Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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graecus Heyden, 1887, Poecilus 482

graecus Apfelbeck, 1904, Pogonistes [Pogonus] 286

graecus Dejean, 1828, Zabrus 571

graellsi Gemminger & Harold, 1868, Platambus [Agabus] 44

grafi Reitter, 1913, Badister 441


grafittii Casale & Giachino, 1988, Sardaphaenops 319

grahami Hatch, 1951, Bembidion 247

grahamianus Mandl, 1975, Carabus 175

graja K. Daniel & J. Daniel, 1898, Amara 563

grajus Jeannel, 1937, Binaghites [Scotodipnus] 239



grajus Reitter, 1896, Carabus 184

grajus K. Daniel & J. Daniel, 1898, Cychrus 204

grajus Dejean, 1828, Pterostichus [Feronia] 507

grajus Jeannel, 1937, Sphodropsis 543

grallatorius Roeschke, 1921, Carabus [Coptolabrus] 122

grammicus Germar, 1827, Hydaticus [Dytiscus] 53

grammophora Chaudoir, 1852, Cylindera [Cicindela] 110

granaria Dejean, 1828, Amara 556

granatensis Vuillefroy, 1866, Calathus 525

granatensis J. Vives & E. Vives, 1982, Laemostenus [Ceuthosthenes] 533

grancanariensis Emden, 1928, Harpalus 370

grancanariensis Machado, 1975, Licinus 442

grancanariensis Machado, 1992, Paradromius 424

grandiceps Bates, 1873, Bradycellus [Tachycellus] 401

grandiceps Reitter, 1900, Eriophonus [Harpalus] 370

grandiceps Bates, 1892, Kareya [Platymetopus] 387

grandiceps Kurnakov, 1961, Lindrothius 529



grandiceps Ganglbauer, 1889, Molops 473

grandiceps Piochard de la Brûlerie, 1876, Platyderus 522

grandiceps Reitter, 1885, Trechus 331

grandicolle Motschulsky, 1850, Bembidion [Philochthus] 264

grandicollis Fairmaire & Laboulbène, 1854, Abax [Feronia] 470

grandicollis C. Zimmermann, 1832, Amara 557

grandicollis Reiche, 1862, Harpalus 375

grandicollis Sturm, 1843, Laemostenus [Pristonychus] 537

grandicollis Audinet‑Serville, 1821, Percus [Harpalus] 478

grandicollis Uéno & Yin, 1993, Perileptus 289

grandicollis Motschulsky, 1850, Platynus [Anchomenus] 462

grandicollis Chaudoir, 1846, Tachyura [Tachys] 277

grandicollis Uéno, 1980, Trechiama 322

grandinella Semenov, 1890, Merizomena [Glycia] 409

grandipenne Schaum, 1862, Bembidion 259



grandis Faldermann, 1836, Acinopus 369

grandis Andrewes, 1937, Apristus 431



grandis Bates, 1883, Carabus 122

grandis Belousov, 1998, Cimmerites 297

grandis Klug, 1834, Dineutus [Gyrinus] 26

grandis Zamotajlov, 1998, Diplous 283

grandis Uéno, 1998, Dongodytes 298

grandis Decou, 1961, Duvalius 300


grandis Knirsch, 1913, Duvalius [Anophthalmus] 299

grandis Laporte, 1835, Hyphydrus 74

grandis Uéno, 1973, Kurasawatrechus 313

grandis Motschulsky, 1850, Laemostenus [Pristonychus] 537

grandis Peyron, 1858, Loxoncus [Stenolophus] 404

grandis Dejean, 1826, Pseudoclivina [Clivina] 221



grandis Dejean, 1828, Pterostichus [Cophosus] 488

grandis Fabricius, 1798, Scarites 233

grandis Fischer von Waldheim, 1828, Scarites 233

grandis Motschulsky, 1844, Taphoxenus [Sphodrus] 544

grandis Ganglbauer, 1891, Trechus 332

grandistyle J. Schmidt, 1996, Xestagonum 468

grandisubtilis W. Horn, 1935, Neocollyris [Collyris] 116



grandivadosa W. Horn, 1935, Neocollyris [Collyris] 116

graniger Palliardi, 1825, Carabus 196



granjanensis Wagner, 1932, Chlaenius 352

granosus Chaudoir, 1844, Carabus 153

granularis Linné, 1767, Graptodytes [Dytiscus] 59

granulata Gebler, 1843, Cicindela 105

granulatocostatus Mandl, 1965, Carabus 162

granulatum Andrewes, 1925, Asaphidion 242

granulatus Falkenström, 1936, Agabus [Gaurodytes] 36

granulatus Linné, 1758, Carabus 136

granulatus Motschulsky, 1850, Cychrus 202

granulatus Dejean, 1826, Licinus 442

granulatus Régimbart, 1883, Noterus 34

granulipennis Apfelbeck, 1918, Aptinus 212

granulipennis Balkenohl, 1994, Haplogaster 232



granulisparsus Bates, 1889, Carabus 155

granulosum Motschulsky, 1844, Calosoma 120

granulum Régimbart, 1895, Bidessus 55

granulum Motschulsky, 1855, Hydroscapha [Limnebius] 25



grapeioides Munster, 1930, Bembidion 272

grapii Gyllenhal, 1827, Bembidion 271

grapii Gyllenhal, 1808, Rhantus [Dytiscus] 45

grassator Coquerel, 1859, Acinopus 368

grassator Andrewes, 1932, Platynus [Anchomenus] 462

grasselyi Antoine, 1941, Carabus 152

gratiae Bucciarelli, 1978, Boldoriella 296

gratiosa Tschitschérine, 1898, Aristochroa 471

gratiosa Ledoux & Roux, 1998, Nebria 92

gratiosum Mannerheim, 1853, Agonum [Anchomenus] 453

gratiosus Mannerheim, 1844, Callistus 347



gratiosus Chaudoir, 1837, Chlaenius 350

gratus Semenov, 1887, Carabus 134



gratus Faldermann, 1836, Laemostenus [Pristonychus] 538

gratus Chaudoir, 1859, Poecilus [Feronia] 481

gratus Jedlika, 1962, Poecilus [Pterostichus] [HN, repl. in vol. 2] 482

gravidula Rosenhauer, 1856, Amara 555

gravidus Lindroth, 1956, Synuchus 545

gravidus Putzeys, 1870, Trechus 332

gravis Jedlika, 1939, Pterostichus 501

gravis Dejean, 1828, Zabrus 569

grayii Wollaston, 1862, Sirdenus [Pogonus] 288

grazianii Schatzmayr, 1937, Cymindis 412

graziolii Cavazzuti, 1987, Carabus 147

graziosii Deuve, 1995, Carabus 150

gredensis Fery, 1999, Hydroporus 60

gredosana Jeanne, 1985, Cymindis 412

gredosanum Jeanne, 1974, Bembidion [Ocydromus] 245

gredosanus Lauffer, 1905, Chlaenius 352

gredosanus Jeanne, 1970, Zabrus 569



greffi A. Gaudin, 1947, Aphoenops 295

gregarius Fauvel, 1897, Nesarpalus [Harpalus] 388

gregarius Reiche, 1862, Platyderus 522

gregarius Chaudoir, 1846, Tachys 276

gregoryi Jeannel, 1937, Deuveotrechus [Stevensius] 298

grenieri Pandellé, 1867, Trechus 332

gressittianus Mandl, 1975, Carabus 156

gressoria Chaudoir, 1871, Lebia 428

gressorius Dejean, 1828, Poecilus [Feronia] 483

gridellii Schatzmayr, 1912, Agonum 451

gridellii Baliani, 1928, Amara 555

gridellii J. Müller, 1931, Anophthalmus 291

gridellii De Monte, 1947, Bembidion 258

gridellii Breuning, 1959, Carabus 155

gridellii Focarile, 1960, Hydroporus 61

grignensis Monzini, 1987, Boldoriella 296



grignensis Born, 1902, Carabus 184

grigorjevi Kryzhanovskij, 1953, Carabus 146

grilli J. Schmidt & Hartmann, 1998, Deltomerodes 281

grinevi Lutshnik, 1935, Pogonistes 286

griseoides Reitter, 1900, Ophonus 390

griseostriatus DeGeer, 1774, Stictotarsus [Dytiscus] 69



griseus Fabricius, 1781, Eretes [Dytiscus] 53

griseus Panzer, 1796, Harpalus [Carabus] 385

grisvardi De Wailly, 1949, Bembidion [Peryphus] 253

grmecensis Born, 1910, Carabus 163

groenlandicus Aubé, 1838, Colymbetes 45

grombczewskii Semenov, 1891, Carabus 153



grombczewskii Semenov, 1891, Nebria 90

grombczewskyi Tschitschérine, 1890, Poecilus [Feronia] 484

grosclaudei Normand, 1940, Bembidion 245

grososae Deuve & Remond, 1998, Carabus 188

grososae Deuve, 1995, Cychrus 203

grossefoveatus G. Hauser, 1913, Carabus [Apotomopterus] 124



grossegranulatus Breuning, 1943, Carabus 137

grossepunctata W. Horn, 1935, Protocollyris [Collyris] 117

grossepunctatus Vorbringer, 1907, Bidessus 55

grossepunctatus A. Fleischer, 1899, Dyschirius 229

grossi Jeannel, 1928, Neotrechus 315



grossus O. F. Müller, 1776, Hyphydrus [Dytiscus] 74

grossus Schaufuss, 1865, Taphoxenus 544

grottagliensis Schauberger, 1927, Ophonus [Harpalus] 388

grottoi Daffner, 1992, Orotrechus 317

groubei Antoine, 1935, Antoinella [Duvalius] 293

grouvellei Jeannel, 1913, Trechus 334

gruardeti Pater, 1938, Ophonus [Metophonus] 391

grumi Tschitschérine, 1894, Amara 562

grumi Semenov, 1900, Calosoma 119

grumi Tschitschérine, 1903, Cicindela 104

grumi Tschitschérine, 1899, Dicheirotrichus [Trichocellus] 403

grumi Glasunov, 1902, Nebria 90

grumi Tschitschérine, 1898, Poecilus [Feronia] 479

grummi Zamotajlov & Kryzhanovskij, 1990, Diplous 283

grummulifer Deuve, 1994, Cychrus 204

grumorum Semenov, 1898, Carabus 142

grumulifer Deuve, 1993, Carabus 144



grundmanni Mandl, 1955, Carabus 144

gryphus Motschulsky, 1844, Carabus 174

gschwendtneri Guignot, 1942, Hyphydrus 74

gschwendtneri Schauberger, 1921, Pterostichus [Platypterus] 508

gspani Knirsch, 1926, Anophthalmus [Haplotrechus] 291

gspani Reitter, 1918, Anophthalmus 292

guadarramense Gautier des Cottes, 1866, Bembidion 264

guadarramensis Wagner, 1932, Chlaenius 352



guadarramensis Graëlls, 1847, Cicindela 106

guadarramus LaFerté‑Sénectère, 1847, Carabus 179

guadarramus Ehlers, 1883, Microtyphlus 240

guangdongicus Deuve, 1992, Carabus 127

guangxicola Deuve, 1998, Carabus 142

guangxicus Deuve, 1989, Carabus 125

guanmenshanus Imura, 1991, Carabus 183

guanpingae Korell & Richter, 1996, Carabus 190



guardoensis Pierre Meyer & Mollard, 1996, Carabus 167

guareschii Moscardini, 1950, Duvalius 303

gudenzii Neri, 1981, Bembidion [Ocydromus] 256



guedeli Sirguey, 1931, Carabus 139

guedeli Sainte‑Claire Deville, 1902, Pterostichus [Platysma] 503



gueneycensis atloš & Moravec, 1999, Carabus 186

guentheri J. R. Sahlberg, 1887, Amara 550

guentheri Seidlitz, 1887, Bembidion 247

guentheri Maan, 1943, Lebia 427



guentheri Sturm, 1824, Pterostichus [Platysma] [NO] 511

gueorguievi Wewalka, 1975, Hydroporus 61



guergentepeensis Blumenthal, 1975, Carabus 193

guerini Fischer von Waldheim, 1842, Carabus 123

guerini Aubé, 1838, Cybister 50

guerini Gaubil, 1844, Porotachys [Bembidium] 274

guerini Dejean, 1826, Scapterus 231

guernei Régimbart, 1891, Hydroporus 61

guerrouchensis Bruneau de Miré, 1958, Laemostenus [Antisphodrus] 534

guerryanus Breuning, 1961, Carabus 129



guerryi Olsoufieff, 1925, Acilius 48

guerryi Born, 1903, Carabus 143



guerryi Born, 1907, Carabus 139

guglielmofonti Deuve & Font, 1999, Carabus 141

guibeicus Deuve & Tian, 1999, Carabus 125

guidongicus Cavazzuti & Ratti, 1998, Carabus 142



guignoti Vazirani, 1970, Hydronebrius 41

guignoti Gschwendtner, 1935, Hydroporus 60

guignoti Vazirani, 1965, Platambus 44

guildensis Alluaud, 1932, Zabrus 572

guinanensis Deuve, 1991, Carabus 135

guineensis Aubé, 1838, Herophydrus [Hyphydrus] 71

guineensis Dejean, 1831, Scarites 233

guiraoi Perez Arcas, 1869, Percus 479

guizhouensis Cavazzuti, 1994, Carabus 125

guizhouensis Sciaky, 1997, Pterostichus 500

guizhouicus Deuve, 1994, Carabus 143

gujoensis Uéno, 1974, Kurasawatrechus 313



gulickai Schweiger, 1970, Leistus 18, 83

gulickai Löbl, 1967, Trechus 332



gumischchanense Eichler, 1924, Bembidion 251

gundermanni Hubenthal, 1908, Pterostichus 488

gurjevae Kabak, 1999, Cymindis 416

gurjevae Kabak, 1994, Poecilus 480

gurkhaorum Lassalle, 1985, Carabus 162

gurungi J. Schmidt, 1998, Trechus 332

gurwani Jedlika, 1968, Bembidion 262



gusevi Zamotajlov & Koval', 1989, Carabus 137

gusevi Belousov & Zamotajlov, 1988, Deltomerus 282

gusevi Belousov, 1989, Duvalius 303

gusevi Belousov, 1990, Trechus 332

gushensis Belousov & Kabak, 1998, Trechus 332

gussakovskii Kryzhanovskij, 1971, Carabus 153



gustavi Shilenkov, 1996, Carabus 170

gustavi Fagniez, 1930, Geotrechus 310

gustavii Crotch, 1871, Dicheirotrichus 402

gustavlohsei Löbl, 1995, Sphaerius [Microsporus] 26



gutiiensis Takács & Lie, 1992, Carabus 172

guttata Wiedemann, 1823, Cicindela 103

guttatus Paykull, 1798, Agabus [Dytiscus] 40

guttatus Bates, 1873, Amblystomus 360

guttatus Aubé, 1836, Haliplus 32

guttiger Gyllenhal, 1808, Ilybius [Dytiscus] 42



guttigera Reiche & Saulcy, 1855, Tachyura [Bembidium] 279

guttula Chaudoir, 1869, Abacetus 347



guttula Audinet‑Serville, 1821, Bembidion 263

guttula Duftschmid, 1812, Bembidion [Elaphrus] 248

guttula Fabricius, 1792, Bembidion [Carabus] 264



guttula Chaudoir, 1842, Brachinus 214

guttula Chaudoir, 1856, Callistoides [Chlaenius] 348

guttulata Ledoux & Roux, 1999, Nebria [add] 89

guttulatum Chaudoir, 1850, Bembidion 267

guttuliforme Deuve, 1985, Broscosoma 235

guttuloides De Monte, 1953, Bembidion 264



guttulus Schönherr, 1808, Agabus [Dytiscus] 38

guttulus Régimbart, 1899, Platambus [Platynectes] 43

gutturosus Buquet, 1840, Acinopus 368

guyicolasi Deuve & Simard, 1976, Carabus 181

guzelolukensis Lassalle, 1997, Laemostenus 534

guzhangensis Kleinfeld, 1998, Carabus 125

gvalijai Retezár, 1992, Carabus 160

gvozdevae Obydov, 1997, Carabus 199

gwendyddae Morvan, 1982, Laemostenus 536

gwervaenus Morvan, 1995, Pterostichus 515

gwiomarchi Morvan, 1982, Trechus 332

gyaisiensis Zamotajlov & Sciaky, 1999, Chinapenetretus 281

gyalicaensis Horvatovich, 1975, Nebria 86



gyleki Breit, 1911, Duvalius [Anophthalmus] 300

gyllenhali C. G. Thomson, 1859, Acupalpus [Stenolophus] 399

gyllenhali Schönherr, 1806, Nebria [Carabus] 88

gyllenhalii Dejean, 1825, Callytron [Cicindela] 101

gyllenhalii Fischer von Waldheim, 1828, Carabus 164


gyllenhalii Schiødte, 1841, Hydroporus 61

gyllenhalii Laporte, 1840, Rhantus [Colymbetes] 46

gyoerffyi Csiki, 1927, Carabus 143

gyotokuensis Tanaka, 1956, Tachys 275



gyrinoides Aubé, 1863, Hydroscapha [Limnebius] 25

gyrosus Motschulsky, 1866, Pterostichus [Argutor] 507



haagi Wehncke, 1876, Cybister [Trogus] 50

haagi Heyden, 1870, Dixus [Aristus] 365

habaensis Deuve, 1992, Carabus 126

habaensis Deuve & Mourzine, 1998, Cychrus 204

habashanicola Farka, 1999, Leistus 80

habei Uéno, 1954, Trechiama [Trechus] 322

habelmanni Schilsky, 1888, Calosoma 120

habelmanni Wehncke, 1876, Hydroporus 64

haberfelneri Ganglbauer, 1891, Carabus 181

haberhaueri Heyden, 1889, Nebria 90

haberhaueri Heyden, 1889, Platyderus 521



habroterus Antoine, 1941, Harpalus 373

habui Jedlika, 1965, Bembidion 265



habui Jedlika, 1962, Pterostichus 494

habui Jedlika, 1958, Trichotichnus 397

habui J. Schmidt, 1996, Xestagonum 468



hachlafensis Fongond, 1983, Amara 567

hacquetii Sturm, 1853, Typhlotrechus [Anophthalmus] 344



hadenia Gistel, 1838, Cicindela 107

haeckeli Farka, 1995, Archastes 79

haeckeli Bezina & Imura, 1997, Carabus 141

haeckeli Deuve, 1997, Cychrus 204

haeckeli Farka, 1995, Leistus 80

haeckeli Sciaky & Wrase, 1997, Straneostichus 518



haeffneri Aubé, 1837, Agabus 38

haekeli Ledoux & Roux, 1995, Archastes 79

haematomerus Kraatz, 1878, Carabus 136

haematomerus Kraatz, 1878, Carabus 196

haematopus Csiki, 1930, Pterostichus 497

haematopus Dejean, 1831, Stereocerus [Feronia] 517



haemorhoidalis Motschulsky, 1864, Eucolliuris [Casnonia] 443

haemorrhoidale Stephens, 1835, Bembidion [Lopha] 248

haemorrhoidalis Fabricius, 1787, Copelatus [Dytiscus] 47



haemorrhoidalis Fabricius, 1787, Lebia [Carabus] 429

haemorrhoum Stephens, 1828, Bembidion [Philochthus] 264

haeneli Netolitzky, 1918, Bembidion 265

haeres Fischer von Waldheim, 1823, Carabus 199

haerii Heinz, 1973, Carabus 193

haesitatus Fairmaire, 1888, Pterostichus 501



hafezi Morvan, 1975, Poecilus [Ancholeus] 479

hafniensis Gmelin, 1790, Pterostichus [Carabus] 487

hagai Takami & Ishikawa, 1997, Carabus 176

hagai Mateu, 1992, Dromius 420

hagai Nemoto, 1989, Odacantha 444

hagenbachii Sturm, 1824, Pterostichus 512

hahni Mandl, 1980, Carabus 158

hahni Reitter, 1908, Oodes 445

hainanensis Deuve & Tian, 1999, Carabus 142

hainanensis Tian & Chen, 1997, Dischissus 447

hainanensis Hendrich, 1998, Lacconectus 48

hainanensis Wewalka, 1997, Microdytes 75

hainanensis Nilsson, 1998, Platynectes 44

haisishanicus Farka & Sciaky, 1998, Leistus 80

hajeki Kobmann, 1925, Carabus 185

hajeki Reitter, 1913, Trechus 332

hakkariensis Wewalka, 1989, Deronectes 58

hakonum Harold, 1878, Nipponagonum [Colpodes] 460

hakusanus Nakane, 1961, Carabus 159

hakusanus Kasahara, 1989, Pterostichus 494



halberti Bullock, 1928, Haliplus 30

halbherri Lapouge, 1898, Carabus 151

halensis Schaller, 1783, Dolichus [Carabus] 530

halensis Fabricius, 1787, Scarodytes [Dytiscus] 68

halfeticus Korell, 1999, Carabus 187

haligena Wollaston, 1860, Orthomus [Pterostichus] 477

haliploides Bates, 1892, Elaphropus 273

hallasanensis Kwon & Lee, 1984, Carabus [Scambocarabus] 191

halmai Apfelbeck, 1904, Neotrechus [Trechus] 316



halophilus Fauvel, 1890, Dyschirius 226

halophilus Nicolai, 1822, Pogonus 287

hamadaensis Uéno, 1958, Trechiama 324

hamanense Jedlika, 1933, Bembidion 265

hamarensis Shilenkov, 1996, Carabus 170



hamasiensis J. Müller, 1947, Harpalus 370

hamatum Kolenati, 1845, Bembidion [Notaphus] 253

hamatus Fischer von Waldheim, 1828, Brachinus 215

hamatus Dejean, 1831, Chlaenius 354

hamatus Uéno, 1990, Trechiama 322



hamatus Jeannel, 1927, Trechus 331

hamburgense Meier, 1899, Bembidion 257

hamifasciata H. Kolbe, 1886, Cicindela 107

hamifer Chaudoir, 1856, Chlaenius 354

hamifera Baehr, 1998, Gunvorita 573



hamilcaris Bernau, 1914, Carabus 184

hammarstroemi Poppius, 1908, Amara 549

hammarstroemi Poppius, 1905, Bembidion 247

hammeri Mandl, 1967, Carabus 201

hampei Küster, 1846, Carabus 172

hampei Gestro, 1875, Cychrus 204

hampei Ganglbauer, 1891, Trechus 332

hamulatus Gyllenhal, 1813, Hydroglyphus [Hyphydrus] 56

hamus Basquin & Legrand, 1991, Carabus 157



hanae Chu, 1967, Carabus 148

hanae Hrka, 1990, Duvalius 308

hanatanii Imura, 1991, Carabus 164

handelmazzettii Mandl, 1955, Carabus 190

hangaicus Shilenkov, 1982, Trechus 332


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