Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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ethnicus Gistel, 1857, Pterostichus [Poecilus] [ND] 515



etholeni Mannerheim, 1849, Carabus 153

etoi Uéno, 1958, Rakantrechus 319

etontii Casale, 1988, Laemostenus 533

etrusca Bassi, 1834, Cymindis 413

etruscus Quensel, 1806, Gynandromorphus [Carabus] 362

etschmiadsina Eichler, 1924, Amara 568

etsumianus Uéno, 1988, Trechiama 322



euarabica Ali, 1978, Calomera [Cicindela] 99

euboeicus Ganglbauer, 1889, Molops 475

eucheres Netolitzky, 1943, Bembidion 256

euchlorus Ménétriés, 1839, Harpalus 376



euchromus Palliardi, 1825, Carabus 151

eucosmus Andrewes, 1937, Brachinus 215



eudeserticola Ali, 1978, Cephalota [Cicindela] 102

euganensis Schatzmayr, 1944, Abax 470

euganeus Pace, 1974, Orotrechus 316

eugenes Jedlika, 1933, Bembidion 262

eugenes Jedlika, 1935, Lebia 430

eugenia Chaudoir, 1865, Heptodonta 115



eugeniae Zaitzev, 1909, Hydroporus 61

eugeniae K. Daniel, 1903, Nebria 85



euglypta Bates, 1883, Tachyura [Tachys] 278

eugrammus Vereschagina, 1989, Reflexisphodrus [Taphoxenus] 543

eulabes Bates, 1889, Platynus [Colpodes] 464

eupages Andrewes, 1934, Bembidion 248



eupatoriae Schauberger, 1932, Harpalus 382

euphratica Putzeys, 1866, Amara 565

euphratica Putzeys, 1866, Clivina 219

euphratica Dejean, 1822, Megacephala 118

euphratica Reitter, 1885, Tachyura [Tachys] 279

euphraticus Mandl, 1972, Chlaenius [Stenochlaenius] 355

euphraticus Tschitschérine, 1899, Dicheirotrichus [Trichocellus] 403

euphraticus Putzeys, 1846, Dyschirius 225



europae Jeanne, 1967, Laemostenus [Ceuthosphodrus] 535

europaea Dejean, 1826, Siagona 218

europaeus Mateu, 1980, Apristus 431

europaeus Ahrens, 1814, Rhysodes 78

europs Bates, 1886, Bembidion 254

eurycephalus Chaudoir, 1842, Acinopus 368

eurydamas Bates, 1883, Agonum [Colpodes] 452



eurydera Tschitschérine, 1895, Amara 565

eurydice Andrewes, 1926, Bembidion 260

eurydice Schaufuss, 1881, Duvalius [Anophthalmus] 307

eurygonum Bates, 1883, Bembidion 270



eurymorphus Tschitschérine, 1893, Pterostichus [Feronia] 494

eurynota Panzer, 1796, Amara [Carabus] 549



eurynotus Lapouge, 1913, Carabus 138

euryodes Bates, 1892, Tachys 275

eurytanicus J. Müller, 1917, Molops 475

eurytus Fischer von Waldheim, 1828, Scarites 234

eutherum Andrewes, 1923, Bembidion 271

eutrechoides Deuve, 1992, Trechus 330

euviridis Ishikawa & Kim, 1983, Carabus [Damaster] 144

euxinicus Nitzu, 1998, Laemostenus 538

euxinum Apfelbeck, 1904, Bembidion 248

euxinus Semenov, 1897, Carabus 137

euxinus Znojko, 1927, Dyschirius 224

euxinus Straneo, 1935, Pterostichus 490

evanida Andrewes, 1925, Tachyura [Tachys] 277

evavartianae Mandl, 1973, Microdaccus 422

evelynae Fery & Hosseinie, 1998, Deronectes 58

everesti Andrewes, 1929, Carabus 162



eversmanni Ballion, 1855, Agabus 36

eversmanni Ménétriés, 1832, Brachinus 215

eversmanni Fischer von Waldheim, 1832, Carabus 165

eversmanni Motschulsky, 1844, Carabus 160

evgeniae Zamotajlov & Gorbachov, 1997, Carabus 191

evstigneevi Obydov, 1997, Carabus 182

ewersmanni Chaudoir, 1850, Calosoma [Callisthenes] 118

exapicalis Codina, 1916, Cicindela 103

exarata Fischer von Waldheim, 1844, Nebria 87


exarata Bates, 1873, Tachyura [Tachys] 277

exaratulus Beheim & Breuning, 1943, Carabus [Megodontus] 164

exaratum Mannerheim, 1853, Agonum [Anchomenus] 452

exaratum Andrewes, 1924, Bembidion 267

exaratus Dejean, 1828, Abax [Feronia] 470

exaratus Quensel, 1806, Carabus 164

exaratus Schaum, 1860, Duvalius [Trechus] 303

exaratus Putzeys, 1866, Dyschirius 225

exaratus Hampe, 1871, Laemostenus [Sphodrus] 534

exaratus Bates, 1892, Pseudognathaphanus [Gnathaphanus] 363



exaratus Dalman, 1823, Rhysodes 78

exarboreus Ishikawa, 1992, Carabus [Leptocarabus] 159

exasperatus Duftschmid, 1812, Carabus 165

exasperatus Klug, 1832, Scarites 234

excavatus Charpentier, 1825, Carabus 187



excavatus Paykull, 1790, Patrobus [Carabus] 285

excavatus Carret, 1903, Pogonus 287

excavatus Boudier, 1844, Pterostichus [Feronia] 511

excellens Rosenhauer, 1856, Bembidion 270

excellens Fabricius, 1798, Carabus 170

excellens Tschitschérine, 1895, Poecilus [Feronia] 482

excentricus Deuve, 1992, Queinnectrechus 318

exceptus J. R. Sahlberg, 1887, Pterostichus [Feronia] 490

excisicollis Bänninger, 1956, Tonkinoscaris 234

excisus Dejean, 1826, Carabus 196

exclamationis Fabricius, 1792, Graphipterus [Carabus] 359



exclamationis Luna de Carvalho, 1977, Platyrhopalus 211

exclamationis Ménétriés, 1848, Syntomus [Dromius] 432

excoffieri Vondel, 1991, Haliplus 31

excoffieri Régimbart, 1899, Hyphydrus 74

excoffieri Régimbart, 1899, Platambus 43

excoffieri Fairmaire, 1897, Pronyssiformia [Cicindela] 115

excostatus Kraatz, 1880, Carabus 154

excultus Wollaston, 1854, Bradycellus 400

exdebilis Gottwald, 1994, Carabus 182

exedithae Gottwald, 1985, Carabus 131

exhalans P. Rossi, 1792, Brachinus [Carabus] 214

exhibitum Netolitzky, 1911, Bembidion 20, 249

exigua A. Morawitz, 1863, Perigona [Pentoplogenius] 448

exiguiceps Casey, 1924, Bembidion 262

exiguus Dejean, 1829, Acupalpus 398

exiguus Semenov, 1898, Carabus 150

exiguus R. F. Sahlberg, 1844, Tachys 276

exiliatus Cavazzuti, 1990, Carabus 193

exilicaudis Kabak, 1998, Poecilus 484


exilipenis Belousov & Kabak, 1994, Trechus 330

exilis Bates, 1883, Carabus 159

exilis Machado, 1992, Dicrodontus 573

exilis Schaum, 1857, Microlestes 422

exilis Schmidt‑Göbel, 1846, Microlestes 422

exilis Sciaky & Wrase, 1997, Stomis 517

exilis Uéno, 1983, Trechiama 322

eximia Dejean, 1828, Amara 555

eximius Csiki, 1906, Carabus 172

eximius Putzeys, 1870, Duvalius [Trechus] 303

eximius Clark, 1863, Haliplus 31



eximius Clark, 1863, Hyphydrus 74

eximius A. Morawitz, 1862, Pterostichus 505

exisonum Lutshnik, 1938, Bembidion 250

exochus Andrewes, 1930, Pterostichus 515



exoletus Andrewes, 1924, Chlaenius 349

exophtalmus Deuve, 1998, Junnanotrechus 313

exophthalmica Andrewes, 1930, Aparupa 456

exornatum Andrewes, 1930, Bembidion 243

exornatus Reiche & Saulcy, 1855, Bidessus [Hydroporus] 55

exornatus Machado, 1992, Paradromius 424



expallidus Zaitzev, 1908, Haliplus 32

expansicollis Ledoux, 1989, Archileistobrius 79



expansus Lapouge, 1905, Carabus 136

expansus Putzeys, 1874, Leistus 18, 84



expansus Mateu, 1957, Orthomus 477

expectatus Zamotajlov, 1992, Deltomerus 282

expectatus Aubry, 1971, Pterostichus 498

expectatus Uéno, 1983, Trechiama 322

expeditus Tschitschérine, 1898, Pterostichus [Feronia] 500

explanatula Casey, 1924, Amara [Celia] 564

explanatus Ledoux & Roux, 1995, Archastes 79

explodens Duftschmid, 1812, Brachinus 214

expolita Baliani, 1934, Amara 554

exposita Casey, 1918, Amara [Celia] 564

exquisitum Andrewes, 1923, Bembidion 263



exquisitus Mulsant & Rey, 1868, Diachromus 362

exsanguis Bedel, 1925, Graptodytes [Hydroporus] 59

exsculpta Hausmann, 1866, Amara 549

exsoletus Forster, 1771, Rhantus [Dytiscus] 46



exspectatus Peyerimhoff, 1905, Dytiscus 52

exsultans Cavazzuti, 1993, Carabus 158

exsuturalis Beuthin, 1899, Cicindela 103

extensicollis Putzeys, 1873, Calathus 527

extensicollis Putzeys, 1846, Clivina 219


extensoides Jedlika, 1936, Tapinopterus [Platysma] 519

extensoides Straneo, 1936, Tapinopterus [HN] 519

extensomarginata Dokhtouroff, 1885, Cicindela 105

extensum Ménétriés, 1849, Agonum 450



extensus Kraatz, 1885, Carabus 179

extensus Mannerheim, 1825, Chlaenius 349

extensus Winkler, 1933, Duvalius 300

extensus Putzeys, 1846, Dyschirius 224



extensus Walker, 1866, Harpalus [Amara] 377

extensus Heyden, 1884, Laemostenus [Antisphodrus] 537

extensus Chaudoir, 1871, Sirdenus 288

extensus Dejean, 1831, Tapinopterus [Feronia] 519



exter Csiki, 1927, Carabus 145

externepunctatus Reitter, 1899, Dicheirotrichus 403



externepunctatus Solsky, 1874, Microderes [Pangus] 387

externepunctatus Dejean, 1828, Pterostichus [Feronia] 503

extraneus Schauberger, 1921, Pterostichus [Lyperophorus] 508

extrema Hieke, 1995, Amara 559



extrema Walker, 1858, Tachyta [Acupalpus] 277

extremus Schaufuss, 1882, Carabus 157

extremus Semenov, 1898, Carabus 145

extremus Chaudoir, 1876, Chlaenius 351

extremus Bedel, 1905, Tetragonoderus 357

exul Peyerimhoff, 1910, Nebria 85

exutus I. Frivaldszky von Frivald, 1845, Chlaenius 349

eylandi Semenov, 1889, Discoptera 356

ezo Nakane, 1989, Agabus 36

fabbrii J. Müller, 1931, Anophthalmus 291



faber Ménétriés, 1832, Harpalus 373

fabianii Gestro, 1900, Orotrechus [Anophthalmus] 316

fabichi Maan, 1934, Asaphidion 20, 242

fabioi Daffner, 1996, Anophthalmus 292



fabrei Colas, 1962, Carabus 195

fabressei Régimbart, 1901, Nebrioporus [Hydroporus] 66



fabricii Beuthin, 1892, Calomera [Cicindela] 100

fabricii Panzer, 1810, Carabus 184



fabricii Crotch, 1871, Harpalus 376

fabricii W. S. MacLeay, 1825, Hydaticus [Colymbetes] 53



fabricii Cristoforis, 1832, Hyphydrus 74

fabricii Crotch, 1871, Licinus 442

fabriciiruspolii Breuning, 1969, Carabus 130

fabriciusi A. Ball, 1932, Hydaticus 53

fabriziobattonii Kirschenhofer, 1986, Lebia 428

fabulosum Semenov & Znojko, 1933, Calosoma 121

facchiniellus Deuve, 1998, Trechus 339


facchinii Toledano, 1998, Bembidion 252

facchinii Farka & Sciaky, 1998, Leistus 80

facchinii Sciaky, 1998, Stomis 517

facetus Semenov, 1903, Carabus 147

facialis Wiedemann, 1819, Catascopus [Carabus] 435

fadriquei Español, 1999, Speleotyphlus 240

faerorensis Born, 1913, Carabus 168

fageli Mateu, 1956, Mesolestes 421

fageticola Deuve, 1994, Carabus 126

fagnensis Grotz, 1978, Carabus 171

fagniezi Chobaut, 1902, Duvalius [Trechus] 301

fagniezi Jeannel, 1937, Laemostenus [Ceuthosphodrus] 535

fagniezi Jeannel, 1937, Nebria 93

fagniezi Jeannel, 1937, Scotodipnus 240

fagniezi Colas & A. Gaudin, 1935, Trechus 22, 327

faillae Ragusa, 1884, Chlaenius 351

faillai Magrini & Vanni, 1984, Duvalius 301



faillai Reitter, 1887, Zuphium 574

fairmaireanus Csiki, 1930, Pterostichus 512

fairmairei C. G. Thomson, 1875, Carabus 180

fairmairei Leprieur, 1876, Deronectes [Hydroporus] 58

fairmairei Régimbart, 1883, Gyrinus 27

fairmairei Schaufuss, 1861, Laemostenus [Sphodrus] 535

fairmairei Sciaky, 1995, Myas [Trigonognatha] 476

fairmairei Ledoux & Roux, 1992, Nebria 92

fairmairei Desbrochers des Loges, 1906, Pterostichus 512

fairmairei Chaudoir, 1868, Tapinopterus [Feronia] 519

fairmairei Pandellé, 1867, Trechus 330

falciger Heinz & Ledoux, 1990, Ledouxius [Deltomerus] [DA] 284

falcipenis Zaballos & Jeanne, 1987, Geocharis 239

falcozi Guignot, 1932, Agabus 39

falcozi Netolitzky, 1918, Bembidion 250

falcus Jedlika, 1935, Platynus [Colpodes] 464

faldermanni Zaitzev, 1927, Agabus 39

faldermanni Dejean, 1829, Carabus 135

faldermanni Gistel, 1838, Cymindis 24, 415

faldermanni Ménétriés, 1832, Nebria 86

faldermanni Schatzmayr, 1929, Pterostichus 493

falkenstromi J. Balfour‑Browne, 1944, Agabus 41

falkenstromi Zaitzev, 1953, Agabus [Gaurodytes] 36

falkenstromi Gschwendtner, 1939, Hyphydrus 74



fallaciosa J. Müller, 1946, Amara [Zabrus] 565

fallaciosa W. Horn, 1897, Cylindera [Cicindela] 111



fallaciosa Antoine, 1939, Cymindis 414

fallaciosa Ledoux & Roux, 1992, Nebria 88



fallaciosum J. R. Sahlberg, 1913, Bembidion 263

fallaciosus J. Müller, 1914, Anophthalmus [Trechus] 291



fallaciosus Guignot, 1932, Graptodytes 59

fallaciosus Jeannel, 1921, Trechus 22, 338

fallax Semenov, 1889, Acalathus 529



fallax Munster, 1932, Agabus 37

fallax A. Morawitz, 1862, Agonum [Anchomenus] 454



fallax Apfelbeck, 1904, Agonum [Platynus] 452

fallax LeConte, 1848, Amara 549

fallax Apfelbeck, 1904, Brachinus 213

fallax Rost, 1892, Carabus [Plectes] 132

fallax Sokolá, 1910, Carabus 181

fallax Peyerimhoff, 1929, Eremolestes [Lionychus] 431



fallax Peyron, 1858, Parophonus [Harpalus] 393

fallax Carret, 1903, Pogonus 287

fallettianus Cavazzuti, 1997, Carabus [Procerus] 185

fallettii Deuve, 1993, Carabus 142

fallettii Sciaky, 1996, Pterostichus 506



falsarius Reitter, 1896, Carabus 130

falsicolor Fairmaire, 1886, Pristosia [Calathus] 531



falsinflatus J. Ochs, 1967, Carabus 168

falsulus Antoine, 1941, Microlestes 422

falsum Puel, 1938, Agonum 449

falsus Bates, 1876, Trichotichnus 395

famelica C. Zimmermann, 1832, Amara 549

famelicus Tschitschérine, 1898, Harpalus 376

familiaris Duftschmid, 1812, Amara [Carabus] 549

faminii Dejean, 1826, Carabus 152

faminii Dejean, 1826, Platytarus [Cymindis] 410



fanaticus Casey, 1924, Harpalus 379

fanensis Kataev & Wrase, 1993, Harpalus 376

fangensis Biström & Satô, 1988, Hyphydrus 74

fani Imura, 1993, Carabus 189

fanianus Imura, 1993, Carabus 150

fantingi Imura & Su, 1998, Carabus 121

farai Jedlika, 1963, Coptodera 436



farai Jedlika, 1951, Lebia 430

farctus C. Zimmermann, 1831, Zabrus [Polysitus] 572



farellensis Beuthin, 1888, Cicindela 106

farellensis Graëlls, 1847, Cicindela 103

farineli P. Machard, 1975, Carabus 148

farinesi Dejean, 1826, Carabus 139

farinosae Zaballos & Ruiz‑Tapiador, 1997, Typhlocharis 241

farkaci Deuve, 1995, Agonotrechus [Bhutanotrechus] 289

farkaci Toledano & Sciaky, 1998, Bembidion 265


farkaci Kataev & Wrase, 1995, Harpalus 376

farkaci Sciaky, 1994, Leistus 80

farkaci Ledoux & Roux, 1996, Nebria 88

farkaci Sciaky, 1997, Pterostichus 514

farkaci Sciaky, 1998, Stomis 517

farkaci Sciaky, 1996, Straneostichus 518



faroensis Mjöberg, 1917, Hydroporus 61

farsense Marggi & Huber, 1999, Bembidion 271

farsica Dostal & Zettel, 1999, Axonya 235

fasciata Geoffroy, 1785, Cicindela [Buprestis] 105

fasciatopunctata C. A. Dohrn, 1886, Cicindela 105

fasciatopunctata Germar, 1844, Cicindela 107

fasciatopunctata L. Miller, 1850, Nebria 86

fasciatopunctatus A. Morawitz, 1862, Pogonus 287

fasciatopunctatus Creutzer, 1799, Pterostichus [Carabus] 512

fasciatum Chaudoir, 1846, Bembidion 249

fasciatum Motschulsky, 1844, Bembidion [Notaphus] 254

fasciatum Petri, 1912, Bembidion 248

fasciatum Solsky, 1874, Bembidion 262

fasciatus DeGeer, 1774, Acilius [Dytiscus] 49

fasciatus Motschulsky, 1844, Agatus 408

fasciatus Fabricius, 1781, Craspedophorus [Pimelia] 446

fasciatus Mäklin, 1855, Notiophilus 97

fasciatus Paykull, 1790, Philorhizus [Carabus] 426

fasciatus Aubé, 1838, Rhithrodytes [Hydroporus] 68

fasciatus Fabricius, 1775, Sandracottus [Dytiscus] 49

fasciatus Motschulsky, 1851, Tachys [Trechus] 275

fascifer Reitter, 1894, Metadromius [Metabletus] 421

fascigerum Chaudoir, 1852, Cymbionotum [Coscinia] 234



fasciipeinnis Küster, 1846, Cymindis 412

fasciola Bates, 1886, Dolichoctis 436

fasciolata Schilsky, 1888, Cymindis 417

fasciolatum Duftschmid, 1812, Bembidion [Elaphrus] 245

fasciolatus P. Rossi, 1790, Polistichus [Carabus] 574

fassatii Jedlika, 1951, Bembidion 271

fassatii Kult, 1949, Dyschirius 225

fassatii Jedlika, 1951, Lebia 430

fassatii Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 540

fassatii Maan, 1947, Typhlotrechus 344

fastidiosa Dejean, 1825, Myriochila [Cicindela] 113

fastidiosum Jacquelin du Val, 1852, Bembidion 258

fastidiosus Mannerheim, 1853, Pterostichus [Cryobius] 490

fastiditus Dejean, 1829, Harpalus 379

fastuosior Deuve, 1990, Carabus 123

fastuosus Palliardi, 1825, Carabus 151



fastuosus Faldermann, 1836, Harpalus 374

fatnickensis Absolon & Maan, 1947, Neotrechus 316

fatrensis Hrka & Pulpán, 1992, Nebria 87

fatrica Roubal, 1922, Amara 550

fatricus Roubal, 1930, Duvalius 307

faucium Andrewes, 1930, Amara 557

fauconneti Reitter, 1899, Carabus [Plectes] 131

fauconneti Pic, 1930, Stictonectes [Deronectes] 69

faunulus Kurnakov, 1972, Carabus 131

faunus Kurnakov, 1972, Carabus 131

faunus Kurnakov, 1962, Pterostichus 504

fausti Reitter, 1888, Amara [Harpalodema] 562

fausti C. A. Dohrn, 1873, Carabus 169

fausti Heyden, 1887, Diacheila 206

fausti Fancello, 1988, Typhloreicheia 223

faustus Brullé, 1836, Carabus 176

faustus Jeannel, 1927, Trechus 330

fauveli Ganglbauer, 1891, Bembidion 257



fauvelii Mathan, 1862, Ophonus 389

favieri Fairmaire, 1859, Carabus 152



favieri Lucas, 1859, Chlaenius 355

favieri Fairmaire, 1859, Cymindis 413

favieri Fairmaire, 1868, Laemostenus [Sphodrus] 539

favieri Fairmaire, 1851, Paussus 210

favonius Uéno, 1971, Masuzoa 314

fayardensis P. Machard, 1974, Carabus 130

feae Bates, 1892, Galerita 359



feana Bates, 1891, Tachyura [Tachys] 278

federicii Born, 1899, Carabus [Platycarabus] 184

fedjedjensis Mayet, 1887, Microlestes [Dromius] 424

fedorenkoi Zamotajlov, 1998, Diplous 283

fedorovi Kryzhanovskij, 1994, Andrewesius 456

fedrenkoi Kataev & N. Ito, 1999, Trichotichnus 394

fedtschenkoi Tschitschérine, 1898, Amara 551


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