Val, 1852, Bembidion 260

dromioides Reitter, 1897, Oroblemites [Trechus] 316

drouxi P. Machard, 1977, Carabus 158



drouxianus P. Machard, 1986, Carabus 188

droveniki Daffner, 1996, Anophthalmus 22, 291

droveniki Krätschmer, 1984, Carabus 163

droveniki Magrini, 1998, Duvalius 299

droveniki M. Etonti, 1995, Typhlotrechus 344

drurei Pic, 1904, Anthracus [Acupalpus] 400

dryas Andrewes, 1936, Brachinus 215

dryas Gistel, 1857, Carabus 165

dryops Bokor, 1913, Duvalius [Anophthalmus] 299

dshungarica Jedlika, 1967, Cymindis 411

dshungaricola Deuve, 1990, Carabus 150

dshungaricus Csiki, 1927, Carabus 174

dsungaricum Gebler, 1833, Calosoma 120

dsungaricus Stichel, 1923, Dixus [Ditomus] 365

dsychvensis Gottwald, 1985, Carabus 132

duarius Fischer von Waldheim, 1844, Carabus 136

dubaulti Lassalle, 1990, Laemostenus 532



dubia Motschulsky, 1844, Amara [NO] 549

dubia Ballion, 1878, Cymindis 415

dubia W. Horn, 1892, Myriochila [Cicindela] 114

dubia R. F. Sahlberg, 1844, Nebria 88

dubia Mateu, 1952, Tachyura 280

dubia Minowa, 1932, Tachyura [Tachys] [DA] 279

dubiolus Zamotajlov, 1992, Deltomerus 282

dubiosa Tschitschérine, 1894, Amara 562

dubiosus Poppius, 1905, Agabus 36

dubiosus Mallász, 1930, Duvalius 23, 299

dubiosus Tschitschérine, 1894, Pterostichus [Feronia] 490

dubiosus Andrewes, 1929, Scarites 233

dubisiensis Cabidoche, 1969, Trichaphaenops 344

dubitans Reitter, 1903, Trechus 330



dubium Heer, 1838, Bembidion 248

dubium Herbst, 1779, Omophron [Carabus] 208

dubius Cuvier, 1833, Abax [Carabus] 470

dubius Schilsky, 1888, Acupalpus 397



dubius Mannerheim, 1843, Agabus 41

dubius Ballion, 1871, Brachinus 213

dubius Motschulsky, 1844, Carabus 172

dubius Hoppe, 1796, Chlaenius [Carabus] 351

dubius Chaudoir, 1846, Deltomerus [Cardiomera] 282



dubius Audinet‑Serville, 1821 (as 1830), Dytiscus [HN] 51

dubius Gyllenhal, 1827, Dytiscus 51

dubius Mateu, 1981, Microdaccus 422

dubius Aubé, 1838, Nebrioporus [Hydroporus] 67

dubius Wollaston, 1857, Ocys [Bembidion] 272

dubius Desbrochers des Loges, 1904, Philorhizus [Dromius] 425

dubius Gautier des Cottes, 1869, Pterostichus [Omaseus] [HN] 499

dubius Heer, 1838, Pterostichus 503

dubius Melsheimer, 1844, Suphrodytes [Hydroporus] 69

dublanskii Belousov, 1998, Taurocimmerites 321

ducalis Schaufuss, 1882, Carabus 157

ducalis Chaudoir, 1876, Chlaenius 349

dudichi Csiki, 1928, Bembidion 256

dudichi Csiki, 1927, Carabus 151

dudkorum Belousov & Kabak, 1996, Trechus 330



duerckianus Ganglbauer, 1890, Carabus 179

duerensis Raynaud, 1974, Carabus 179

dufouranus Csiki, 1927, Carabus 194

dufouri Dejean, 1829, Carabus 167

dufouri Chaudoir, 1869, Cychrus 204

dufourii Perris, 1864, Bembidion 266

dufourii Dejean, 1828, Pterostichus [Feronia] 498

duftschmidi J. Schmidt, 1994, Agonum 450



duftschmidi Géhin, 1876, Carabus 196

duftschmidi Lapouge, 1922, Carabus 166

duftschmidii Sturm, 1818, Harpalus 382

duftschmidti Rye, 1872, Hydroporus 64

dulat Belousov & Kabak, 1992, Trechus 330

dulcigradus Bates, 1873, Synuchus [Pristodactyla] 545

dulcis Bates, 1883, Pterostichus [Lagarus] 486

dulcis Andrewes, 1925, Tachyura [Tachys] 277

duluchus Morvan, 1992, Deltocolpodes 457

dumbirensis Pulpán, 1957, Nebria 87

dumetorum Faldermann, 1836, Cicindela 105

dumetorum Motschulsky, 1840, Cicindela 103

dumitrescui Decou, 1959, Trechus 330

dumonti Español, 1977, Geotrechus 309

dundai Sciaky, 1994, Pterostichus 500

duodecimguttata Bonelli, 1813, Anthia 347

duodecimmaculatus Régimbart, 1877, Stictotarsus [Hydroporus] 69

duodecimpunctatum O. F. Müller, 1776, Agonum [Carabus] 451

duodecimpustulatus Fabricius, 1792, Stictotarsus [Dytiscus] 69



dupeuxi Deuve, 1974, Carabus 171

duplanatio Cavazzuti & Ratti, 1999, Carabus 124

duplex Bates, 1883, Lebia 429

duplicatus Uéno, 1986, Trechiama 322



duponcheli Barthélemy, 1837, Carabus 157

duponcheli Dejean, 1831, Carabus 185

duponchelii Dejean, 1831, Tapinopterus [Feronia] 519

duponti Lebis, 1949, Carabus [Chrysocarabus] 138

duponti Dejean, 1826, Cosmodela [Cicindela] 108



dura Casey, 1924, Amara [Curtonotus] 561

durani Raynaud, 1974, Carabus 130

duratii A. Villa & G. B. Villa, 1835, Pterostichus 503

dureli Maindron, 1899, Chlaenius 354

durescans Casey, 1924, Harpalus 377

duripshensis Kurnakov, 1972, Carabus 132

durmitorensis Apfelbeck, 1904, Carabus 163

durmitorensis Apfelbeck, 1904, Duvalius [Trechus] 299

durmitorensis Drovenik, 1984, Laemostenus 534

durmitorensis J. Müller, 1917, Molops 474

durmitorensis Apfelbeck, 1904, Nebria 86

durmitorensis Nonveiller & Pavievi, 1987, Winklerites 240

durmitorus Mandl, 1985, Carabus 165

durudense Marggi & Huber, 1999, Bembidion 255

durus Reitter, 1896, Carabus 196

durvillei A. Morawitz, 1889, Carabus 157

duvali Bedel, 1893, Bembidion 270

duvalii Dejean, 1828, Pterostichus 488

duvergeri Gobert, 1874, Acilius 49

dux Tschitschérine, 1894, Amara 559

dux Andrewes, 1935, Bembidion 268



dux Semenov, 1898, Carabus 143

dux Schaufuss, 1862, Pterostichus 512

dvoraki P. Moravec, 1986, Duvalius 303

dvorshaki Casale & Vereschagina, 1986, Eremosphodrus 532



dybowskii J. L. omnicki, 1903, Notiophilus 96

dyrrhachii De Monte, 1947, Bembidion 254

dyscheres Netolitzky, 1943, Bembidion 256



dyschirius Bonelli, 1813, Dyschirius [Clivina] 228

dyseimatus Cavazzuti, 1995, Carabus 156



dzambuli Jedlika, 1966, Amara 562

dzambuli Jedlika, 1966, Harpalus 379

dzermukensis Iablokoff‑Khnzorian, 1963, Trechus 330



dzhajloensis Deuve, 1991, Carabus 123

dzhalair Belousov & Kabak, 1994, Trechus 330



dzhambazishvili Kryzhanovskij, 1968, Pterostichus 496

dzhebaglicus Gottwald, 1989, Carabus 146

dzhentui Zamotajlov, 1997, Carabus 137

dzhergalanicus Belousov & Kabak, 1994, Trechus 342

dzhugensis Zamotajlov, 1988, Carabus 132


dzhungarica Kryzhanovskij, 1974, Amara [Curtonotus] 559

dzhungaricus Belousov & Kabak, 1992, Trechus 330

dzosonicus Pawowski, 1974, Tachys 275

dzykhvensis Belousov, 1990, Trechus 330

earinum Obydov & Pütz, 1996, Calosoma [Callisthenes] 118

ebenina Nietner, 1858, Tachyura [Bembidium] 279

ebeninus Heyden, 1870, Harpalus 376

ebenus Quensel, 1806, Pterostichus [Carabus] 514

eberlovi O. Berlov, 1996, Harpalus 376



eberti Jedlika, 1965, Amara 564

eberti Jedlika, 1965, Broscus 236



eberti Mandl, 1965, Carabus 162

eberti Mandl, 1965, Cylindera [Cicindela] 110

eberti Jedlika, 1965, Dicranoncus [Agonum] 458

eberti Jedlika, 1966, Harpalus 385

eberti Jedlika, 1965, Lebia 428

eberti G. Ochs, 1966, Orectochilus 29

eberti Jedlika, 1965, Orthotrichus 461

eberti Jedlika, 1965, Peliocypas [Risophilus] 434

eberti Jedlika, 1965, Platynus [Colpodes] 464

eberti Jedlika, 1965, Syntomus [Metabletus] 432

ebneri Netolitzky, 1929, Bembidion 252

ecchelii Bertolini, 1887, Abax 469

eccoptopterus Kraatz, 1894, Carabus [Apotomopterus] 124

echarouxi Lassalle, 1996, Carabus 147

echigo Ishikawa & Takami, 1996, Carabus 176

echigonis Uéno, 1972, Trechiama 322

echigonum Habu & Baba, 1957, Bembidion 255

echigonus Habu & Baba, 1962, Apatrobus [Patrobus] 280

echigonus Habu & Baba, 1960, Oodes 445

echinatus Régimbart, 1895, Synchortus 34

eckerleini Heinz, 1970, Carabus 182



edai Jedlika, 1952, Trechus 325

edai Jedlika, 1952, Trichotichnus [Velmius] 395



edai Jedlika, 1952, Xestagonum [Colpodes] 469

edax Andrewes, 1934, Pristosia 531

edentatus Uéno, 1982, Yamautidius 345

edithae Reitter, 1893, Carabus 131

edithae Janák & P. Moravec, 1989, Duvalius 299

edithae Reitter, 1914, Lebia 428

edmondsi Moore, 1956, Tachys [DA] 275



edmontonense Casey, 1924, Bembidion 248

edmundi Semenov, 1897, Carabus 200

edurus Dejean, 1828, Molops [Feronia] 475

edwardsi Sharp, 1914, Gyrinus 27


efflans Dejean, 1830, Brachinus (as etslans) 213

efflatouni Schatzmayr & Koch, 1934, Porotachys [Tachys] [DA] 274

effluviorum Peyron, 1858, Bembidion 267

effrenus Lutshnik, 1930, Poecilus [Platysma] 484



egesippei LaFerté‑Sénectère, 1847, Carabus 167

egonis J. Müller, 1923, Anophthalmus [Trechus] 291



egorovi Lafer, 1978, Amara 567

egorovi Lafer, 1989, Harpalus 376

egorovi Lafer, 1973, Mastax 217

egorovi Belousov & Kabak, 1996, Trechus 330



egregius Casey, 1914, Harpalus 378

egregius Jeannel, 1927, Trechus 330

egrissicus Belousov, 1992, Carabus 169

ehdenensis Deuve, 1992, Carabus 158

ehikoense Habu, 1984, Bembidion 271

ehikoense Habu, 1954, Negreum [Agonum] 460

ehlersi Putzeys, 1872, Cymindis 412

ehlersi Abeille de Perrin, 1872, Hydraphaenops [Anophthalmus] 311

ehlersi Heyden, 1881, Pterostichus [Haptoderus] 490

ehrenbergi Fischer von Waldheim, 1829, Carabus 154

ehrenbergii Klug, 1832, Carabus 188

eichhoffii Bach, 1851, Bembidion 266

eichhoffii Kraatz, 1888, Bembidion [Pseudolimnaeum] 267

eichingeri Jedlika, 1965, Harpalus 371

eichingeri Jedlika, 1965, Harpalus [Xenodus] 377

eichingeri Jedlika, 1965, Pterostichus 513

eichwaldi Fischer von Waldheim, 1828, Carabus 192

eichwaldianus Géhin, 1885, Carabus 192

eidami Wagner, 1932, Chlaenius 352

eikoae Morita, 1997, Trichotichnus 395

eiregi Jedlika, 1964, Pseudotaphoxenus [Taphoxenus] 542

eiselti Mandl, 1967, Cephalota 102

ejaculans Fischer von Waldheim, 1828, Brachinus 213

elainus Bates, 1883, Diacanthostylus [Colpodes] 458

elatum Andrewes, 1930, Bembidion 265

elatus Fabricius, 1801, Molops [Carabus] [NP] 472

elbursensis Breuning, 1946, Carabus 169

elbursensis Morvan, 1974, Ocys 272

elbursiaca B. Bodemeyer, 1927, Nebria 86

elbursiacum Morvan, 1973, Bembidion [Trepanedoris] 248

elbursianus Mandl, 1958, Carabus 185

elbursicum Netolitzky, 1939, Bembidion 247

elburzensis Morvan, 1981, Deltomerus 282

elburzensis Morvan, 1981, Laemostenus [Cryptoxenus] 537

elburzensis Morvan, 1977, Olisthopus [Odontonyx] 461



elburzensis Morvan, 1977, Synuchus 545

elburzensis Morvan, 1974, Trechus 330



elburziacum Morvan, 1971, Bembidion [Bembidionetolitzkya] 246

eldorensis Casey, 1918, Amara [Celia] 564

elegans Dejean, 1829, Acupalpus [Stenolophus] 398



elegans Rylands, 1841, Amara 549

elegans Germar, 1824, Bembidion 247


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