Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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hyrcanus Heinz, 1970, Calathus 527



hyrcanus Heinz, 1978, Carabus 137

hyrcanus Tschitschérine, 1904, Stomis 517

hysteron Netolitzky, 1943, Bembidion 270

hystrix Reitter, 1894, Ophonus 390

hystrix Westwood, 1850, Paussus 210

iberiae Schauberger, 1929, Ophonus [Harpalus] 389

iberica Mandl, 1935, Cicindela 106

iberica Jeanne, 1985, Cymindis 412

iberica Paulino de Oliveira, 1876, Nebria 93

ibericum Piochard de la Brûlerie, 1868, Bembidion 253

ibericus Jaeger, 1988, Acupalpus 398

ibericus Falkenström, 1939, Bidessus 55

ibericus Fischer von Waldheim, 1823, Carabus [Plectes] 174

ibericus Jeanne, 1976, Cychrus 204

ibericus Fedorenko, 1996, Dyschirius [Dyschiriodes] 229



ibericus Rosenhauer, 1856, Dytiscus 52

ibericus Pantel, 1888, Harpalus 375

ibericus Holdhaus, 1912, Microlestes 423



ibericus Tschitschérine, 1893, Poecilus [Feronia] 483

ibericus Régimbart, 1901, Scarodytes [Hydroporus] 68

iberiparcus Zaballos & García‑Munez, 1991, Parophonus 393

iberlovi O. Berlov, 1996, Pterostichus 506

iblis Peyerimhoff, 1910, Duvalius [Aphaenops] 308

iborensis Zaballos, 1990, Geocharis 239

icalari Nakajima, 1993, Carabus [Acoptolabrus] 122

ichangensis Bates, 1889, Carabus 125

ichihashii Uéno, 1959, Kurasawatrechus 313

iconiensis Ganglbauer, 1905, Zabrus 571

ictis Antoine, 1931, Bembidion 267

ida Mandl, 1954, Therates 115

idae Bates, 1873, Lebia 430

idae Maan, 1934, Tapinopterus 519

idaeus Schweiger, 1977, Calathus 524

idaeus Schweiger, 1968, Zabrus 571

idiotus Bates, 1889, Harpalus 370

idolon Semenov, 1898, Carabus 121

idriae Meschnigg, 1934, Bembidion 270


idriss Peyerimhoff, 1924, Trechus 332

ienishteai Nitzu, 1988, Pterostichus 503

ifniensis Zarco, 1941, Carabus [Cathoplius] 137

ifnii Fery, 1999, Hydroporus 63

ifranensis Fery, 1999, Hydroporus 63

ifranensis Antoine, 1941, Platyderus 522

igai Nakane & Uéno, 1953, Reicheiodes [Dyschirius] 231

igai Uéno, 1955, Trephionus 546

ignatovitschi Tschitschérine, 1894, Amara 560



ignavus Duftschmid, 1812, Harpalus [Carabus] 377

ignavus Stephens, 1828, Harpalus 381

ignavus Tschitschérine, 1897, Pterostichus [Feronia] 501

ignavus Csiki, 1907, Zabrus 572

ignicinctus Bates, 1883, Catascopus 435



ignicolor Reitter, 1893, Carabus 132

ignifer Haury, 1889, Carabus 139

igniferescens Deuve, 1992, Carabus 122

ignigenus G. Hauser, 1914, Carabus [Coptolabrus] 141

ignimitella Bates, 1888, Carabus 142

ignitum Kraatz, 1875, Xenion [Pterostichus] 521

ignitus Reitter, 1896, Carabus [Iniopachys] 156

ignoratus Stichel, 1923, Carterus [Odogenius] 364

ignotus Mulsant & Rey, 1861, Graptodytes [Hydroporus] 59

ignotus Mateu, 1971, Microlestes 423

igori Vereschagina & Kabak, 1996, Pseudotaphoxenus 541



ihai Habu, 1968, Harpalus 386

ii Uéno, 1970, Gotoblemus 311

ijimai Nilsson & Nakane, 1993, Hydroporus 61

ikazuchiense Habu, 1954, Agonum 453

ikedai Nilsson, 1997, Platambus [Agabus] 43

ikedai Kasahara, 1991, Platynus [Colpodes] 465

ikezakii Nakane, 1989, Jujiroa 459

ikiensis Nakane, 1968, Carabus [Apotomopterus] 177

ikutanii Tanaka, 1961, Pterostichus 494

ikutanii Uéno, 1961, Trechus 325

ilandjievi V. B. Guéorguiev, 1964, Pheggomisetes 318

ilcevi Knirsch, 1924, Pheggomisetes 318

ilgasdensis Jedlika, 1962, Harpalus 382

ilgazdaghensis Maan, 1932, Cychrus 202

ilgazdensis Jedlika, 1961, Poecilus [Pterostichus] 482

ilgazicus Breuning & Ruspoli, 1970, Carabus 131

ilgazicus Pawowski, 1976, Trechus 332

iliense Iablokoff‑Khnzorian, 1970, Bembidion 248

iliensis Kabak, 1994, Carabus 146

iliensis Glasunov, 1913, Mnuphorus 358



iljinskii Lutshnik, 1922, Poecilus [Platysma] 479

iljukhini Dolzhanski & Ljovuschkin, 1985, Duvalius [Jeannelius] 303



illairei H. Kolbe, 1895, Siagona 218

illautus Machado, 1992, Paradromius 424

illecebrosa Dokhtouroff, 1885, Cylindera [Cicindela] 110

illigeri Netolitzky, 1914, Bembidion 253

illigeri Dejean, 1826, Carabus 173

illigeri Crotch, 1871, Chlaenius 348

illigeri Luna de Carvalho, 1951, Platyrhopalus 211

illigeri Panzer, 1803, Pterostichus [Carabus] 511



illustris Putzeys, 1868, Broscus 236

illustris Mihók, 1914, Duvalius [Trechus] 300

illyricum Netolitzky, 1918, Bembidion 258



illyricus Kraatz, 1878, Carabus [Orinocarabus] 180

illyricus Kraatz, 1880, Carabus 129

illyricus Jeannel, 1921, Trechus 343



illyriensis Géhin, 1885, Carabus 129

ilmingi Schmid, 1965, Arctaphaenops 296

ilseanus Mandl, 1955, Carabus 148

ilvensis Holdhaus, 1924, Typhloreicheia [Reicheia] 223

imadatei Uéno & Shibanai, 1954, Trechiama [Trechus] 322

imafukui Nakane, 1962, Carabus [Apotomopterus] 177

imafukui Habu, 1961, Trichotichnus 396

imaicus Jeannel, 1923, Perileptus 289

imaicus Jeannel, 1923, Trechus 332

imaii Habu, 1972, Microlestes 423

imanensis Lafer, 1989, Carabus 134

imasakai Kasahara & Ohtani, 1988, Pterostichus 512

imathiae Casale, Giachino, Jali & Vailati, 1998, Reicheadella 222

imbricata Wollaston, 1871, Meladema [Scutopterus] 45

imbricatum Klug, 1832, Calosoma 121

imbricatus Klug, 1853, Synchortus [Noterus] 34

imereticum Belousov & Sokolov, 1996, Bembidion 259

imereticus Belousov, 1990, Trechus 332

imeretinus Kryzhanovskij, 1989, Pterostichus 494

imeretinus Dolzhanski & Ljovuschkin, 1985, Troglocimmerites [Birsteiniotrechus] 344



imitans Schauberger, 1926, Ophonus [Harpalus] 391

imitator K. Daniel, 1902, Bembidion 262



imitator Schweiger, 1969, Bradycellus 400

imitator Reitter, 1883, Carabus 181



imitator Reitter, 1895, Chlaenius 351

imitator Breit, 1914, Leistus 83

imitator J. Müller, 1917, Molops 472

imitator Zamotajlov, 1990, Penetretus 285

imitator Andrewes, 1930, Platynus [Colpodes] 465


imitator Belousov, 1998, Porocimmerites 318

imitator Sainte‑Claire Deville, 1902, Pterostichus [Platysma] 509



imitatoria V. E. Jakovlev, 1890, Cymindis 417

imitatrix Lutshnik, 1935, Amara 550

imitatrix Apfelbeck, 1904, Cymindis 413



imitatrix Tschitschérine, 1896, Stereocerus [Feronia] 517

imitatus Reitter, 1895, Chlaenius 352

imitatus Morvan, 1978, Pterostichus 515

immaculata Wanach, 1907, Cicindela 107

immaculata Dejean, 1829, Cymindis 418

immaculata Souverbie, 1855, Eurynebria [Nebria] 79

immaculatum J. R. Sahlberg, 1875, Bembidion 254

immaculatus Gschwendtner, 1927, Agabus [Gaurodytes] 40

immaculatus Schrank, 1781, Agabus [Dytiscus] 39

immaculatus Gerhardt, 1877, Haliplus 30



immaculatus Paulino de Oliveira, 1876, Lionychus 431

immaculicornis Dejean, 1826, Brachinus 214

immanis Bates, 1874, Hypaetha [Cicindela] 112

immarginatum J. Schmidt, 1997, Xestagonum 468

immarginatus Putzeys, 1866, Dyschirius 229

immarginatus Straneo, 1977, Pterostichus 510



imminutus Kraatz, 1879, Carabus [Procrustes] 187

immundus Reitter, 1900, Acupalpus 398

immune Stephens, 1828, Bembidion [Tachys] [HN] 265

immune Marsham, 1802, Bembidion [Carabus] (as immunis) 271



immunis Stephens, 1828, Ilybius [Colymbetes] 42

impar Lapouge, 1910, Carabus 168

impar Bonadona, 1955, Duvalius 306



impar Schauberger, 1927, Nebria 86

impar Andrewes, 1930, Syntomus [Metabletus] 432



impensus Reitter, 1896, Carabus [Iniopachys] 155

imperator Stark, 1890, Carabus 133

imperfecta Brown, 1930, Amara [Curtonotus] 560

imperfecta Chaudoir, 1852, Myriochila [Cicindela] 113

imperfectus Meier, 1899, Agabus 37

imperfectus Semenov, 1887, Carabus 178

imperialis Fischer von Waldheim, 1823, Carabus 164

imperialis Klug, 1834, Cephalota [Cicindela] 102



imperialis Zoubkoff, 1837, Cymindis 411

imperialis Germar, 1824, Demetrias [Dromius] 418

imperialis Reitter, 1886, Tapinopterus 519

imperialis Sciaky & Pavesi, 1994, Trechus 338

impexus Zamotajlov & Sciaky, 1999, Chinapenetretus 281



impicta Chaudoir, 1873, Cymindis 417

impiger Duftschmid, 1812, Harpalus [Carabus] 373

impiger Panzer, 1809, Synuchus [Carabus] 546

importunoides Jeanne, 1996, Dyschirius 229

importunus Schaum, 1857, Dyschirius 229

impressa Motschulsky, 1844, Amara 548

impressa Reitter, 1893, Cymindis 415



impressa Newman, 1833, Nebria [Helobia] 88

impressiceps Louwerens, 1953, Platynus [Colpodes] 465



impressicollis Visart, 1890, Calathus 525

impressicollis Fairmaire & Laboulbène, 1854, Pterostichus [Feronia] 512



impressicollis Zoubkoff, 1837, Scarites 234

impressifrons Fedorenko, 1993, Dyschirius 229



impressifrons Chaudoir, 1842, Elaphrus 207

impressifrons Joseph, 1870, Laemostenus [Sphodrus] 535

impressifrons A. Morawitz, 1862, Notiophilus 97



impressipennis Dejean, 1829, Harpalus 375

impressipennis Motschulsky, 1844, Harpalus 378

impressipennis Schilsky, 1891, Platynus [Anchomenus] 464

impressipennis Motschulsky, 1859, Tachys 275

impressopunctatus Schaller, 1783, Hygrotus [Dytiscus] 72

impressostriatum Motschulsky, 1865, Agonum [Agonothorax] 449

impressulus Schweiger, 1970, Carabus 187

impressum Panzer, 1796, Agonum [Carabus] 450

impressum Geoffroy, 1785, Asaphidion [Buprestis] 242

impressum Panzer, 1797, Bembidion [Elaphrus] 247

impressus Guignot, 1932, Agabus 38

impressus Zoubkoff, 1833, Agabus [Colymbetes] 37

impressus Klug, 1832, Carabus 188

impressus Zamotajlov & Sciaky, 1999, Chinapenetretus 281

impressus Putzeys, 1846, Dyschirius 227

impressus Fabricius, 1787, Haliplus [Dytiscus] 31

impressus J. Roth, 1851, Harpalus 370

impressus Fischer von Waldheim, 1829, Ophonus 392

impressus Fairmaire & Laboulbène, 1854, Pterostichus [Feronia] 503

impressus Fabricius, 1801, Scarites 233

impressus Dejean, 1825, Syntomus [Dromius] 432



improbus Schaufuss, 1872, Carabus 173

impubis H. Kolbe, 1886, Loxoncus [Anoplogenius] 404

impudicus J. Müller, 1931, Anophthalmus 293

impudicus Gautier des Cottes, 1866, Carabus 186

impunctata Schauberger, 1932, Acinopus 369

impunctata Reiche & Saulcy, 1855, Amara 568

impunctata Westhoff, 1881, Cicindela 103

impunctatus Puel, 1926, Acinopus 24, 368

impunctatus Kraatz, 1886, Carabus 165

impunctatus Andrewes, 1923, Platynus [Colpodes] 465



impunctatus Schilsky, 1888, Pterostichus 508

impunctatus Motschulsky, 1844, Syntomus [Dromius] 433

impunctatus Stephens, 1827, Syntomus [Dromius] 433

impuncticeps Reitter, 1900, Harpalus [Ophonus] 386

impunctipennis Dawson, 1854, Dyschirius 225

impunctoides Deuve, 1988, Trechus 325

impunctus Casale, 1979, Trechus 332



impustulatum Schilsky, 1888, Bembidion 267

imunada Uéno, 1993, Jujiroa 459

imurai Uéno, 1999, Trechus 332

imurai Morita, 1997, Trichotichnus 396

inaense Habu, 1956, Bembidion 270

inaequalis Abeille de Perrin, 1872, Aphoenops [Anophthalmus] 294

inaequalis Fabricius, 1777, Hygrotus [Dytiscus] 73

inaequalis Panzer, 1796, Laemostenus [Carabus] 539

inaequalis Reitter, 1900, Ophonus [Harpalus] 392

inaequalis Marsham, 1802, Pedius [Carabus] 478

inaequalis Wollaston, 1863, Syntomus [Metabletus] 432

inagakii Deuve, 1991, Carabus 125

inanis Casey, 1918, Amara [Curtonotus] 558

inapertus Motschulsky, 1850, Carabus 199



inapertus Faldermann, 1836, Pterostichus [Abax] 496

inauratum Mannerheim, 1825, Agonum 453

incatenatus Mannerheim, 1830, Carabus 194

incelatus Jeannel, 1923, Duvalius [Duvalites] 300

incerta Gautier des Cottes, 1868, Amara 565

incertans Mateu, 1952, Platyderus 522



incerticeps Chaudoir, 1850, Bembidion 251

incertum Dejean, 1831, Bembidion 258

incertus Brullé, 1834, Brachinus 216

incertus Haury, 1885, Carabus [Procrustes] 188

incertus Meyer‑Darcis, 1902, Carabus [Coptolabrus] 143

incertus Motschulsky, 1844, Carabus 169

incertus Dejean, 1829, Harpalus 379

incertus Régimbart, 1888, Hydaticus 53



incertus Aubé, 1838, Hydroporus 62

incertus Wollaston, 1864, Philorhizus [Dromius] 425



inchanicus Lapouge, 1918, Carabus 199

incidatus Ledoux & Roux, 1995, Archastes 79



incilis Dawson, 1849, Trechus 325

incisa Grimmer, 1841, Amara 550

incisa Dokhtouroff, 1888, Cylindera [Cicindela] 109

incisipenis Belousov & Kabak, 1999, Trechus 332

incisum Andrewes, 1921, Bembidion 243

incisus Dejean, 1829, Ophonus [Harpalus] 389



incisus Andrewes, 1927, Platynus [Anchomenus] 463

inclarus Darge, 1994, Carabus 154

inclavatus Israelson, 1983, Olisthopus 461

includens Walker, 1871, Rhantus 46

inclusa Chaudoir, 1865, Lophyra [Cicindela] 113

incognita Fassati, 1946, Amara 567

incognitum J. Müller, 1931, Bembidion 261

incognitus Bourgin, 1953, Carabus 195

incognitus Sharp, 1869, Hydroporus 61



incognitus Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 540

incognitus Scholz, 1927, Rhantus 46

incola Peyerimhoff, 1909, Trechus 332

incommodum Netolitzky, 1926, Bembidion 264

incommodus Schaum, 1858, Pterostichus 495

incompleta Fairmaire, 1885, Cylindera [Cicindela] 111

incompletus Fischer von Waldheim, 1828, Carabus 154

incompsus Kraatz, 1880, Carabus 173

incomptus Bates, 1873, Brachinus 215

inconcinnus Chaudoir, 1876, Harpalus 370

inconspectus Leprieur, 1876, Deronectes [Hydroporus] 58

inconspicua Blackburn, 1877, Endynomena [Saronychium] 437

inconspicuum Wollaston, 1864, Bembidion 248



inconspicuus Chaudoir, 1848, Carabus 130

inconspicuus Fall, 1917, Laccophilus 76

inconspicuus Chaudoir, 1855, Scarites 232

inconstans Régimbart, 1892, Hydroglyphus [Bidessus] 56

inconstans Mateu, 1981, Microdaccus 422

incrassata Baliani, 1934, Amara 558

incrassatus C. G. Thomson, 1871, Hydroporus 63

incrassatus Ahrens, 1814, Zabrus [Carabus] 571

incrematum LeConte, 1860, Bembidion 250

incultus Kraatz, 1875, Pterostichus 495

indagator Fabricius, 1787, Calosoma [Carabus] 121

indagi Vigna Taglianti, 1980, Sbordoniella 320



indentatus Newman, 1833, Leistus 82

inderiensis Motschulsky, 1850, Amara [Celia] 555

inderiensis Motschulsky, 1850, Chlaenius 351



inderiensis Motschulsky, 1848, Harpalus [Pseudoophonus] 383

inderiensis Zaitzev, 1915, Hydaticus 54

inderiensis Motschulsky, 1850, Poecilus [Lyrothorax] 479

indiae Louwerens, 1953, Platynus [Colpodes] 465

indica Putzeys, 1866, Amara [Leiocnemis] 564

indica Chaudoir, 1861, Galerita 359

indica Darlington, 1968, Gunvorita 573

indica Liebke, 1938, Lebia 429



indica Hope, 1831, Nirmala [Omaseus] 476

indica Thunberg, 1784, Ophionea [Attelabus] 444



indica Putzeys, 1846, Pseudoclivina [Clivina] 221

indicola Bates, 1878, Harpalus 377

indicum Chaudoir, 1850, Asaphidion [Tachypus] 242

indicum Hope, 1831, Calosoma 121



indicus Régimbart, 1899, Agabus 38

indicus Fairmaire, 1889, Carabus 162

indicus Sharp, 1882, Copelatus 47

indicus Aubé, 1838, Cybister 50

indicus Aubé 1838, Dineutus [Dineutes] 26

indicus Andrewes, 1923, Dromius 421

indicus Putzeys, 1866, Dyschirius 229

indicus Aubé, 1838, Gyrinus 27

indicus Bates, 1891, Harpalus 385



indicus Gschwendtner, 1936, Hyphydrus 74

indicus Gschwendtner, 1936, Laccophilus 76

indicus Régimbart, 1892, Leiodytes [Bidessus] 56

indicus Sharp, 1882, Nebrioporus [Deronectes] 66

indicus Bates, 1891, Orthotrichus 461

indicus Chaudoir, 1863, Paropisthius [Opisthius] 98

indicus Jeannel, 1923, Perileptus 289

indicus Putzeys, 1870, Trechus 332



indigena Knirsch, 1926, Anophthalmus [Haplotrechus] 291

indigestus Semenov, 1898, Carabus 190

indigonacea Miwa, 1935, Cicindela 108

indistincta Mannerheim, 1853, Amara [Celia] 564

indivisa Putzeys, 1871, Amara 557

indochinensis Bates, 1889, Dioryche [Platymetopus] 370

indus Kataev, 1997, Idiomelas 404

indus Tschitschérine, 1903, Leistus 80

indus Tschitschérine, 1900, Lesticus 471

indus Olivier, 1795, Scarites 232



indus Chaudoir, 1852, Sphodrus [Sphodrus] 543

induta Chaudoir, 1876, Siagona 218



ineditus Dejean, 1829, Harpalus 370

inefficiens Walker, 1859, Laccophilus [Hydroporus] 76



inepta LeConte, 1855, Amara 550

ineptus Sharp, 1882, Hydroporus 61

ineptus Coquerel, 1859, Pedius [Feronia] 478

inermis Curtis, 1831, Dyschirius 226

inermis Dawson 1854, Dyschirius [HN] 225

inermis Baehr, 1998, Gunvorita 573

inermis Uéno, 1980, Trechiama 322

inessae O. Berlov, 1997, Cychrus 19, 202


inexpectatior Deuve, 1991, Cychrus 204

inexpectatus Kryzhanovskij & Abdurachmanov, 1983, Abacetus 346

inexpectatus Sciaky & Benes, 1997, Morionidius 443

inexpectatus Uéno, 1980, Trechiama 322

inexpectus Schauberger, 1921, Pterostichus [Calopterus] 488

inexspectatum Kryzhanovskij, 1994, Xestagonum [Polychaetagonum] 468

inexspectatus Nilsson, 1990, Agabus 36

inexspectatus Basquin & Darge, 1988, Carabus 153

inexspectatus Kataev, 1989, Harpalus 377

inexspectatus Farka, 1999, Leistus 80

infans Andrewes, 1930, Bembidion 270

infans Netolitzky, 1934, Bembidion 268

infans Abeille de Perrin, 1909, Dixus [Aristus] [DA] 365

infans Lutshnik, 1930, Pseudotaphoxenus [Taphoxenus] 541

infans Bates, 1886, Tachyura [Tachys] 278

infantile Netolitzky, 1943, Bembidion 268

infantulus A. Morawitz, 1886, Carabus 123


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