Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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elegans Stephens, 1835, Bembidion [Peryphus] 260

elegans Chaudoir, 1842, Brachinus 213



elegans Bates, 1883, Broscosoma 235

elegans Kirsch, 1859, Calosoma [Callisthenes] 118



elegans Kwon & Lee, 1984, Carabus [Damaster] 142

elegans Weber, 1801, Catascopus [Elaphrus] 435

elegans Fischer von Waldheim, 1823, Cephalota [Cicindela] 102

elegans Belousov, 1998, Cimmerites 297

elegans Chaudoir, 1871, Deltomerus 282

elegans Uéno, 1989, Epaphiopsis 309

elegans Landin, 1955, Gunvorita 573

elegans Preller, 1862, Harpalus 372

elegans A. Gaudin, 1945, Hydraphaenops 311



elegans Brullé, 1832, Laemostenus [Pristonychus] [HN] 537

elegans Dejean, 1828, Laemostenus [Pristonychus] 532

elegans Rost, 1891, Leistus 82

elegans Andrewes, 1925, Nebria 94

elegans Panzer, 1794, Nebrioporus [Dytiscus] 66

elegans Bedel, 1900, Platyderus 522



elegans Coye, 1870, Pseudaristus [Aristus] 367

elegans Uéno, 1960, Rakantrechus 319

elegans Coiffait, 1962, Sphodropsis 543

elegans Chaudoir, 1861, Stomis 517

elegans Putzeys, 1847, Trechus 330

elegans Dodero, 1916, Typhloreicheia [Reicheia] 223

elegantellus Lorenz, 1998, Platynus [Colpodes] 464

eleganticollis Deuve & Imura, 1991, Carabus 126

eleganticolor Deuve, 1997, Carabus 182

elegantula Tschitschérine, 1899, Amara 554



elegantula Schmidt‑Göbel, 1846, Coptodera 436

elegantula Antoine, 1962, Geocharis [DA] 239

elegantula Schmidt‑Göbel, 1846, Mastax 216

elegantulum R. F. Sahlberg, 1844, Bembidion 254

elegantulus Motschulsky, 1850, Carabus 193

elegantulus Ménétriés, 1832, Harpalus 375

elegantulus Sciaky, 1987, Ophonus 390



elegantulus Louwerens, 1953, Platynus [Colpodes] 464

elegantulus Normand, 1916, Pseudanillus [Anillus] 238



elegantulus Sciaky & Pavesi, 1997, Pseudotaphoxenus 540

elegantulus Chaudoir, 1844, Pterostichus [Feronia] 505

elemeri Mihók, 1911, Duvalius [Anophthalmus] 300

eleonorae Bonavita & Vigna Taglianti, 1993, Bembidion [Ocydromus] 245

elephas Putzeys, 1872, Carabus 168

elevata Motschulsky, 1844, Amara [Bradytus] 557

elevata Lindroth, 1956, Pristosia 531



elevatum Puel, 1938, Agonum 449

elevatum Motschulsky, 1844, Bembidion [Leja] 251

elevatus Panzer, 1793, Brychius [Dytiscus] 30

elevatus Schmidt‑Göbel, 1846, Catascopus 435

elevatus Sharp, 1882, Hydrovatus 71

elevatus Sharp, 1882, Hygrotus [Coelambus] 72

elevatus Sharp, 1882, Hyphoporus 73



elevatus Sharp, 1882, Rhantus 46

elevatus Fabricius, 1792, Somotrichus [Carabus] 439

elferi G. Ochs, 1940, Orectochilus 29

elgae Hendrich, Balke & Wewalka, 1995, Microdytes 75

elikasensis Morvan, 1976, Trechus 329

elisabethae Semenov, 1908, Carabus 145



elisabettianus Font, 1997, Carabus 143

elisae Motschulsky, 1859, Cylindera [Cicindela] 110



eljasi Wirén, 1968, Suphrodytes [Hydroporus] 70

ellipsoideus Balkenohl, 1995, Reicheiodes 230



elliptica Heyden, 1885, Amara 549

elliptica Reitter, 1894, Amara [Pterostichus] 558

ellipticocurtum Netolitzky, 1935, Bembidion 264



ellipticus Cuvier, 1833, Abax [Carabus] 470

ellipticus Reitter, 1889, Calathus 524



ellipticus Ballion, 1878, Harpalus 380

ellipticus Bates, 1878, Harpalus [Hypsinephus] 381

ellipticus Régimbart, 1889, Laccophilus 76

ellipticus Schaufuss, 1862, Laemostenus [Pristonychus] 533

ellipticus Reitter, 1885, Leistus 82

ellipticus Wollaston, 1857, Leistus 83

ellipticus Andrewes, 1937, Miltodes 445

ellipticus Porta, 1901, Percus [Abax] 478

ellipticus Bedel, 1902, Platyderus 522



ellipticus Heer, 1841, Pterostichus 513

elliptipennis Antoine, 1961, Atlantomasoreus 357

elliptipennis Deuve, 1995, Carabus 124

elliptipennis Bates, 1874, Nebria 85

elliptipennis Wollaston, 1864, Philorhizus [Dromius] 425



ellypticus Petri, 1912, Hydroporus 61

elmbergi Poppius, 1908, Pterostichus 505



elmii Fery & Hosseinie, 1998, Deronectes 58

elonensis Schweiger, 1970, Carabus 157



elongata Baliani, 1937, Amara 568

elongata LeConte, 1850, Amara [Curtonotus] 560

elongata Randall, 1838, Clivina 219

elongata Heer, 1841, Cymindis 413

elongata Bates, 1873, Diplocheila [Rembus] 439



elongata Chaudoir, 1844, Drypta 358

elongata Uéno, 1962, Epaphiopsis 309

elongata Dupuis, 1913, Hexagonia 407

elongata Uéno, 1955, Jujiroa 459

elongata Uéno, 1999, Kusumia 314

elongata Mannerheim, 1823, Pelophila 98

elongaticeps Deuve, 1992, Cychrus 204



elongatior Deuve, 1991, Carabus 150

elongatipennis Imura & Yamaya, 1994, Carabus 159

elongatosignata W. Horn, 1922, Calomera [Cicindela] 100



elongatulus Duftschmid, 1812, Demetrias [Lebia] 419

elongatulus Dawson, 1856, Dyschirius 224

elongatulus Sturm, 1835, Hydroporus 61

elongatulus Fischer von Waldheim, 1829, Platynus 464

elongatulus Uéno, 1979, Ryugadous 319

elongatulus Casale, 1983, Sphodropsis 543

elongatulus Dejean, 1831, Tachys [Bembidium] 275

elongatulus Putzeys, 1870, Trechus 330



elongatum Fischer von Waldheim, 1823, Agonum 454

elongatum Dejean, 1831, Bembidion 267



elongatus Lucas, 1846, Acinopus 368

elongatus Gyllenhal, 1826, Agabus [Dytiscus] 36



elongatus Tournier, 1864, Brachinus 214

elongatus Motschulsky, 1860, Bradycellus [Stenolophus] 401



elongatus Dejean, 1828, Calathus 527

elongatus Fischer von Waldheim, 1842, Carabus 136

elongatus Motschulsky, 1859, Carabus 188

elongatus Stark, 1894, Carabus [Cechenus] 137

elongatus Audinet‑Serville, 1821, Cychrus 203

elongatus Dejean, 1831, Deltomerus [Platynus] 282



elongatus Ahrens, 1830, Dyschirius [Clivina] 226

elongatus Fischer von Waldheim, 1828, Elaphrus 206

elongatus Jeannel, 1946, Geotrechus 310



elongatus Jeannel, 1946, Geotrechus 310

elongatus Aubé, 1838, Gyrinus 27

elongatus Dejean, 1828, Laemostenus [Pristonychus] 534



elongatus J. Ochs, 1949, Laemostenus [Antisphodrus] 532

elongatus Winkler, 1926, Neotrechus 315



elongatus Wollaston, 1854, Olisthopus 461

elongatus Chaudoir, 1859, Orthomus [Feronia] 477

elongatus Chaudoir, 1842, Panagaeus 447

elongatus Kanô, 1930, Paussus 210



elongatus Porta, 1901, Percus [Abax] 478

elongatus Uéno, 1996, Phreatodytes 34

elongatus Motschulsky, 1850, Pseudotaphoxenus [Taphoxenus] 542

elongatus Duftschmid, 1812, Pterostichus [Carabus] 499



elongatus A. Costa, 1847, Zabrus 570

elongatus Ménétriés, 1832, Zabrus 572

eluta Andrewes, 1923, Coptodera 435

elysii J. Thomson, 1856, Carabus 141



elytralis Obenberger, 1917, Molops 474

elytralis Straneo, 1983, Pterostichus 510

elytrolargus Morvan, 1981, Laemostenus [Pristonychus] 538

emanuelei Imura, 1997, Carabus 128



emarginata Gmelin, 1790, Drypta [Carabus] 358

emarginata Nietner, 1858, Tachyura [Bembidium] 277

emarginatibasis N. Ito, 1998, Trichotichnus 394



emarginatum Gyllenhal, 1827, Agonum [Harpalus] 449

emarginatus Chaudoir, 1842, Acinopus 368

emarginatus Duftschmid, 1812, Carabus 196



emarginatus Olivier, 1795, Licinus [Carabus] 442

emarginatus Chaudoir, 1876, Pheropsophus 217

emarginatus Herbst, 1806, Scarites 234

emarginatus Andrewes, 1930, Trichotichnus 395

emarginatus Uéno, 1982, Yamautidius 345

embersoni Deuve, 1995, Trechiotes [Archaeotrechiama] 324

emblema Marseul, 1871, Platyderus 522

emdeni Kult, 1949, Dyschirius 227

emei Sciaky, 1994, Pterostichus 500

emeicola Deuve & Imura, 1992, Carabus 128

emeiensis Kataev & N. Ito, 1999, Trichotichnus 394

emerita Péringuey, 1898, Tachyura [Tachys] 279

emetzi Mikhailov, 1977, Cymindis 415



emgei Reitter, 1884, Brachinus 214

emgei Ganglbauer, 1888, Carabus [Procrustes] 187

emiratensis Wiesner, 1993, Hypaetha 112



emmerichi Baliani, 1932, Amara 554

emmerichi Reitter, 1895, Carabus 153

emmerichi Falkenström, 1936, Haliplus 31

emmerichi Falkenström, 1936, Orectochilus 29

emmerichi Jedlika, 1932, Platynus [Colpodes] 464

emmerichi Falkenström, 1936, Stictotarsus [Deronectes] 69

empetricola Dejean, 1828, Pterostichus [Feronia] 490


empiricus Wollaston, 1865, Nesacinopus [Cratognathus] 388

encaustus Fairmaire, 1868, Calathus 528

encephalus Lucas, 1858, Scarites 233

enci Jedlika, 1969, Harpalus 383

encopoleus Solsky, 1873, Poecilus 482

endogaeus Uéno & Baba, 1965, Kurasawatrechus 313

endymion Andrewes, 1935, Bembidion 260

enigmatica Omer‑Cooper, 1954, Yola 57

enigmaticus Belousov, 1998, Alanorites 290

enigmaticus Heinz, 1980, Carabus 193

enigmaticus Coiffait, 1971, Trechus 330

enk Morvan, 1999, Xestagonum 468

enleensis Mandl, 1992, Chlaenius 353

enneagrammus Ahrens, 1833, Hygrotus [Hydroporus] 71

enops Schaufuss, 1882, Carabus 160

enricoi S. Battoni & Blumenthal, 1973, Carabus 158

ensiger Kurnakov, 1960, Deltomerus 282

eobius Tschitschérine, 1899, Pterostichus [Platysma] 507

eohalensis Jeannel, 1937, Dolichus 530

eokirgisicus Kabak, 1990, Carabus 150

eos Uéno, 1982, Yamautidius 345

eous A. Morawitz, 1889, Carabus 150



eous Znojko, 1928, Dyschirius 224

eous Tschitschérine, 1901, Harpalus 385

eous Tschitschérine, 1894, Myas [Feronia] 476

ephippiatus Fairmaire, 1884, Metadromius [Dromius] 421

ephippiatus Bates, 1873, Trechus 325

ephippiger Duftschmid, 1812, Anthracus [Carabus] 399

ephippiger Andrewes, 1932, Metadromius [Metabletus] 421



ephippium Dejean, 1829, Acupalpus 398

ephippium Marsham, 1802, Bembidion [Carabus] 253

ephippium Escalera, 1914, Cymindis 412

epigaeus Puisségur, 1952, Laemostenus [Ceuthosphodrus] 533

epimethaeus Kurnakov, 1972, Carabus 132

epipleuralis Roeschke, 1907, Carabus [Imaibius] [HN] 155

epipleuralis Semenov, 1907, Carabus 155



epipleuralis Munster, 1925, Gyrinus 27

epipleuralis J. R. Sahlberg, 1887, Pterostichus [Feronia] 490

epipleuricus Seidlitz, 1887, Stictonectes [Hydroporus] 69



epirensis Purkyn, 1928, Carabus 160

epirotes Colas, 1957, Trechus 330

epiroticus Csiki, 1930, Pterostichus 515

episcopalis J. Müller, 1931, Anophthalmus 291



episcopalis Reiche, 1863, Ophonus 389

episcopus Drapiez, 1819, Laemostenus [Carabus] 539

episternalis Gerhardt, 1888, Badister 440

epsteini Heinertz, 1978, Carabus 162

eques Sturm, 1825, Bembidion 249

eques Schrank, 1781, Callistus [Carabus] 347

eques Lapouge, 1925, Carabus 195

equestris Duftschmid, 1812, Amara [Carabus] 565



equestris Gaubil, 1849, Cylindera [Cicindela] 110

equestris Gebler, 1825, Cymindis 417



equestris Geoffroy, 1785, Panagaeus [Buprestis] 447

equiphilus Deuve, 1998, Cychrus 205

erberi Heinz, 1983, Carabus 155

erberii Schaufuss, 1863, Laemostenus [Sphodrus] 534

erebia Bates, 1892, Kareya [Platymetopus] 387

erebus Krauss, 1906, Anophthalmus [Trechus] 291

eregliense Jedlika, 1961, Bembidion 264

eregliensis Breuning, 1974, Carabus 130

eremicola Kryzhanovskij, 1962, Amara 562



eremita Tschitschérine, 1894, Amara 559

eremita Quéinnec, 1984, Amerizus [Tiruka] 241

eremita Fischer von Waldheim, 1823, Carabus 135

eremita Dejean, 1825, Dixus [Ditomus] 24, 365

eremita Jeannel & J. Ochs, 1938, Duvalius 304

eremnus Andrewes, 1933, Dromius 420

erenhabergaensis Deuve, 1993, Carabus 150

erenleriensis Schweiger, 1964, Carabus 157



erhardti Nowicki, 1873, Pedius 478

ericae Wollaston, 1854, Olisthopus 461

ericeti Panzer, 1809, Agonum [Carabus] 450

erichsoni Fischer von Waldheim, 1844, Carabus 123

erichsoni Gemminger & Harold, 1868, Ilybius [Agabus] 42

erichsoni Schmidt‑Göbel, 1846, Pentagonica 448

erichsonii Guérin‑Méneville, 1849, Copelatus 47

erichsonii Schaufuss, 1864, Duvalius [Anophthalmus] 306

erillensis Español & Mateu, 1945, Pterostichus [Haptoderus] 490

eriophorus Yoshida & S. Nomura, 1952, Kurasawatrechus 313

erivanus Reitter, 1900, Harpalus 382

eriwanicus Tschitschérine, 1897, Pterostichus [Feronia] 485

erlangensis Uéno, 1998, Epaphiopsis 308

ermaki Zaitzev, 1953, Agabus [Gaurodytes] 36

ermaki Lutshnik, 1924, Carabus 163

ernsti Kataev, 1995, Harpalus 382



erosoides Jedlika, 1953, Harpalus 373

erosus Motschulsky, 1866, Carabus 192

erosus Mannerheim, 1825, Harpalus 376

erotyloides Bates, 1883, Lioptera 436



errabundus Deuve & Imura, 1990, Carabus 127

errans Fischer von Waldheim, 1823, Carabus 199



errans Gory, 1839, Carabus 179

errans Tschitschérine, 1897, Harpalus 377

errans Sharp, 1882, Hydroporus 61



errata P. Rossi, 1790, Lebia [Carabus] 428

erratica Duftschmid, 1812, Amara [Carabus] 556



erraticus Coiffait, 1962, Aphoenops 294

erraticus Jeannel, 1938, Aphoenops 295

erraticus Uéno, 1971, Nipponaphaenops 316

erraticus Sharp, 1884, Rhantus 46



erratoambiguus Verhoeff, 1891, Calathus 527

erratum De Monte, 1949, Bembidion 264

erratus C. R. Sahlberg, 1827, Calathus [Harpalus] 527

errensis Rousselle, 1974, Carabus [Chrysocarabus] 140

erro Newman, 1838, Badister [Trimorphus] 441

ertli Born, 1910, Carabus [Coptolabrus] 142

erudita Wiedemann, 1823, Cylindera [Cicindela] 110



eruditus Dejean, 1828, Pterostichus [Feronia] 508

ervini Antoine, 1941, Ophonus [Harpalus] 389

erwini Mandl, 1975, Carabus 162

erymanthius Ganglbauer, 1915, Zabrus 570

erythrocephala Sturm, 1815, Nebria 91

erythrocephalum Chaudoir, 1862, Zuphium 574



erythrocephalus Buysson, 1901, Demetrias 419

erythrocephalus Linné, 1758, Hydroporus [Dytiscus] 61



erythrocephalus Bassi, 1834, Platynus 462

erythrocerus Fischer von Waldheim, 1829, Harpalus 386

erythrocnema Dejean, 1828, Amara 567

erythrocnemus Fischer von Waldheim, 1828, Carabus 170

erythroderus Gemminger & Harold, 1868, Calathus 527

erythrodes Fischer von Waldheim, 1828, Carabus 170

erythromerus Dejean, 1826, Carabus 170

erythromerus Semenov, 1898, Carabus 198

erythromerus Ganglbauer, 1891, Pterostichus 503

erythropa Stephens, 1828, Amara 549

erythropus Fischer von Waldheim, 1829, Agonum 455



erythropus Fischer von Waldheim, 1820, Carabus 199

erythropus Fischer von Waldheim, 1828, Chlaenius 350

erythropus Motschulsky, 1844, Miscodera 237

erythropus Dejean, 1828, Poecilus 482



erythropus A. Villa & G. B. Villa, 1833, Pterostichus 503

erythropus Marsham, 1802, Pterostichus [Carabus] 508

erythropyga Putzeys, 1866, Clivina 219

erythropyga Putzeys, 1866, Clivina [bis] 220


erythrostomus Deuve, 1987, Trechus 330

erzeliki Schweiger, 1977, Calathus 524

erzeliki Schweiger, 1962, Carabus 198

esakianus Nakane, 1961, Carabus [Apotomopterus] 176

esakii Csiki, 1927, Carabus 177

esakii Nakane, 1956, Coptodera [Coptoderina] 436

esakii Habu, 1969, Parena 434

esakii Habu, 1954, Platyagonum [Agonum] 462

esakii Ishida, 1959, Pterostichus 512

esakii Uéno, 1969, Stygiotrechus 320

esau Heyden, 1885, Eocarterus [Sabienus] 365

escalerae Schauberger, 1932, Acinopus 368

escalerae Csiki, 1927, Carabus 137

escalerae Abeille de Perrin, 1903, Trechus 330



escalerae Ganglbauer, 1915, Zabrus 569

escalerai Morvan, 1977, Microdaccus 422



escalerai Régimbart, 1900, Platambus 44

escalevae Tschitschérine, 1898, Pterostichus [Feronia] 489

escheri Palliardi, 1825, Carabus 139

escheri Heer, 1837, Nebria 93



escheri Motschulsky, 1844, Nebria 89

escheri Heer, 1837, Pterostichus 503

escheri Aubé, 1838, Stictonectes [Hydroporus] 69

escherichi Ganglbauer, 1897, Bembidion 264

eschscholtzi Chaudoir, 1837, Amara [Leirus] 561

eschscholtzi Mannerheim, 1827, Carabus 169

eschscholtzii Zoubkoff, 1829, Blethisa 206

eschscholtzii Mannerheim, 1823, Pelophila 98

eschscholtzii Parry, 1849, Physodera 438

eschscholtzii Germar, 1824, Pterostichus [Platysma] 509

escorialensis Gautier des Cottes, 1866, Amblystomus 360



escorialensis Jedlika, 1958, Cymindis 411

escorialensis Brisout de Barneville, 1885, Microlestes [Blechrus] 422

eserensis Bolívar y Pieltain, 1918, Zabrus 569



esertus Motschulsky, 1850, Dicheirotrichus [Bradycelus] 402

esfandiari Morvan, 1974, Nebria 90

esfandiarii Heinz, 1973, Carabus 192

esfandiarii Morvan, 1974, Laemostenus [Antisphodrus] 534



esfandiarii Morvan, 1974, Trechus 339

eskilos Schatzmayr, 1909, Bembidion 255

eskualduna Coiffait, 1959, Aphoenops 294



esmeraldae Breuning & Ruspoli, 1977, Carabus 186

esoxensis Reitter, 1896, Carabus 181

espagnoli Duran, 1971, Amara [Leiocnemis] 563

espagnoli Lagar Mascaró, 1965, Percus 478


espanoli Salgado Costas, 1996, Apoduvalius 295

espanoli Jeanne, 1985, Microlestes 422

espanoli J. Vives & E. Vives, 1977, Paradromius [Trichodromius] 425

espanoli Mateu, 1952, Platyderus 522

espanoli J. Vives & E. Vives, 1977, Pterostichus [Haptoderus] 490

esperanzae Heinz, 1970, Carabus 193

espinassei Puel, 1937, Bembidion 260

estagnousensis Coiffait, 1952, Trechus 325

estarragnensis Jeannel, 1947, Pterostichus [Haptoderus] 490

estegeicus Cavazzuti, 1989, Carabus [Procerus] 185

estreicheri Fischer von Waldheim, 1820, Carabus 198

estrelensis Jeanne, 1976, Calathus 524

estrellae Zaballos & Ruiz‑Tapiador, 1997, Typhlocharis 241

estrellanus Heyden, 1880, Zabrus 569



estrellensis Schaufuss, 1882, Hydroporus 64

estriatus Fairmaire, 1887, Scarites 234

etainn Morvan, 1996, Xestagonum 467

etelkae Ormay, 1888, Pterostichus 488


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