Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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hiraii Nakane, 1961, Carabus [Apotomopterus] 177

hirakei Uéno, 1979, Kurasawatrechus 313

hirakei Uéno, 1982, Yamautidius 345

hiranishi Morita, 1997, Trichotichnus 396

hiranoi Habu, 1972, Agonum 454

hiranoi Morita, 1996, Bembidion [Ocydromus] 265

hirasawai Morita, 1997, Trichotichnus 396

hirashimai Habu, 1954, Agonum 449

hirmocaelum Chaudoir, 1850, Bembidion 250

hirmocoelus Chaudoir, 1879, Platynus [Colpodes] 465

hirobane Habu & Baba, 1958, Pterostichus 502


hiroshimae Habu, 1975, Diacanthostylus [Platynus] 458

hirsutulus Bates, 1892, Apotomus 237

hirsutulus Dejean, 1829, Parophonus [Harpalus] 392

hirsutum Bates, 1873, Lachnoderma [Singilis] 438



hirticornis Krynicki, 1829, Stenolophus 406

hirtipennis Pic, 1901, Phloeozetus [Singilis] 438

hirtipes Panzer, 1796, Harpalus [Carabus] 377

hirtus Sturm, 1853, Anophthalmus 291

hirtus Dieck, 1869, Scotodipnus 240

hirus Morvan, 1976, Trechus 329

hisakai Imura, 1990, Carabus [Damaster] 122



hisamatsui Satô, 1961, Copelatus 48

hisamatsui Ishida & Shibata, 1961, Pterostichus 498



hisarensis Darge, 1990, Carabus 157

hispalensis Rambur, 1837, Brachinus 212

hispanica Csiki, 1929, Amara 555

hispanica Gory, 1833, Cephalota [Cicindela] 101



hispanica Motschulsky, 1844, Cicindela 108

hispanicum Dejean, 1831, Bembidion 261

hispanicus Gautier des Cottes, 1866, Calathus 525

hispanicus Rosenhauer, 1856, Deronectes [Hydroporus] 58



hispanicus Sharp, 1873, Ilybius 42

hispanicus Dejean, 1826, Lonchosternus [Oodes] 445



hispanicus Motschulsky, 1864, Microlestes [Blechrus] 423

hispanicus Dejean, 1828, Olisthopus 461



hispanicus Schauberger, 1926, Ophonus [Harpalus] 392

hispanicus Dejean, 1828, Orthomus [Feronia] 477

hispanicus Dejean, 1824, Pheropsophus [Brachinus] 217

hispanicus Sharp, 1882, Rhantus 46

hispanicus Matthews, 1899, Sphaerius 26

hispanus Puel, 1931, Anisodactylus 361

hispanus Fabricius, 1787, Carabus 140

hispanus Putzeys, 1866, Dyschirius 225

hispanus Jeanne, 1967, Laemostenus [Ceuthosphodrus] 535

hispanus Rambur, 1838, Parophonus [Harpalus] 393

hispanus Motschulsky, 1849, Scarites [Parallelomorphus] 232

hispanus Freude, 1988, Zabrus 573

hissarianus Semenov, 1900, Carabus 148

hissaricum Netolitzky, 1943, Bembidion 256

hissariensis Lapouge, 1925, Carabus 201

hissariensis Lutshnik, 1922, Ophonus [Harpalus] 388

histrio Andrewes, 1930, Amara 552

histrio Clark, 1864, Hydaticus 53

histrio Tschitschérine, 1903, Lophyra [Cicindela] 113

histrio Fabricius, 1801, Mastax [Brachinus] 217


hitleri Scheibel, 1937, Anophthalmus 291

hittita Sciaky, 1987, Ophonus 390

hiurai Kamiyoshi & Mizoguchi, 1960, Carabus 159

hiurai Satô, 1972, Hygrotus [Coelambus] 72

hiurai Uéno, 1985, Trechiama 322

hlisnikovskyi Mlyná, 1977, Molops 472



hnicolasi Chobaut, 1908, Carabus 195

hoberlandti Jedlika, 1951, Bembidion 271



hoberlandti Maan, 1952, Carabus [Procrustes] 158

hoberlandtianum Fassati, 1959, Bembidion 256

hobhousae Jeannel, 1930, Anillidius [Nannotrechus] 290

hochetlingeri Winkler, 1912, Anophthalmus [Trechus] 292

hochhuthi Chaudoir, 1846, Carabus 169

hochhuti Zaitzev, 1908, Agabus 37

hochstetteri Born, 1919, Carabus 140

hockingi Larson, 1975, Hydroporus 63

hodeberti Deuve, 1998, Trechus 332



hoeltzerae Lutshnik, 1916, Poecilus [Platysma] 482

hoelzeli Schweiger, 1950, Trechus 330

hoemorroidalis Ponza, 1805, Tachyura [Carabus] 278

hoenggandoensis Kwon & Lee, 1984, Carabus [Damaster] 144

hoepfneri Dejean, 1826, Nebria 90

hoffmanni Liebke, 1927, Brachinus 214



hoffmanni Motschulsky, 1845, Carabus 199

hoffmanni Schauberger, 1928, Harpalus 378

hoffmanni Netolitzky, 1917, Ocys [Bembidion] 272

hoffmanni Schauberger, 1927, Pterostichus 488

hoffmannseggii Panzer, 1803, Licinus [Carabus] 442

hoffmanseggii Dejean, 1828, Pterostichus [Feronia] 514

hohlbecki Jeannel, 1962, Trechus 337

hokkaidensis Habu & Baba, 1963, Harpalus 383

hokkaidensis Satô, 1972, Hygrotus 73

hokkaidoensis Lapouge, 1925, Carabus 136

hokurikuensis Ishikawa & Kubota, 1994, Carabus 177

holcartensis Genest, 1977, Geotrechus 310

holconotum Andrewes, 1935, Bembidion 20, 271



holdhausi Apfelbeck, 1904, Agonum [Platynus] 450

holdhausi Fassati, 1952, Bembidion 261

holdhausi Born, 1911, Carabus 168

holdhausi Gridelli, 1930, Microlestes 423

holdhausi J. Müller, 1917, Molops 474

holdhausi Ganglbauer, 1904, Orotrechus [Trechus] 317

hollandei Ledoux & Roux, 1993, Nebria 92

hollbergi Mannerheim, 1827, Carabus 192



holmbergi Putzeys, 1866, Amara [Curtonotus] 561

holmeni Vondel, 1991, Haliplus 32

holmeni Biström, 1983, Hyphydrus 74

holmeni Brancucci, 1983, Laccophilus 76

holofernes Semenov & Znojko, 1929, Odotoncarus [Carterus] 366

hololeucus Harald Lindberg, 1950, Apristus 431

holomera Chaudoir, 1871, Lebia 428



holosericeus Fabricius, 1787, Chlaenius [Carabus] 351

holosericeus Marsham, 1802, Hydroporus [Dytiscus] 64

holosericeus Chaudoir, 1850, Tilius [Lionychus] 433



holsaticus Reitter, 1908, Haliplus 31

holtzi Schatzmayr, 1909, Laemostenus 539

holtzi K. Daniel, 1903, Nebria 92

holzschuhi Wrase, 1995, Broscodera [Sinobrosculus] 235

holzschuhi Kirschenhofer, 1990, Calathus 528

holzschuhi Brancucci, 1986, Lacconectus 48

holzschuhi Morvan, 1997, Lepcha 459

holzschuhi Straneo, 1985, Lesticus 471

holzschuhi Wewalka, 1975, Platambus 44

holzschuhi F. Battoni, 1982, Pristosia 531

holzschuhi Morvan, 1995, Skoeda 466

holzschuhi Kirschenhofer, 1992, Trichotichnus 395

holzschuhi Casale, 1984, Vachinius 356

holzschuhi Morvan, 1996, Xestagonum 468

holzschuhianum Morvan, 1996, Xestagonum 468

holzun Shilenkov & Sokolov, 1987, Trechus 332



homagrica Duftschmid, 1812, Cymindis [Lebia] 412

homalonotus Tschitschérine, 1894, Pterostichus [Feronia] 490

homodynamus Deuve, 1994, Carabus 125

honamensis Kwon & Lee, 1984, Carabus 183

honanensis G. Hauser, 1921, Carabus [Coptolabrus] 144

hondorum Habu, 1974, Agonum [Platynus] 452

honestoides Reitter, 1900, Harpalus 377

honestus Duftschmid, 1812, Harpalus [Carabus] 377

hongdoensis Kwon & Lee, 1984, Carabus [Damaster] 144

hongyuanicus Deuve, 1991, Carabus 191

hongyuanicus Farka, 1999, Leistus 80

hongyuanus Zamotajlov & Sciaky, 1996, Parapenetretus 284

honnoratii Dejean, 1826, Carabus 195

honnoratii Dejean, 1828, Pterostichus [Feronia] 512

honshuense Habu, 1975, Agonum [Platynus] 453

honshuensis Habu & Baba, 1958, Pterostichus 487



hookeri Nodier, 1821, Carabus 153

hookeri Bates, 1875, Cymindis 413



hopei Parry, 1844, Heptodonta 115

hopei Gistel, 1837, Myriochila [Cicindela] 114

hopffgarteni Kraatz, 1877, Carabus [Procrustes] 187

hopffgarteni Kraatz, 1880, Carabus 172

hopffgarteni Schilsky, 1892, Graptodytes [Hydroporus] 59

hopffgarteni Heyden, 1881, Molops 473

hopffgartenianus Beuthin, 1885, Carabus [Orinocarabus] 179

hoplites Bates, 1883, Pterostichus [Allotriopus] 495

hoppei Germar, 1824, Carabus 180

hoppei Ganglbauer, 1891, Cychrus 203

hoppi Wagner, 1932, Chlaenius 352

hoppi Winkler, 1926, Nannotrechus 315

hoppi Jeannel, 1927, Trechus 332

horikawae Kanô, 1930, Paussus 210

horioi Nakane, 1961, Carabus 159

horioi Nakane, 1960, Nippononebria 95

horioi Nakane & Ishida, 1958, Pterostichus 507

horioni Fassati, 1957, Bembidion 245



horioni Korge, 1968, Carabus 131

hormuzakii Born, 1907, Carabus 182

horni Jedlika, 1932, Agonotrechus 289



horni Csiki, 1929, Amara 564

horni Andrewes, 1923, Anthracus [Acupalpus] 400

horni Jedlika, 1931, Beckeria 457

horni Jedlika, 1932, Catascopus 435



horni Jedlika, 1928, Dyschirius 228

horni Jedlika, 1928, Harpalus 384

horni Schauberger, 1938, Loxoncus [Anoplogenius] 404

horni G. Ochs, 1933, Orectochilus 29

horni Jedlika, 1932, Peliocypas [Risophilus] 434



horni Jedlika, 1934, Platynus [Colpodes] 465

hornschuchi Hoppe & Hornschuch, 1825, Carabus 198

horricoma Kryzhanovskij & Mikhailov, 1971, Petrimagnia 418



horsfieldii J. Thomson, 1857, Calochroa [Cicindela] 99

horsti Reitter, 1888, Lindrothius [Pristodactyla] 529



hortensis Fourès, 1954, Aphoenops 294

hortensis Linné, 1758, Carabus 179



hortensoides Sokolá, 1910, Carabus 181

hortulanus Darge, 1994, Carabus 130

horvathi Csiki, 1927, Carabus 144

horvathi Csiki, 1912, Duvalius [Anophthalmus] 300

horvathi Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 541

horvatovichi Deuve, 1992, Carabus 159

horvatovichi Kirschenhofer, 1991, Pterostichus 490

hoschecki Scheibel, 1933, Anophthalmus 293

hospes Sturm, 1818, Harpalus 377



hosti Depoli, 1929, Carabus 151

hostilis Sharp, 1884, Nebrioporus [Deronectes] 66

hottentotta Duftschmid, 1812, Harpalus [Carabus] 373

houhuanshanus N. Ito, 1985, Bradycellus 401



houlberti Bleuse, 1914, Carabus 195

houskai Jedlika, 1952, Cymbionotum [Graniger] 234



houskai Jedlika, 1952, Microdaccus 422

houskai Jedlika, 1955, Ophonus [Macrophonus] 389

houskai Jedlika, 1952, Poecilus [Pterostichus] 479

houzhenzi Farka, 1999, Leistus 80



hozari Maan, 1952, Carabus [Procrustes] 157

hozgargantae E.‑G. Burmeister, 1983, Ilybius [Agabus] 42

hozumii Ishida, 1961, Pterostichus 502

hroni Deuve, 1997, Carabus 153

hroni Dvoák, 1998, Laemostenus 534

hsifanicus Schauberger, 1932, Harpalus 383

hsuehshana Uéno, 1989, Epaphiopsis 309

huairenensis Deuve & Li, 1998, Carabus 122

huangi Deuve, 1992, Carabus 146

huangi Deuve, 1996, Cimmeritodes [Gotoblemus] 297

huangi Deuve, 1992, Cychrus 205

huanglongensis Deuve, 1990, Carabus 154

huangmaogeng Imura, 1999, Carabus 126

huashanus Sciaky, 1994, Pterostichus 500

huashixiaensis Mazzi, 1995, Carabus 190

huayangzhen Cavazzuti, 1999, Carabus 128

hubbardi Spangler, 1996, Sinodytes 57

hubeicus Deuve, 1991, Carabus 125

hubenthali Wagner, 1926, Pheropsophus 217

huchangensis Kwon & Lee, 1984, Carabus 183

hudsonica Hayward, 1908, Amara 561

hudsonica Casey, 1924, Blethisa 206

hudsonicus Leech, 1938, Agabus 37

hudsonicus Lapouge, 1925, Carabus 136

hudsonicus Motschulsky, 1866, Carabus 154

hudsonicus Fall, 1919, Hygrotus [Coelambus] 72

huebneri Fairmaire, 1878, Endynomena 437

huedepohli Mandl, 1983, Chlaenius 353

huegeli Roeschke, 1907, Carabus [Imaibius] 155

huemmleri Beuthin, 1899, Carabus [Euporocarabus] 179

huetei Ortuño, 1997, Hypotyphlus 239

huetheri Jeannel, 1934, Duvalius 303

huetheri Jeannel, 1934, Duvalius 303

huiphilus Deuve, 1994, Carabus 124

huisanensis Deuve & Li, 1999, Carabus 122

hulae Wewalka, 1984, Ilybius [Agabus] 42



hulunbeierensis Li, 1992, Cylindera [Eugrapha] 110

humboldtii Faldermann, 1836, Carabus 160



humerale Heer, 1838, Bembidion 263

humerale Sturm, 1825, Bembidion 244



humeralis Heyden, 1892, Acupalpus 398

humeralis J. Jacquet, 1938, Acupalpus 398

humeralis Audinet-Serville, 1821, Agabus (Dytiscus) [add, syn of nebulosus (Forter)] 40

humeralis Chaudoir, 1843, Aspidoglossa [Dyschirius] 219

humeralis Bonelli, 1813, Badister 441

humeralis Dodero, 1924, Boldoriella [Duvalius] 296

humeralis Ahrens, 1812, Brachinus 215

humeralis Beuthin, 1905, Cicindela 107

humeralis W. Horn, 1900, Cylindera [Cicindela] 111

humeralis Geoffroy, 1785, Cymindis [Buprestis] 413



humeralis Paykull, 1790, Cymindis [Carabus] 418

humeralis Marsham, 1802, Hydroporus [Dytiscus] 64

humeralis Uéno, 1957, Ishikawatrechus 312

humeralis Dejean, 1825, Lebia 429

humeralis Semenov, 1909, Pseudotaphoxenus [Taphoxenus] 541



humeralis Wollaston, 1854, Scarites 232

humeralis Uéno, 1999, Shenaphaenops 320



humeralis Motschulsky, 1844, Syntomus [Dromius] 433

humeralis Stephens, 1827, Syntomus [Dromius] 433

humeralis J. R. Sahlberg, 1900, Tachys 276

humeralis Dejean, 1831, Trechus 339

humeralis Uhagón, 1904, Zabrus 568

humerangula Hieke, 1995, Amara 560

humeratus Chaudoir, 1850, Dyschirius 228

humeratus Uéno, 1991, Masuzonoblemus 314

humeratus Mulsant & Godart, 1860, Stenolophus 406

humerosa Semenov, 1889, Agonops [Anchomenus] 449



humerosa Srnka, 1890, Cicindela 103

humerosa Quensel, 1806, Cymindis [Carabus] 413

humerosa Glasunov, 1902, Nebria 91

humerosus Knirsch, 1926, Duvalius [Neoduvalius] 307

humerosus Wollaston, 1858, Olisthopus 461

humerosus Tschitschérine, 1896, Poecilus [Feronia] 480

humerosus Uéno, 1960, Rakantrechus 319

humerula W. Horn, 1905, Cylindera [Cicindela] 111

humerulus Jeannel & J. Ochs, 1938, Duvalius 302

humi Roubal, 1936, Duvalius 307

humicola Löbl, 1995, Sphaerius [Microsporus] 26



humilis Baliani, 1934, Amara 554

humilis Casey, 1918, Amara 549

humilis Bernau, 1911, Carabus 184

humilis A. Morawitz, 1863, Clivina 220

humilis Klug, 1834, Hydroporus 61



humilis Sharp, 1882, Hydrovatus 70

humilis Casale, 1988, Pseudotaphoxenus 542

humiolcus Chaudoir, 1850, Dyschirius 228

hummeli Falkenström, 1936, Agabus [Gaurodytes] 36

hummeli Jedlika, 1935, Amara 554

hummeli Fischer von Waldheim, 1823, Carabus 170



hummeli Liebke, 1935, Cymindis 417

hummeli Falkenström, 1932, Haliplus 31

hummeli Falkenström, 1932, Hydroglyphus [Bidessus] 56

hummeli Jedlika, 1935, Pterostichus 515

hummelioides Deuve, 1989, Carabus 175

hummleri J. Müller, 1918, Bembidion 260

hummleri Jeannel, 1927, Trechus 332



hunanensis Born, 1910, Carabus [Coptolabrus] 141

hunanensis L. C. Pu, 1948, Hydroscapha 25

hunanicola Deuve & Yu, 1992, Carabus 156

hunanicus Deuve, 1995, Carabus 143

hunanus Morvan, 1997, Vachinius 356



hungarica Csiki, 1908, Cymindis 413

hungaricum J. Frivaldszky, 1877, Zuphium 574

hungaricus Fabricius, 1792, Carabus 182

hungaricus Csiki, 1903, Duvalius [Anophthalmus] 307

hungaricus Dejean, 1828, Pterostichus [Feronia] 495

hunnorum Lapouge, 1909, Carabus 172

hunteri Crotch, 1872, Sandracottus [Hydaticus] 49

hunyadensis Csiki, 1923, Carabus 172

huoditangicus Cavazzuti, 1999, Carabus 190

hupeensis G. Hauser, 1924, Carabus [Apotomopterus] 125

hureishanus Schweiger, 1970, Carabus 187

hurkai Gottwald, 1980, Carabus 200

hurkai Janák & P. Moravec, 1989, Duvalius 299

hurkai Divoký, Pulpán & Rébl, 1990, Harpalus 374

hurrita Pavesi & Sciaky, 1990, Trechus 332

hustachei Jeannel, 1917, Aphoenops 294

hustachei Antoine, 1923, Bembidion 264



hustachei Fagniez, 1927, Pterostichus 512

hustacheianus Puel, 1936, Pterostichus [Haptoderus] 490



hutchinsoni Andrewes, 1934, Bembidion 254

huxianensis Imura, 1993, Cychrus 203

huyuenii Lassalle & Remond, 1998, Carabus 191

hwangtienyuni Shilenkov & Kryzhanovskij, 1983, Archileistobrius 79

hweisiensis Jedlika, 1937, Pristosia [Eucalathus] 531

hweisinensis Schauberger, 1929, Harpalus 385


hyalina Semenov, 1889, Amara 562

hyalinus DeGeer, 1774, Laccophilus [Dytiscus] 76



hyalinus Marsham, 1802, Laccophilus [Dytiscus] [HN] 76

hyaloptera Semenov, 1891, Cymindis 415

hyantis Reitter, 1899, Nebria 86

hybneri Fabricius, 1787, Hydaticus [Dytiscus] 54

hybrida Linné, 1758, Cicindela 105



hybrida Heer, 1841, Cymindis 413

hybrida Rottenberg, 1874, Nebria 86

hybridum Apfelbeck, 1904, Bembidion 258

hybridus Reitter, 1908, Anchomenus [Chlaeniomimus] 456

hybridus Ganglbauer, 1887, Carabus [Procrustes] 187

hybridus Apfelbeck, 1902, Molops 473

hydaticoides Régimbart, 1877, Neptosternus [Laccophilus] 77

hydrophilus A. Gaudin, 1945, Aphoenops 294

hydrophilus Reitter, 1896, Carabus 154

hydrophilus Coiffait, 1962, Geotrechus 310



hydropica Putzeys, 1866, Clivina 220

hygrobius Uéno, 1955, Awatrechus 296

hygrotoides Fery, 2000, Hydroporus 61

hylonomus Deuve, 1992, Carabus 190

hylonomus A. Gaudin, 1935, Trechus 340

hyperboraeorum Munster, 1923, Bembidion 265

hyperborea Dejean, 1831, Amara 560

hyperborea Gyllenhal, 1827, Nebria 88

hyperboreus Motschulsky, 1844, Harpalus 381

hyperboreus Gyllenhal, 1826, Nebrioporus [Hyphydrus] 66

hyperboreus Kryzhanovskij, 1995, Notiophilus 97



hyperboreus Dejean, 1828, Patrobus 285

hyphydroides Perris, 1864, Herophydrus [Hydroporus] 71

hypochlorus Gozis, 1911, Rhantus 47

hypochora Tarrier, 1971, Carabus 166

hypocrita Roubal, 1937, Abax 469

hypocrita Apfelbeck, 1904, Agonum [Platynus] 450

hypocrita Dejean, 1831, Bembidion 257

hypocrita Chaudoir, 1869, Brachichila 435

hypocrita Semenov, 1926, Elaphrus 207

hypocrita Dejean, 1829, Harpalus 375

hypocrita Putzeys, 1866, Notiophilus 97

hypocrita Binaghi, 1936, Scotodipnus 240

hypogaeus Aubé, 1861, Anillus 238

hypogaeus Laneyrie, 1952, Duvalius 306



hypogaeus Fairmaire, 1856, Laemostenus [Pristonychus] 533

hypogeomysis Huang, 1993, Harpalus 385

hypogeus A. Gaudin, 1947, Geotrechus 310


hypsela Andrewes, 1923, Amara 552

hypseloides Baliani, 1934, Amara 553



hypselus Jedlika, 1958, Pterostichus 508

hypsibia Schauberger, 1934, Miscodera 237

hypsobius Apfelbeck, 1918, Carabus 195

hypsophila Antoine, 1953, Amara 567

hyrcana Tschitschérine, 1896, Cymindis 415


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