Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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hangtschouensis G. Hauser, 1919, Carabus [Coptolabrus] 143

hanhaica Tschitschérine, 1894, Amara 552

hanhaicus Tschitschérine, 1894, Poecilus [Feronia] 480

hannemanni Hieke, 1991, Amara 556

hanseatica W. Horn, 1927, Salpingophora [Cicindela] 114

hansi Jedlika, 1932, Bembidion 263

haptoderoides Tschitschérine, 1889, Pterostichus [Eurythorax] 494

haraldi Håkan Lindberg, 1933, Agabus 36

haraldi Daffner, 1990, Orotrechus 317

haraldianus Daffner, 1992, Anophthalmus 291

haraldmuelleri Mandl, 1979, Carabus 162

harambureae Breuning & Ruspoli, 1974, Carabus 147

haranti Verdier, Quézel & Rioux, 1951, Microlestes 423

harbinensis Straneo, 1942, Pterostichus 507

harcyniae Sturm, 1815, Carabus 168

hardei Heinz, 1990, Carabus 193

hardwickii Hope, 1831, Metacolpodes [Colpodes] 460

hardwickii Hope, 1831, Paussus 210



hardyi Chaudoir, 1861, Miscodera 237

hariensis Machado, 1992, Paradromius 424

harmandi Lapouge, 1909, Carabus 149

harmandi Tschitschérine, 1900, Lesticus 471

harmandi Tschitschérine, 1900, Pterostichus [Platysma] 510

harminae Vondel, 1990, Haliplus 30

haroldii Schaum, 1863, Acinopus [Acmastes] 368

harpagon Farka, 1998, Leistus 80

harpalinus Audinet‑Serville, 1821, Bradycellus [Trechus] 401

harpaloides A. Fleischer, 1897, Amara 561

harpaloides Dejean, 1828, Amara 560



harpaloides Lapouge, 1925, Carabus 178

harpaloides Audinet‑Serville, 1821, Ocys [Bembidion] 272

harpaloides Wollaston, 1864, Paraeutrichopus [Pterostichus] 521


harponifer Tanaka, 1987, Pterostichus 513

harrarense Alluaud, 1918, Metagonum [Agonum] 460



harrisoni Baliani, 1934, Amara 555

hartmanni Hieke, 1997, Amara 564

hartmanni Sciaky & Facchini, 1997, Microzargus 439

hartmanni J. Schmidt, 1995, Pterostichus 510



hartmannorum Schmid, 1966, Arctaphaenops 296

hasandaghensis Darge & Basquin, 1990, Carabus 193

hasegawai Habu, 1975, Agonum [Platynus] 450

hasegawai Satô, 1988, Copelatus 48

hasemiya Morita, 1990, Apatrobus 280

hashimotoi Uéno, 1961, Trechus 325

hashinaukamui Nakajima, 1993, Carabus [Damaster] 122

haslundi Emden, 1954, Pseudotaphoxenus [Taphoxenus] 541

hassani Antoine, 1933, Bembidion 244

hassani Antoine, 1941, Ophonus [Harpalus] 389

hastatus Andrewes, 1931, Syntomus [Metabletus] 432

hastii C. R. Sahlberg, 1827, Bembidion 246

hasurada Andrewes, 1924, Bembidion 255

hattusae Franzen, 1996, Cephalota 102

hauckei P. Moravec & Lompe, nom. nov., Anophthalmus 22, 291

hauckei Ganglbauer, 1913, Typhlotrechus [Trechus] 344

haupti Reitter, 1908, Bembidion 258



hauryi Géhin, 1885, Carabus 123

hauseri Schauberger, 1929, Anisodactylus 361

hauseri A. Fleischer, 1914, Anthracus 400

hauseri Jedlika, 1958, Bembidion 268

hauseri Lapouge, 1925, Carabus 201

hauseri Reitter, 1894, Carabus 145



hauseri Jedlika, 1931, Chlaenius 354

hauseri Jedlika, 1931, Colpoides 457



hauseri Reitter, 1894, Cymindis [Dromius] 415

hauseri Reitter, 1894, Dicheirotrichus [Trichocellus] 403



hauseri Jedlika, 1931, Dromius 420

hauseri A. Fleischer, 1898, Dyschirius 225



hauseri Schauberger, 1928, Harpalus 372

hauseri Jedlika, 1933, Myas [Trigonognatha] 476

hauseri W. Horn, 1898, Myriochila [Cicindela] 113

hauseri Spaeth, 1900, Notiophilus 97

hauseri Schauberger, 1933, Parophonus 392

hauseri Jedlika, 1931, Pristosia [Kanoldia] 531

hauseri Jedlika, 1933, Pseudotaphoxenus [Taphoxenus] 541

hauseri Reitter, 1899, Pterostichus [Lyperosomus] 499

hauseri Jeannel, 1962, Trechus 332

hauseri Jedlika, 1931, Xestagonum [Colpodes] 468


hauseri Reitter, 1895, Zuphium 574

hauserianum Netolitzky, 1918, Bembidion 253



hauserianus Born, 1922, Carabus [Damaster] 148

hauserianus Reitter, 1896, Carabus 192

hauserianus Lorenz, 1998, Dromius 420

hauserianus Schauberger, 1929, Harpalus 385

hauserianus Casale, 1988, Taphoxenus 544

havelkai Fassati, 1955, Bembidion 261

havelkai Kult, 1949, Poecilus [Pterostichus] 484

hayachinense Nakane, 1979, Bembidion [Bembidion] 259

hayachinensis Nakane, 1968, Apatrobus [Patrobus] 280

hayakawai Morita, 1997, Trichotichnus 396

hayashii Uéno, 1977, Epaphiopsis 309

hayashii Uéno, 1999, Ishikawatrechus 312

hayashii N. Ito, 1999, Trichotichnus 394

hazara Andrewes, 1935, Bembidion 261

hazardi Andrewes, 1930, Brachinus 215

hazarensis Deuve, 1982, Carabus 154

hearseyanus Westwood, 1842, Paussus 210

hebaueri Fery & Hosseinie, 1998, Deronectes 58

hebaueri Hendrich, 1990, Hydroporus 61

hebraicus Reiche & Saulcy, 1855, Brachinus 214

hebraicus Geoffroy, 1785, Platambus [Dyticus] 44

hector G. Hauser, 1924, Carabus [Apotomopterus] 123

hedini Heinz, 1973, Carabus 194

hedini Zaitzev, 1908, Cybister 51

hedini Liebke, 1935, Cymindis 417

hedini Andrewes, 1935, Holosoma [Chlaenius] 444

hedini Schauberger, 1936, Trichotichnus 394

hedjazica Hieke, 1988, Amara 564

hedkeinekus Morvan, 1995, Pterostichus 515

hedwigae Reitter, 1897, Hydroporus 62

heegeri Dejean, 1826, Nebria 88

heeri K. Daniel, 1903, Nebria 93

heeri J. Jacquet, 1925, Stictotarsus [Hydroporus] 69

heeri Tournier, 1870, Trechus 331

heerianus Géhin, 1876, Carabus 184

heerii Kelch, 1846, Bembidion 259

heerii Germar, 1836, Carabus 184

heerii Heer, 1837, Pterostichus 508

hegeduesii J. Frivaldszky, 1880, Duvalius [Anophthalmus] 299

hei Cavazzuti, 1996, Carabus 129

heideni Putzeys, 1873, Calathus 524

heidenii Dejean, 1828, Pterostichus 503

heikertingeri Mandl, 1955, Carabus 131



heilongjiangensis Deuve & Li, 1998, Carabus 122

heilongjiangensis Li, 1992, Pterostichus 515

heinertzi Deuve, Lassalle & Quéinnec, 1985, Calathus 525

heinigeri Morvan, 1992, Deltocolpodes 457

heinigeri Morvan, 1991, Leistus 80

heinigeri Morvan, 1997, Lepcha 459

heinigeri Morvan, 1995, Skoeda 466

heinrichi Mandl, 1966, Carabus 164

heinrichi Schatzmayr, 1936, Dyschirius 225

heinzi Jedlika, 1963, Amaroschema 521

heinzi Wrase, 1992, Apotomus 237

heinzi Korge, 1971, Bembidion 256

heinzi Jaeger, 1990, Bradycellus 401



heinzi Jedlika, 1965, Calathus 526

heinzi Breuning, 1964, Carabus 193

heinzi Zamotajlov & Wrase, 1997, Chinapenetretus 281

heinzi Casale & Giachino, 1991, Duvalius 303

heinzi Casale, 1988, Laemostenus 536

heinzi Farka, 1995, Leistus 80



heinzi Jedlika, 1965, Nebria 86

heinzi Dostal, 1986, Notiophilus 97

heinzi Wrase, 1991, Ophonus 391

heinzi F. Battoni, 1984, Pristosia 531

heinzi Jedlika, 1965, Pterostichus 496

heinzi Kirschenhofer, 1979, Pterostichus [HN] 497

heinzi Straneo, 1984, Pterostichus 494

heinzi Casale, 1982, Sphodropsis 543

heinzi Straneo, 1983, Tapinopterus 519

heinzi Pawowski, 1976, Trechus 343

heinzi Jedlika, 1965, Zabrus 571



heinziana Shilenkov, 1983, Nebria 85

heinzianus F. Battoni, 1986, Calathus 525

heinzianus Wrase, 1996, Ophonus 390

heinzianus Sciaky, 1996, Pterostichus 494

heinzianus Pawowski, 1979, Trechus 332

heinzorum Hieke, 1997, Amara 564



heldreichi Roeschke, 1891, Cicindela 104

heldreichi Schaum, 1856, Nebria 93

helena Kleinfeld, 1999, Carabus 125



helferi Jedlika, 1958, Pterostichus 507

helferi Schaum, 1863, Salpingophora [Cicindela] 114



helgae Schmid, 1975, Arctaphaenops 296

helleni J. Müller, 1922, Dyschirius 226

hellenica Cassola, 1973, Cylindera [Eugrapha] 111

hellenicus Heinz, 1982, Acinopus 368


hellenicus Mazzoldi & Toledo, 1992, Hydroporus 61

hellenicus Apfelbeck, 1904, Poecilus [Pterostichus] 483

hellenicus J. Müller, 1932, Tapinopterus 520

hellenicus Heyden, 1883, Zabrus 571



helleri Gredler, 1868, Amara 560

helleri Ganglbauer, 1891, Carabus 173



hellieseni Munster, 1922, Haliplus 30

hellmichi Jedlika, 1965, Amara 564

hellmichi Jedlika, 1965, Batoscelis [Pachytrachelus] 400

helluo Dejean, 1826, Carabus 167

hellwigii Panzer, 1803, Nebria [Carabus] 86

helmutschuetzei Kleinfeld, 1995, Carabus 124

helopioides Heer, 1838, Amara 563

helopioides Motschulsky, 1844, Amara [Bradytus] 567

helopioides Faldermann, 1836, Harpalus 376

helopioides Fabricius, 1792, Oodes [Carabus] 445

helopioides Reiche & Saulcy, 1855, Zabrus 571

helva Tschitschérine, 1898, Amara 552



helveticus Heer, 1837, Carabus 171

helveticus Heer, 1838, Licinus 442

helveticus Régimbart, 1901, Oreodytes [Hydroporus] 68

helveticus Jeannel, 1950, Trichaphaenops 344



helvitarsis Lutshnik, 1922, Parophonus [Harpalus] 392

helvolus Klug, 1834, Eretes [Eunectes] 53

hemicallisthenes Kabak & Kaláb, 1997, Carabus 147

hemicalosoma Semenov, 1903, Carabus 169

hemichlorus Fairmaire, 1888, Chlaenius 349



hemipterus Sirguey, 1931, Carabus 140

hemprichi Klug, 1832, Nebria 92

hemprichii Dejean, 1826, Carabus 157

hendrichi Fery & Hosseinie, 1998, Deronectes 58

hengduanicola Deuve, 1996, Carabus 175

hengduanicola Farka, 1999, Leistus 80

heniochicus Ljovuschkin, 1970, Trechus 332

henningi Fischer von Waldheim, 1817, Carabus 170



henoni Fairmaire, 1867, Amara 556

henoni Bedel, 1907, Calodromius [Dromius] 419

henoni Bedel, 1899, Dicheirotrichus [Trichocellus] 403

henoni Fairmaire, 1859, Glycia [Cymindis] 409

henrii P. Moravec & Lompe, nom. nov., Aphoenops 22, 294

henroti Coiffait, 1959, Geotrechus 310

henroti Jeannel, 1950, Henrotius [Hypogeobium] 471

henroti Jeannel, 1938, Oreonebria 96

henroti Cerruti, 1973, Speluncarius 516

henroti Jeannel, 1957, Typhloreicheia 223


henryi Andrewes, 1919, Chlaenius 349

henseleri B. Bodemeyer, 1922, Carabus 197

henvelikum Morvan, 1996, Xestagonum 468

henvelus Morvan, 1996, Platynus [Batenus] 463

hepaticus Faldermann, 1836, Laemostenus [Pristonychus] 538



hephaestus Zamotajlov, 1988, Carabus 132

heptapotamicum Tschitschérine, 1895, Bembidion 254

heptapotamicus Lutshnik, 1927, Pterostichus [Platysma] 502

heracleotes Jeannel, 1953, Speluncarius 516

herbacea Klug, 1832, Cicindela 105

herbertfranzi Toledano, 1998, Bembidion 252

herbeti Peytoureau, 1894, Cybister [Dytiscus] 51

herbsti Gmelin, 1790, Calosoma [Carabus] 120

herbsti Dejean, 1826, Carabus 151

herbstii Gistel, 1838, Collyris 116

hercegovinensis Knirsch, 1926, Duvalius [Neoduvalius] 307

hercegovinensis Krauss, 1906, Duvalius [Anophthalmus] 300

hercegovinensis Winkler, 1925, Winklerites [Scotodipnus] 240

hercegovinus Born, 1913, Carabus 165

herculeana Tschitschérine, 1894, Amara 560

herculis J. Frivaldszky, 1889, Duvalius [Trechus] 299



hermanni Born, 1902, Carabus 166

hermanni Fabricius, 1775, Hygrobia [Dytiscus] 35

hermensis Jeannel, 1920, Geotrechus 310

herminae Reitter, 1889, Carabus [Tribax] 200

herminius Jeanne, 1970, Platyderus 523

hermonensis Schweiger, 1970, Carabus 169

hermoniensis Hieke, 1997, Amara 566

hermonis Piochard de la Brûlerie, 1875, Leistus 83

heroicus Semenov, 1907, Carabus 155

heros Fabricius, 1801, Anisodactylus [Carabus] 361

heros Lapouge, 1925, Carabus 184

heros Kirschenhofer, 1995, Holosoma 444



heros Latreille, 1827, Scarites 234

heros Faldermann, 1836, Zabrus 569

herrmanni Mannerheim, 1827, Carabus 170

hervei Schuler, 1964, Amara 563

hervei Laneyrie & J. Ochs, 1948, Duvalius 302

hervei J. Ochs, 1953, Polistichus 574

herzegovinensis J. Müller, 1930, Molops 474

herzi Poppius, 1906, Pterostichus [Cryobius] 490



hesperia Casey, 1918, Amara 549

hesperica Motschulsky, 1849, Myriochila [Cicindela] 114

hespericus Rosenhauer, 1856, Harpalus 373

hespericus Motschulsky, 1849, Orthomus 477

hespericus Jeanne, 1985, Parophonus 393

hespericus Bucciarelli & Sopracordevole, 1958, Pterostichus [Platysma] 511



hespericus Dejean, 1831, Scarites 233

hesperidum Wollaston, 1861, Calomera [Cicindela] 99

hessei Kult, 1954, Dyschirius 225



heterocerum C. G. Thomson, 1868, Bembidion 259

heterodynamus Deuve, 1991, Carabus 156

heterolata Hieke, 1997, Amara 564

heteromorphus J. Müller, 1923, Anophthalmus [Trechus] 291

heteromorphus K. Daniel, 1896, Carabus [Orinocarabus] 180

heterostictus Tschitschérine, 1898, Microderes [Neopangus] 387

hetschkoi Reitter, 1911, Duvalius [Trechus] 303



hetzeri C. Müller, 1884, Abax 470

hexacoelum Chaudoir, 1850, Agonum [Anchomenus] 450

hexagrammus Chaudoir, 1876, Brachinus 215

heydeni Wehncke, 1872, Agabus 40



heydeni Schatzmayr, 1909, Amara 549

heydeni Ganglbauer, 1891, Bembidion 267



heydeni Paulino de Oliveira, 1876, Carabus 130

heydeni Kraatz, 1890, Cephalota [Cicindela] 102

heydeni Kraatz, 1883, Cymindis 416

heydeni Paulino de Oliveira, 1882, Cymindis 413

heydeni A. Fleischer, 1899, Dyschirius 225

heydeni Kraatz, 1890, Graphipterus 359

heydeni Wehncke, 1875, Haliplus 30

heydeni Hopffgarten, 1883, Molops [Tanythrix] [HN] 475

heydeni Kraatz, 1875, Molops 472

heydeni Krausse, 1915, Omophron 208

heydeni Semenov, 1890, Scarites 233

heydeni Ganglbauer, 1915, Zabrus 571

heydenianus Stark, 1889, Carabus [Cechenus] 137

heydenianus Jakobson, 1907, Pterostichus [Platysma] 512

heydenii Dejean, 1831, Nebria 92



heyerii Sturm, 1824, Pterostichus [Platysma] 507

heylensis Cabidoche, 1964, Trechus 327

heyrovskyi Jedlika, 1932, Bembidion 258

heyrovskyi Maan, 1936, Dyschirius 228

heyrovskyi Jedlika, 1928, Harpalus 377

heyrovskyi Jedlika, 1940, Platynus [Colpodes] 465

heyrovskyi Jedlika, 1932, Pristosia [Kanoldia] 531

heyrovskyi Jedlika, 1939, Tapinopterus 519

hibernicus Lindroth, 1956, Carabus 136

hickeri Breuning, 1927, Carabus 180

hickeri Knirsch, 1913, Duvalius [Anophthalmus] 299

hicksi Lindroth, 1968, Amara 556


hidakamontanus Ishikawa, 1966, Carabus [Procrustes] 164

hidakana Habu, 1972, Amara [Shunichius] 567

hidakanus Ishikawa & Takami, 1996, Carabus 176

hidalgoi Español & Comas, 1985, Aphoenops 294



hidamontanus Ishikawa, 1968, Carabus 159

hidaosa Ishikawa & Kubota, 1994, Carabus 177

hiekei Kryzhanovskij & Mikhailov, 1987, Amara [Curtonotus] 560

hiekei Müller‑Motzfeld, 1986, Bembidion 255

hiekei Kabak & Kryzhanovskij, 1990, Carabus 150

hiekei Jedlika, 1969, Cymindis 413

hiekei Shilenkov, 1982, Nebria 88

hiekei Naviaux, 1994, Neocollyris 117



hiekei Casale, 1988, Pseudotaphoxenus 542

hiekei Zaballos & Farinós, 1995, Typhlocharis 241

hiekeianus Deuve, 1991, Carabus 125

hiemalis Kryzhanovskij & Mikhailov, 1987, Dromius 420

hiemalis Semenov & Arnoldi, 1937, Notiophilus 97

hiemalis Apfelbeck, 1902, Trechus 333

hiemeieri Morvan, 1996, Platynus [Batenus] 463

hienfoungi J. Thomson, 1857, Carabus 156

hierichontica Reiche & Saulcy, 1855, Cymindis [Iscariotes] 415



hierichonticus Reiche & Saulcy, 1855, Licinus 441

hierroensis Machado, 1992, Platyderus 521

higonis Kasahara, 1994, Pterostichus 512

higonis Morita, 1997, Trichotichnus 396

hikosana Habu, 1955, Lebia 430

hikosana Habu, 1956, Nebria 94

hikosanum Habu & Uéno, 1955, Bembidion [Peryphus] 259

hikosanus Habu, 1953, Apatrobus [Penetretus] 280

hikosanus Kasahara, 1994, Pterostichus 512

hikosanus Habu, 1955, Synuchus [Calathus] 545

hilare Schmidt‑Göbel, 1846, Scalidion 430

hilariola Bates, 1874, Lophyra [Cicindela] 113

hilaris Tschitschérine, 1899, Acupalpus 399

hilaris Friederichs, 1903, Notiophilus 97

hilaris Fabricius, 1798, Pheropsophus [Carabus] 217

hilfi Born, 1907, Carabus 163

hilfi Reitter, 1903, Neotrechus [Anophthalmus] 315



himalaica Bates, 1878, Amara [Leiocnemis] 564

himalaicus Fedorenko, 1997, Dyschirius [Dyschiriodes] 229

himalayae Bates, 1891, Calathus 528

himalayae Habu, 1973, Xestagonum [Agonum] 468

himalayanum Andrewes, 1924, Bembidion 258

himalayanum Gestro, 1875, Calosoma 119

himalayanus Semenov, 1910, Craspedonotus 236


himalayanus Uéno, 1972, Trechus 325

himalaycus Jedlika, 1970, Platynus [Colpodes] 465

himalayensis Della Beffa, 1931, Hemiaulax [Idiomelas] [DA] 403

himalayensis G. Ochs, 1955, Orectochilus 28

himalayensis Vazirani, 1984, Orectochilus [HN!] 29

himalayica Jedlika, 1965, Agelaea 449

himalayica Uéno & Pawowski, 1983, Epaphiopsis 308

himalayica Andrewes, 1924, Lebidia 426

himalayica Bates, 1889, Nebria 94

himalayicus Bänninger, 1935, Haplogaster 232

himalayicus Jedlika, 1966, Harpalus 386

himalayicus Andrewes, 1931, Lionychus 431

himalayicus Andrewes, 1923, Peliocypas [Risophilus] 434

himalayicus Jedlika, 1935, Synuchus [Pristodactyla] 545



himalchuliensis Lassalle, 1985, Carabus 162

himalchuliensis Perrault, 1986, Leistus 80

himalchuliensis Lassalle, 1985, Trigonotoma 520

himalensis Deuve & Quéinnec, 1985, Trechus 332



himaleyica L. Redtenbacher, 1844, Calomera [Cicindela] 100

himaleyicus L. Redtenbacher, 1844, Lesticus [Omaseus] 471

hindukushi Casale, 1982, Pseudotaphoxenus [Taphoxenus] 541

hingstoni Baliani, 1934, Amara 552

hingstoni Andrewes, 1930, Bembidion 265

hingstoni Andrewes, 1930, Cymindis 413

hingstoni Andrewes, 1929, Dyschirius 229

hingstoni Jeannel, 1928, Trechus 332

hingstoni Andrewes, 1930, Trichotichnus 395

hiogoense Bates, 1873, Bembidion 246

hiogoensis Bates, 1873, Amara [Curtonotus] 560

hiogoensis Bates, 1873, Dyschirius 229

hiogoensis Kanô & Kamiya, 1931, Haliplus 31

hipponensis Géhin, 1885, Carabus 152

hipponensis Pic, 1894, Dyschirius 225

hiradonis Imura, Dejima & Mizusawa, 1993, Carabus 177


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