Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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aeruginosus Chaudoir, 1856, Chlaenius 354

aeruginosus Stephens, 1828, Pogonus 287

aerumnosa Lutshnik, 1933, Amara 557

aesculanus Pantel, 1888, Harpalus 371

aesculapius Imura & Kezuka, 1989, Carabus [Protodamaster] 188

aesculoides Jedlika, 1959, Ophonus [Harpalus] 391

aestivus Duftschmid, 1812, Harpalus [Carabus] 379

aestuans Dejean, 1826, Notiophilus 96

aestuarii Uéno & Habu, 1954, Cillenus [Armatocillenus] 272

aethiopica Kult, 1959, Clivina 219

aethiopicus Bänninger, 1933, Scarites 232

aethiops Cristoforis & Jan, 1837, Carabus 182

aethiops Stephens, 1828, Nebria [Helobia] 88

aethiops Panzer, 1796, Pterostichus [Carabus] 493

aetolica Apfelbeck, 1901, Nebria 84

aetolicum Apfelbeck, 1901, Bembidion 254



aetolicus Schaum, 1857, Carabus 136

aetolicus Apfelbeck, 1904, Omphreus 446

aetolicus Apfelbeck, 1901, Platyderus 523

aetolicus Ganglbauer, 1889, Tapinopterus 520

aetolus Schaum, 1864, Zabrus 570

aezcoaensis Dupré, 1988, Aphoenops 293

afer Coquerel, 1859, Leistus 84

affine Stephens, 1828, Agonum 453

affine L. Redtenbacher, 1849, Bembidion 245

affine Stephens, 1835, Bembidion [Peryphus] 259

affinis Paykull, 1798, Agabus [Dytiscus] 38

affinis Dejean, 1828, Amara 554

affinis Motschulsky, 1844, Amara 568

affinis Motschulsky, 1850, Amara 556

affinis Baudi di Selve, 1871, Binaghites [Scotodipnus] 238

affinis F. Battoni, 1986, Calathus 526

affinis Duftschmid, 1812, Carabus 196

affinis Gistel, 1857, Carabus 196

affinis Kwon & Lee, 1984, Carabus [Tomocarabus] 149


affinis Panzer, 1809, Carabus 171

affinis Rambur, 1837, Carterus 364

affinis Fischer von Waldheim, 1820, Cicindela 103

affinis Motschulsky, 1844, Cicindela 104

affinis Stein, 1881, Colymbetes 45

affinis Rambur, 1837, Cymindis 412



affinis C. R. Sahlberg, 1827, Dicheirotrichus [Harpalus] 403

affinis Stephens, 1828, Haliplus 31

affinis Ballion, 1878, Harpalus 382

affinis Schrank, 1781, Harpalus [Carabus] 23, 371



affinis Régimbart, 1895, Hydrovatus 70

affinis Stephens, 1828, Hygrotus 73

affinis Chaudoir, 1843, Masoreus 357



affinis Aubé, 1838, Nebrioporus [Hydroporus] 67

affinis Sturm, 1835, Nebrioporus [Hydroporus] 66

affinis Sturm, 1824, Poecilus [Platysma] 482

affinis Stephens, 1828, Pterostichus [Omaseus] 499

affinis Bach, 1851, Stenolophus 406

affinissimus Ali, 1967, Poecilus [Pterostichus] 481



afflatus Scholz, 1917, Hydroporus 64

afgana Jedlika, 1956, Caphora 357

afgana Jedlika, 1956, Cymindis 24, 416

afgana Jedlika, 1956, Glycia 409

afganistana Jedlika, 1956, Klapperichella 459

afganum Lapouge, 1930, Calosoma [Campalita] 120

afganus Jedlika, 1956, Chlaenius 349

afghana Mandl, 1955, Calomera [Cicindela] 100

afghana Kult, 1961, Tachyura 278

afghana Mateu, 1976, Zolotarevskyella 426



afghanica Emetz, 1972, Cymindis 416

afghanicus Wewalka, 1971, Deronectes 57



afghanistanica Jedlika, 1967, Tachyura [Tachys] 278

afghanistanicum Fassati, 1957, Bembidion 244

afghanistanicum Jedlika, 1967, Bembidion [HN!] [ND] 271

afghanistanum Jedlika, 1968, Bembidion 267



afghanistanus Jedlika, 1967, Chlaenius 356

afghanistanus Jedlika, 1965, Daptus 369

afghanistanus Jedlika, 1967, Licinus [Colpostoma] 442

afghanus Jedlika, 1961, Agatus 24, 408

afghanus Jedlika, 1967, Anisodactylus [HN] 361

afghanus Schauberger, 1929, Anisodactylus 361

afghanus Jedlika, 1967, Brachinus 215

afghanus Jedlika, 1967, Dyschirius 224

afghanus Mateu, 1976, Microlestes 424

afghanus G. Ochs, 1955, Orectochilus 28



afghanus Jedlika, 1961, Pseudotaphoxenus [Taphoxenus] 540

afghanus Jedlika, 1967, Scarites 233



africana Chaudoir, 1873, Cymindis 413

africana Boheman, 1848, Drypta 358

africana Dejean, 1825, Galerita 359



africana Solier, 1835, Lebia 427

africanum Netolitzky, 1911, Bembidion 256

africanus Pederzani & Schizzerotto, 1998, Agabus 38



africanus Sharp, 1882, Copelatus 48

africanus Laporte, 1835, Cybister 50



africanus Putzeys, 1846, Dyschirius 228

africanus Sharp, 1882, Hyphydrus 74

africanus Dejean, 1825, Pheropsophus [Brachinus] 217

africanus Rocchi, 1976, Prodaticus 54

afrum Duftschmid, 1812, Agonum [Carabus] 449

agaboides Sharp, 1882, Hydaticus 53

agakhaniantzi Mikhailov, 1972, Harpalus [Loxophonus] 372

agamemnon Breuning, 1943, Carabus 128

agathense Mayet, 1908, Parazuphium [Zuphium] 573

ageneralovi E. Berlov & O. Berlov, 1996, Pterostichus 505

agiensis Uéno, 1975, Kurasawatrechus 313

agile Jacquelin du Val, 1851, Bembidion 269

agile Stephens, 1828, Bembidion [Peryphus] 255

agilis Rylands, 1841, Amara 551

agilis Chaudoir, 1856, Chlaenius 353

agilis Peyron, 1858, Chlaenius 352

agilis Fabricius, 1792, Copelatus [Dytiscus] 47

agilis Fabricius, 1787, Dromius [Carabus] 419

agilis Ledoux & Roux, 1996, Nebria 89

agilis Sharp, 1884, Orectochilus 28



agilis Motschulsky, 1850, Trechus 339

agilis Tschitschérine, 1897, Trichotichnus 396

agiloides Jedlika, 1935, Chlaenius 353

aglaia Andrewes, 1930, Lebia 430

agnata Fleutiaux, 1890, Cylindera [Cicindela] 110

agnatus Ganglbauer, 1889, Carabus [Plectes] 200

agnatus Motschulsky, 1844, Dyschirius 224

agnatus Chaudoir, 1846, Ophonus 388

agnatus Chaudoir, 1876, Pheropsophus 217

agni Deuve & Quéinnec, 1985, Trechus 325

agnolettii Vigna Taglianti, 1999, Laemostenus 533

agnus Foster, 1992, Rhithrodytes 68

agona Tschitschérine, 1898, Amara 558

agonaderus Chaudoir, 1850, Tapinopterus [Feronia] 519

agoniothorax Bedel, 1906, Cymindis 414


agonoderus Putzeys, 1878, Harpalus 386

agonoides Bates, 1883, Stenolophus 405

agonoides Bates, 1889, Synuchus [Pristodactyla] 545

agonus Puel, 1914, Bradycellus 400

agonus Tschitschérine, 1894, Harpalus 381

agonus G. H. Horn, 1880, Pterostichus 497

agonus Tschitschérine, 1895, Synuchus [Calathus] 545

agostini Jeannel, 1937, Rhegmatobius 240

agrestis Creutzer, 1799, Carabus 199

agricola Olivier, 1790, Licinus [Carabus] 442

agrorum Geoffroy, 1785, Agonum [Buprestis] 450

agrorum Olivier, 1795, Chlaenius [Carabus] 350

agurense Lutshnik, 1938, Bembidion 246

agyzimbanus Fairmaire, 1869, Stenolophus 405

aharonii Reitter, 1909, Scarites 233

ahascoanus Lapouge, 1925, Carabus 167

ahenus Ahrens, 1830, Dyschirius [Clivina] 224

ahmetustaensis Cavazzuti, 1987, Carabus 157



ahngeri Poppius, 1908, Lebia 428

ahngeriana Tschitschérine, 1903, Amara 562

ahrensi J. Schmidt, 1995, Xestagonum 467

ai Barševskis, 1993, Syntomus 432



aibgensis Stark, 1890, Carabus 133

aibgensis Belousov, 1998, Cimmerites 297

aibgensis Zamotajlov, 1992, Deltomerus 283

aibgensis Stark, 1890, Pterostichus 500

aida Schatzmayr, 1936, Dyschirius 224

aida Mandl, 1959, Megacephala 118

aidovskanus Ganglbauer, 1913, Anophthalmus [Trechus] 290



aimaki Jedlika, 1964, Amara 552

aimaki Jedlika, 1964, Apristus [Metabletus] 431



aimaki Jedlika, 1964, Bembidion 244

aimaki Jedlika, 1968, Harpalus 376

aimaki Jedlika, 1964, Pseudotaphoxenus [Taphoxenus] 542

aimaki Jedlika, 1968, Pterostichus 510

aimonissabaudiae Baliani, 1932, Amara 564

aino Rost, 1908, Carabus 164

aino Lewis, 1891, Cicindela 105

ainu Habu & Baba, 1968, Bembidion 253



ainus Habu & Baba, 1963, Harpalus 378

airumlus Kolenati, 1845, Nebrioporus [Hydroporus] 65

aishanensis Kleinfeld, 1999, Carabus 137

aisrrensis Belousov, 1992, Carabus 131



aizuensis Jedlika, 1955, Pterostichus 494

ajax Britton, 1948, Badister 440



ajax Breuning, 1933, Carabus 123

ajmasianus Bolívar y Pieltain, 1922, Laemostenus 535

ajmonis Netolitzky, 1935, Bembidion 270

akaishicus Tanaka, 1958, Pterostichus 501

akaishiensis Ishikawa, 1986, Carabus [Leptocarabus] 149

akane Imura & Yamaya, 1994, Carabus 141

akbensis Jedlika, 1958, Harpalus 372

akbensis Mateu, 1955, Orthomus 476

akbesianus Breuning, 1975, Carabus 185

akbesiensis Toulgoët, 1987, Carabus 185

akcakocae Breuning, 1969, Cychrus 202

akensis Haury, 1889, Carabus 153

akibensis Belousov, 1990, Trechus 325

akinini A. Morawitz, 1886, Carabus 145

akinini Tschitschérine, 1895, Harpalus 372

akinini Tschitschérine, 1887, Poecilus 481

akinobui Uéno, 1986, Trechiama 321

akirai Uéno, 1990, Accoella 289

akitanus Ishikawa, 1992, Carabus [Leptocarabus] 158

akiyamai Mandl, 1981, Protocollyris [Neocollyris] 117

akkus Jedlika, 1963, Platyderus 521

akkusensis Breuning, 1964, Carabus 193

akkusianus Kirschenhofer, 1981, Pterostichus 489

akozlovi E. Berlov & O. Berlov, 1996, Pterostichus 505

akozyrevi O. Berlov & E. Berlov, 1999, Pterostichus 486

aksautensis Belousov, 1998, Alanorites 290

aksekianus Straneo, 1988, Pterostichus [Haptotapinus] 496

akserai Jedlika, 1961, Cymindis 24, 416

aksuanus Gottwald, 1987, Carabus 145

aksuensis Jedlika, 1958, Pseudotaphoxenus [Taphoxenus] 541

aksuensis Belousov & Kabak, 1996, Trechus 325

aktashiensis Kirschenhofer, 1986, Brachinus 213

alacer Andrewes, 1937, Holcoderus 436

alacer Boheman, 1848, Megalonychus [Anchomenus] 460

alacer Coquerel, 1859, Platyderus [Feronia] 522

alacer A. Morawitz, 1862, Pterostichus 493

alacris Tschitschérine, 1899, Amara 567

alacris Chevrolat, 1861, Harpalus 382

alaf Britton, 1948, Bembidion 260



alagoensis Kraatz, 1897, Carabus [Mimocarabus] 169

alagoensoides Mandl, 1975, Carabus 194



alaiense Lapouge, 1930, Calosoma [Campalita] 120

alaiensis Tschitschérine, 1894, Amara 562

alaiensis Kabak, 1995, Leistus 83

alaiensis Glasunov, 1902, Nebria 91


alajanicus Jedlika, 1957, Harpalus 377

alajensis Semenov, 1897, Carabus 182



alajensis Tschitschérine, 1896, Cymindis 415

alajensis Znojko, 1930, Dyschirius 224

alajensis Tschitschérine, 1898, Harpalus 372

alajensis Belousov & Kabak, 1994, Trechus 325



alamannicus Csiki, 1927, Carabus 152

alanicus Belousov, 1990, Trechus 325

alanstivelli Morvan, 1981, Carabus 161

alanstivelli Morvan, 1999, Xestagonum 467



alasanica Motschulsky, 1839, Calomera [Cicindela] 100

alaskana Csiki, 1929, Amara 558

alaskanus Obydov, 1996, Carabus 165

alaskense Lindroth, 1962, Bembidion 247



alaskensis Basilewsky, 1937, Carabus 134

alaskensis Lindroth, 1968, Harpalus 384

alatavicus Semenov, 1908, Taphoxenus 544

alatus Schauberger, 1922, Bradycellus 400

alatus Verhoeff, 1891, Calathus 526

alatus Wollaston, 1854, Laemostenus [Pristonychus] 536

alatus Reitter, 1905, Philorhizus [Dromius] 425

alatus Uéno, 1979, Trechiama 321



albanensis Géhin, 1885, Carabus 151

albanensis Schauberger, 1933, Ophonus [Harpalus] 389

albanica Apfelbeck, 1909, Cicindela 106

albanicum J. Müller, 1935, Bembidion 260

albanicus Apfelbeck, 1906, Calathus 526



albanicus Breuning, 1943, Carabus [Procrustes] 187

albanicus Hopp, 1929, Cychrus 205



albanicus J. Müller, 1922, Dyschirius 226

albanicus Reitter, 1900, Harpalus 372

albanicus J. Müller, 1914, Laemostenus 534

albanicus J. Müller, 1932, Licinus 443

albanicus Apfelbeck, 1904, Molops 472

albanicus Apfelbeck, 1906, Omphreus 446

albanicus Tschitschérine, 1900, Speluncarius [Platysma] 516

albanicus Apfelbeck, 1907, Trechus 326

albanicus Apfelbeck, 1904, Zabrus 570

albarracina Hieke, 1984, Amara 563

albarracinensis Fery, 1986, Ilybius [Agabus] 41

albarracinus Wagner, 1926, Brachinus 212



albarracinus Ganglbauer, 1886, Carabus 167

albarracinus Wagner, 1926, Ophonus [Harpalus] 388

alberichae Español, 1971, Apoduvalius 295

albericus Jeanne, 1985, Pterostichus 512


albertana Casey, 1924, Amara [Curtonotus] 561

alberti Jeannel, 1939, Aphoenops 294

alberti Jeannel, 1942, Pterostichus [Oreophilus] 503

alberti Burgeon, 1935, Tachyura [Tachys] 277

albertianus Guignot, 1959, Hydrovatus 71

albianus Depoli, 1926, Molops 473

albicornis Schaum, 1860, Tachyura [Tachys] 278

albida Dejean, 1825, Chaetodera [Cicindela] 103

albidipenne Casey, 1918, Bembidion 262

albidivalis Uéno, 1994, Trechiama 321

albiensis J. Müller, 1925, Abax 469

albina Wiedemann, 1819, Chaetodera [Cicindela] 103



albinus Reitter, 1911, Haliplus 30

albipes Heer, 1838, Bembidion 268

albipes Stephens, 1829, Bembidion [Peryphus] 268

albipes Sturm, 1825, Bembidion 267

albipes Fabricius, 1796, Paranchus [Carabus] 462

albissoni Reitter, 1908, Chlaenius 355

albocyanescens W. Horn, 1912, Neocollyris [Collyris] 116

alboequus Deuve & Imura, 1993, Carabus 176

alboguttata Klug, 1832, Calomera [Cicindela] 99

albomaculata Habu, 1979, Parena 433

albomaculatum J. R. Sahlberg, 1900, Bembidion 244

albomaculatus Lucas, 1846, Lionychus [Dromius] 431

albomaculatus Ballion, 1871, Mnuphorus [Lebia] 357

albonotatus Dejean, 1825, Lionychus [Dromius] 431

alboobliquatus W. Horn, 1909, Therates 115



albopilosa Dokhtouroff, 1885, Cicindela 107

albopunctata Chaudoir, 1852, Cylindera [Cicindela] 110

alboreducta W. Horn, 1934, Cephalota [Cicindela] 102

albovittata Naviaux, 1985, Cylindera 112

albowi Reitter, 1896, Carabus 132

albrechti A. Morawitz, 1862, Carabus 176



albuferensis Báguena Corella, 1924, Lophyra [Cicindela] 112

alcaracinus Kraatz, 1886, Carabus [Hadrocarabus] 167

aldanicum Poppius, 1906, Agonum [Platynus] 452

aldarkose Kabak, 1992, Carabus 123

aldensis Jeannel, 1955, Geotrechus 309

alecrinensis Franz, 1981, Trechus 335

alecto Andrewes, 1930, Amara 552



alecto Bates, 1883, Leistus 82

alegrei Español & Comas, 1985, Aphaenotyphlus 238

alei Abdul‑Karim & Ali, 1986, Hygrotus 73

aleppense Fassati, 1947, Bembidion 262

ales Semenov & Znojko, 1929, Daer 427


alesi Jedlika, 1935, Chlaenius 349

alesi Jedlika, 1935, Dromius 421

alesi Jedlika, 1937, Pristosia [Eucalathus] 530

alessiensis Apfelbeck, 1901, Carabus 195

alexandrae Semenov, 1887, Carabus 121

alexandrae Kataev, 1990, Harpalus 372

alexandrae Tschitschérine, 1889, Poecilus 481

alexandri F. Battoni, 1984, Brachinus 213

alexandri Kalashyan, 1999, Chilotomus 364

alexandriensis Hieke, 1988, Amara 558

alexandrinus Schatzmayr & Koch, 1934, Tachys [DA] 276

alexandrovi Lutshnik, 1915, Blemus [Trechus] 296

alexandrovi Lafer, 1979, Pterostichus 495

alexeevi Zamotajlov, 1992, Deltomerus 282

alexeevi Belousov, 1992, Duvalius 301

alexeevi Kataev, 1990, Harpalus 372

alexeji Zamotajlov & Kryzhanovskij, 1992, Pterostichus 499

alfambrai Lagar Mascaró, 1979, Hydraphaenops 311

alfierii Peyerimhoff, 1927, Dromius 419

alfierii Schatzmayr & Koch, 1933, Torretassoa 231



algarbica Serrano, 1988, Cicindela 106

algarvensis Jeanne, 1985, Trechus 340

algarvensis Coiffait, 1971, Typhlocharis 241

algens Andrewes, 1934, Calathus 528



algerianus Luna de Carvalho, 1957, Paussus 211

algericus Raffray, 1873, Chlaenius 352

algerinus Gory, 1833, Chlaenius 355

algerinus Gory, 1833, Laemostenus [Pristonychus] 537

algesiranus Dieck, 1870, Platyderus 521

algibensis Fery & Fresneda, 1988, Deronectes 58



algida Andrewes, 1930, Amara 548

algidum Andrewes, 1935, Bembidion 243



algidus Poppius, 1904, Poecilus [Pterostichus] 480

algirica Desmarest, 1849, Megacephala 118

algiricum Géhin, 1885, Calosoma 120

algiricus Gautier des Cottes, 1866, Calathus 524

algiricus Géhin, 1885, Carabus 152

algiricus Putzeys, 1846, Dyschirius 228

algiricus Jedlika, 1958, Laemostenus 537

algiricus Jeannel, 1932, Lymnastis 274



algiricus Csiki, 1928, Ocys [Bembidion] 272

algiricus Csiki, 1930, Poecilus [Pterostichus] 481

algiricus Lucas, 1846, Polyderis [Bembidium] 274

algiricus Jeannel, 1922, Trechus 326

algirinum Motschulsky, 1864, Cymbionotum [Graniger] 235


algirinus Reitter, 1887, Amblystomus 360

algirinus Motschulsky, 1864, Graniger 366

algirinus Buquet, 1840, Platynus [Anchomenus] 462

alhambrae Fery, 1999, Hydroporus 63

alhamillensis Cobos, 1961, Platyderus 521

aliai Escalera, 1944, Carabus [Cathoplius] 137

alicantinus Ganglbauer, 1886, Carabus 167

alicantinus Español, 1971, Trechus 326

aliense Habu, 1973, Bembidion 268

aliensis Habu, 1978, Synuchus 545

alienus A. Zimmermann, 1919, Bidessus 54

alienus LeConte, 1879, Harpalus 378

alienus Schauberger, 1928, Harpalus 375

alienus Sharp, 1873, Oreodytes [Hydroporus] 68

alii Marggi & Huber, 1999, Bembidion 269

alikhelicum Kirschenhofer, 1989, Bembidion 249

alinae Lafer, 1988, Agabus [Gaurodytes] 38

alinae J. Müller, 1932, Molops 475

alini Jedlika, 1942, Amara 567

allax Peschet, 1923, Orectochilus 29

alliciens Deuve, 1990, Carabus 189

alligator Normand, 1933, Agabus 38

allobrogica Jeanne, 1976, Nebria 93

alloligops Uéno, 1978, Epaphiopsis 308

alluaudi Antoine, 1941, Acupalpus 399

alluaudi Paulian & Villiers, 1939, Amara 558

alluaudi Antoine, 1925, Bembidion [Peryphus] 253



alluaudi Colas, 1936, Carabus 168

alluaudi Antoine, 1939, Cymindis 411

alluaudi Mateu, 1952, Dicrodontus 573

alluaudi Antoine, 1922, Harpalus 379

alluaudi Bedel, 1899, Laemostenus 539

alluaudi Jeannel, 1926, Perileptus 288



alluaudi Jeannel, 1946, Tachyura [Tachyphanes] 278

alluaudi Peschet, 1926, Yola 57


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