Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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ischioxantha Bates, 1892, Loxocrepis [Colpodes] 460

iseranicus Deuve & Simard, 1977, Carabus 180



iserlei Kult, 1940, Dyschirius 229

isfahanensis Hieke, 1993, Amara 562

isfanensis Kabak, 1998, Poecilus 484

isfanensis Belousov & Kabak, 1998, Trechus 332

ishidai Ohkura & Shibata, 1963, Agonum [Colpodes] 452

ishidai Imura & Mizusawa, 1995, Carabus 177

ishidai Tanaka, 1961, Pterostichus 494

ishidai N. Ito, 1996, Trichotichnus 396

ishigakiensis Habu, 1975, Badister 441

ishigakiensis Habu, 1971, Loxocrepis [Agonum] 460

ishiharai Kasahara & Y. Itô, 1994, Apatrobus 280

ishiharai Uéno, 1994, Ishikawatrechus 312

ishiharai Ishida & Shibata, 1961, Pterostichus 491

ishiii Habu, 1961, Ophionea 444

ishikarinus Ishikawa, 1992, Carabus [Leptocarabus] 159

ishikawai Uéno, 1951, Ishikawatrechus [Nipponotrechus] 312

ishikawai Perrault, 1984, Leistus 83

ishikawai Nemoto, 1990, Morphodactyla [Calathus] 530

ishikawai Nemoto, 1988, Pterostichus 497

ishikawai Habu, 1950, Ryugadous 319

ishikawai Uéno, 1958, Trechiama 324

ishikawai Kasahara, 1992, Trichotichnus 396

ishikawaianus Breuning & Ruspoli, 1970, Carabus 201

ishikawaorum Uéno, 1982, Yamautidius 345

ishizuchianus Nakane, 1953, Carabus [Apotomopterus] 177



ishizuchiensis Habu, 1976, Apatrobus [Patrobus] 280

ishizuchiensis Uéno, 1962, Epaphiopsis 309

ishizuchiensis Kasahara, 1985, Pterostichus 512

ishizukai Deuve & Ohshima, 1989, Carabus 142

ishizukai Kasahara, 1995, Pterostichus 502

isignyensis P. Machard, 1977, Carabus 194

iskerensis Jedlika, 1964, Harpalus 379

islamiticus Sharp, 1882, Nebrioporus [Deronectes] 66



islamitus Reitter, 1899, Carabus 196

islandicum Sharp, 1900, Bembidion 271

islandicus Lindroth, 1968, Carabus 168



islei Barthe, 1921, Duvalius [Duvalites] 304

islei Barthe, 1922, Duvalius [Trechus] 304

ismenia Gory, 1833, Homodela [Cicindela] 112

ispartanum Netolitzky, 1930, Bembidion 255

ispiratus Cavazzuti, 1986, Carabus 185

ispirensis P. Machard, 1991, Carabus 201

ispiriacus Deuve, 1993, Carabus 147

ispulensis Kabak, 1994, Poecilus 480

ispulensis Belousov & Kabak, 1992, Trechus 332

israelita Ravizza, 1971, Bembidion 258

israelita Piochard de la Brûlerie, 1876, Ophonus 390



israelitus Reitter, 1899, Scarites 232

issykatae Chromý, 1993, Carabus 145

issykensis Lapouge, 1925, Carabus 178

issykkulensis Jedlika, 1969, Amara 564

istratii Hormuzaki, 1901, Carabus 139

istrianus Depoli, 1939, Abax 470

istrianus Ganglbauer, 1913, Anophthalmus [Trechus] 293

istrianus J. Müller, 1916, Molops 473

istriensis J. Müller, 1909, Anophthalmus [Trechus] 292



istriensis Schulz, 1909, Calomera [Cicindela] 100

istrus J. Müller, 1926, Graptodytes 59

istrus J. Müller, 1926, Typhlotrechus 344

isumiensis Kasahara & Saito, 1997, Pterostichus 502

italianus Csiki, 1927, Carabus 187

italica Klug, 1834, Cylindera [Cicindela] 109

italica Holdhaus, 1924, Reicheia 222

italicum De Monte, 1943, Bembidion 259

italicus Dejean, 1831, Brachinus [Aptinus] 212

italicus Dejean, 1826, Carabus 151

italicus Lapouge, 1925, Carabus 187

italicus Bonelli, 1810, Cychrus 204



italicus A. Fiori, 1913, Leistus 84

italicus Puel, 1925, Licinus 442



italicus Chaudoir, 1837, Pterostichus 513

italicus Dejean, 1828, Pterostichus [Feronia] 495

italicus K. Daniel & J. Daniel, 1898, Trechus 332

italus Schaum, 1860, Harpalus 377

iteratus Breuning, 1934, Carabus [Trachycarabus] 178

ithae Reitter, 1888, Trechus 332



itoensis Antoine, 1939, Carabus 152

itoi Ishikawa & Takami, 1996, Carabus 176

itoi Kasahara, 1986, Pterostichus 512

itoi Uéno, 1995, Trechiama 322

itoshimaensis Habu, 1954, Pogonus 287

itoshimanus Habu, 1954, Harpalus [Ophonus] 384

iturupensis Lafer, 1978, Amara 556

itzingeri Breuning, 1934, Carabus [Eucarabus] 190

iucundus Machado, 1992, Paradromius 424

iuliae Casale, Giachino, Vailati & Vigna Taglianti, 1996, Duvalius 303

iulianae Vigna Taglianti & Casale, 1973, Duvalius 303

ivanensis Belousov & Kabak, 1999, Trechus 339

ivicensis Knirsch, 1929, Neotrechus 315

iwasakii Morita, 1987, Apatrobus 280

iwasakii Uéno, 1979, Kurasawatrechus 313

iwasakii Kasahara, 1990, Pterostichus 494

iwasakii Uéno, 1988, Trechiama 322

iwatai Uéno, 1970, Daiconotrechus [Stygiotrechus] 297

iwatai Uéno, 1954, Trechiama [Trechus] 322

iwawakianus Nakane, 1953, Carabus [Apotomopterus] 178

iwawakianus Ishida, 1958, Pterostichus 507

ixion Andrewes, 1934, Bembidion 261

ixion Tschitschérine, 1902, Pterostichus [Platysma] 498

izeldoareum Morvan, 1996, Xestagonum 468

jablanicensis Maan, 1940, Molops 474

jablanicensis Apfelbeck, 1904, Neotrechus [Trechus] 315

jablanicensis Maan, 1935, Tapinopterus 520

jablanicensis Maan, 1939, Zabrus 570

jacobsoni Zaitzev, 1905, Agabus 37

jacobsoni Mikhailov, 1988, Bembidion 258

jacobsoni Semenov, 1908, Carabus 144

jacobsoni Sokolov & Shilenkov, 1987, Epaphiopsis 309

jacobsoni Zaitzev, 1927, Hydroporus 61

jacobsoni Semenov, 1922, Omophron 208

jacobsoni Pliginskiy, 1912, Pseudaphaenops [Trechus] 318

jacobsoni Poppius, 1906, Pterostichus [Cryobius] 491

jacobsoni Jedlika, 1952, Taphoxenus 544

jacquelini Boeildieu, 1859, Laemostenus [Pristonychus] 533

jacqueti Jeannel, 1941, Bembidion [Peryphus] 269



jacqueti Puel, 1936, Harpalus 383

jacqueti Pic, 1903, Pterostichus 512

jaculans Dejean, 1824, Brachinus [Aptinus] 212

jacupicensis Jedlika, 1935, Tapinopterus 520



jacuticus Znojko, 1932, Dyschirius 228

jacutorum Poppius, 1904, Poecilus [Pterostichus] 480



jacutus Mäklin, 1881, Carabus 172

jadodraconis Deuve, 1995, Trechus 333

jaechi Kirschenhofer, 1988, Apristus 431

jaechi Kirschenhofer, 1991, Chlaenius [Stenochlaenius] 355

jaechi Wewalka & Biström, 1987, Clypeodytes 55

jaechi Wewalka, 1989, Deronectes 58



jaechi Vondel, 1991, Haliplus 32

jaechi Fery & Nilsson, 1993, Ilybius [Agabus] 42

jaechi Sciaky, 1995, Myas [Trigonognatha] 476

jaechi Toledo, 1998, Nebrioporus 67

jaechi Wewalka, 1989, Neohydrocoptus [Hydrocoptus] 34

jaechi Mazzoldi, 1998, Orectochilus 29

jaechi Kirschenhofer, 1997, Pterostichus 491

jaechi Kirschenhofer, 1988, Syntomus 432

jaechi Kirschenhofer, 1991, Tapinopterus 519

jaegeri Kataev, 1996, Acupalpus 398

jaegeri Morita, 1997, Bradycellus 401

jaegeri J. Schmidt, 1996, Broscus 236



jaegeri Ménétriés, 1848, Carabus 199

jaegeri Fischer von Waldheim, 1832, Cicindela 105

jaegeri Balkenohl & J. Schmidt, 1997, Reicheiodes 230

jahni Schönmann, 1937, Trechus 330

jahorina Horvatovich, 1973, Nebria 87

jahorinae Jeannel, 1928, Duvalius 300

jailensis Breit, 1914, Laemostenus 534

jailensis Schauberger, 1926, Ophonus [Harpalus] 389

jailensis Winkler, 1911, Trechus 334

jajalaensis Hieke, 1995, Amara 553

jakeschi Jedlika, 1967, Chlaenius 355

jakeschi Jedlika, 1967, Pseudotaphoxenus [Taphoxenus] 542

jakeschi Jedlika, 1967, Singilis 438

jakobsonianus Lutshnik, 1928, Pterostichus [Platysma] 504

jakovlevi Tschitschérine, 1903, Notiophilus 97

jakovlevi Zaitzev, 1905, Oreodytes [Hydroporus] 67

jakowlewi Tschitschérine, 1898, Amara 552

jakowlewi Semenov, 1897, Carabus 137

jakowlewi Semenov, 1895, Cephalota [Cicindela] 102

jakowlewi Semenov, 1889, Cymindis 415

jakowlewi Semenov, 1895, Elaphrus 206


jakowlewi Semenov, 1898, Haliplus 30

jakowlewi Semenov, 1891, Mnuphorus [Hypercosmeton] 358

jakuchiensis Habu, 1977, Apatrobus [Patrobus] 280

jakupicaensis Roubal, 1920, Trechus 337

jakupicensis Maan, 1939, Molops 474

jakupicensis Maan, 1939, Zabrus 570



jakutskensis Csiki, 1930, Pterostichus 505

jalepensis Morvan, 1992, Deltocolpodes 457

jaloricus Andrewes, 1927, Licinus 441

jalzici Daffner, 1996, Anophthalmus 291

jalzici Pretner, 1979, Lovricia 241

jamae G. Etonti & M. Etonti, 1979, Orotrechus 317



jamagati Jedlika, 1956, Agonum [Colpodes] 453

jamata Motschulsky, 1866, Nebria 88

janae Farka & Plutenko, 1992, Leistus 82



janae Kult, 1943, Poecilus [Pterostichus] 484

janaki P. Moravec, 1993, Trechus 333

janakpurensis Kirschenhofer, 1992, Metacolpodes [Agonum] 460

janatai Bezina, 1996, Carabus 135

janatai Sciaky & Wrase, 1998, Doliodactyla 530

janatai Farka, 1999, Leistus 80

janatai Sciaky & Wrase, 1997, Pterostichus 500

janbritoi Morvan, 1994, Pterostichus 510

janczyki Casale, 1988, Laemostenus 537

jandiensis Machado, 1992, Philorhizus 425

janetscheki Mandl, 1970, Cychropsis 202

janetscheki Jedlika, 1970, Harpalus 385

janfarkaci Ledoux & Roux, 1997, Archastes 79

jani Toledano, 1998, Bembidion 252

jani Géhin, 1885, Carabus 158

jani Farka, 1995, Leistus 80

janinae Jeanne, 1984, Harpalus 370

janitor Jeannel & J. Ochs, 1938, Duvalius 305

jankowskianus Breuning, 1964, Carabus 191

jankowskiellus Deuve, 1991, Carabus 133

jankowskii Lafer, 1992, Agonum 450

jankowskii Oberthür, 1883, Carabus [Coptolabrus] 142

jankowskyi Tschitschérine, 1897, Pterostichus [Feronia] 507

jannonei Binaghi, 1955, Platyderus 523

jannuense Jedlika, 1968, Xestagonum [Agonum] 468

jannui Hieke, 1988, Amara 553

janoanus Jeannel, 1939, Trechiama [Trechus] 321

janoi Jeannel, 1937, Epaphiopsis [Trechus] 309

janoi Jedlika, 1952, Pterostichus 494

janschneideri Ledoux & Roux, 1999, Nebria 90



jansoni Kraatz, 1890, Carabus 160

jansoniana Wollaston, 1858, Perigona [Trechus] 449

janthinipennis Solsky, 1874, Poecilus 484

janthinus Ganglbauer, 1887, Carabus [Sphodristocarabus] 194

janthinus L. Redtenbacher, 1844, Chlaenius 355



janthinus Gautier des Cottes, 1872, Harpalus 377

janthinus Duftschmid, 1812, Laemostenus [Carabus] 536



janus Fairmaire, 1856, Harpalus 386

janus Reitter, 1900, Harpalus 377

janus Newman, 1833, Leistus 82

japana Motschulsky, 1858, Cicindela 106



japanensis Jedlika, 1962, Brachinus 216

japanensis Chaudoir, 1863, Cicindela 107

japanensis Lutshnik, 1922, Pterostichus [Platysma] 494

japanus Motschulsky, 1864, Stenolophus 406

japonica Motschulsky, 1860, Amara 548

japonica Jedlika, 1960, Calleida 408

japonica Guérin‑Méneville, 1847, Cicindela 106

japonica Thunberg, 1781, Cicindela 108

japonica Bates, 1883, Coptodera 436

japonica Bates, 1873, Drypta 358

japonica Bates, 1892, Eustra 208

japonica Bates, 1873, Galerita 359

japonica Bates, 1873, Lachnocrepis 445



japonica Chaudoir, 1871, Lebia 430

japonica Bates, 1883, Nebria 94

japonica Jedlika, 1946, Parena 434

japonica Bates, 1873, Perigona [Trechicus] 449

japonica Jedlika, 1935, Rupa 466

japonica Tamanuki, 1928, Tenomerga [Cupes] 25

japonicum A. Morawitz, 1862, Agonum [Dyscolus] 452

japonicum Motschulsky, 1860, Agonum [Tanystola] 452

japonicum Jedlika, 1961, Bembidion 257

japonicus Brinck, 1939, Acilius 48

japonicus Sharp, 1873, Agabus 36

japonicus Tschitschérine, 1898, Caelostomus [Stomonaxus] 471

japonicus Motschulsky, 1858, Carabus 177

japonicus Sharp, 1884, Copelatus 47

japonicus Sharp, 1873, Cybister 51

japonicus Andrewes, 1933, Dischissus 447

japonicus Uéno, 1954, Elaphrus 207



japonicus Sharp, 1873, Graphoderus [Hydaticus] 49

japonicus Sharp, 1873, Gyrinus 27

japonicus Sharp, 1873, Haliplus 31

japonicus A. Morawitz, 1862, Harpalus 385


japonicus Jedlika, 1958, Harpalus 384

japonicus Sharp, 1873, Hydroglyphus [Hydroporus] 56

japonicus Takizawa, 1933, Hydrovatus 70

japonicus Sharp, 1873, Hyphydrus 74

japonicus Bates, 1883, Morion 443

japonicus Sharp, 1873, Noterus 34

japonicus Chaudoir, 1861, Panagaeus 447

japonicus Jedlika, 1962, Panagaeus [Craspedophorus] 447

japonicus Bates, 1873, Perileptus 289

japonicus Putzeys, 1875, Pogonus 287

japonicus Motschulsky, 1860, Pterostichus [Omaseus] 493

japonicus Straneo, 1953, Stomis 517

jarkendis Jedlika, 1965, Pterostichus 511

jaroslawi A. Fleischer, 1898, Carabus 152

jarrigei Colas, 1971, Carabus 136

jarrigei Ledoux & Roux, 1991, Nebria 90

jarrigei Morvan, 1972, Trechus 333

jasilkowskii Born, 1905, Carabus 170

jason Semenov, 1898, Carabus 131



javanica Doesburgh, 1893, Calochroa [Cicindela] 99

javanus Wehncke, 1883, Canthydrus 34

javanus Csiki, 1938, Hydrovatus 71

javanus Klug, 1834, Miscelus 436

javanus Dejean, 1825, Pheropsophus [Brachinus] 217

javeti Jacquelin du Val, 1852, Bembidion 263

javetii Chaudoir, 1861, Cicindela 106

javurkovae Fassati, 1944, Bembidion 263

jaxartis Tschitschérine, 1903, Pterostichus [Platysma] 509

jeani Puel, 1925, Chlaenius 350

jeannei Ortuño & Arribas, 1992, Abax 470

jeannei J. Vives & E. Vives, 1989, Acinopus 368

jeannei Aubry, 1970, Agonum 449

jeannei Coiffait, 1960, Aphoenops 293

jeannei Lassalle & Pham, 1981, Carabus 141

jeannei Casale, 1988, Laemostenus 539

jeannei Zaballos, 1990, Platyderus 522

jeannei Sciaky, 1998, Trechus 333

jeannei Zaballos, 1989, Typhlocharis 241

jeanneli Abeille de Perrin, 1905, Aphoenops [Trechus] 293



jeanneli De Wailly, 1951, Bembidion [Peryphus] 253

jeanneli Razet, 1951, Brachinus 213

jeanneli Deuve, 1994, Carabus 155

jeanneli Kurnakov, 1960, Deltomerus 282

jeanneli Jedlika, 1934, Diacanthostylus [Colpodes] 458



jeanneli Binaghi, 1939, Duvalius 304

jeanneli Coiffait, 1973, Duvalius [Trechopsis] 303

jeanneli Winkler, 1933, Duvalius 300

jeanneli A. Gaudin, 1938, Geotrechus 310

jeanneli Vinson, 1956, Halocoryza 221

jeanneli Guignot, 1935, Hyphydrus 75

jeanneli Carret, 1907, Laemostenus 535

jeanneli Pater, 1939, Lebia 427

jeanneli Winkler, 1933, Neotrechus 315

jeanneli Sciaky, 1987, Ophonus 390

jeanneli Jedlika, 1952, Perileptus 289

jeanneli Mateu, 1950, Pterostichus [Haptoderus] 489

jeannelianum Casale & Vigna Taglianti, 1993, Bembidion [Ocydromus] 253

jeannelianus Paulian & Villiers, 1939, Laemostenus [Ceuthosphodrus] 535

jedlikai Louwerens, 1955, Platynus [Colpodes] 465

jedlickai Baliani, 1932, Amara 564

jedlickai Fassati, 1945, Bembidion 261



jedlickai Mandl, 1991, Chlaenius 354

jedlickai Schauberger, 1932, Chydaeus 362

jedlickai Zamotajlov, 1996, Diplous 283

jedlickai Kult, 1940, Dyschirius 224

jedlickai Schauberger, 1929, Harpalus 384

jedlickai Lafer, 1989, Lioholus 404



jedlickai Nemoto, 1990, Morphodactyla [Calathus] 530

jedlickai Ali, 1976, Perileptus 289

jedlickai Uéno, 1976, Perileptus 289

jedlickai Maan, 1935, Platyderus 522

jedlickai Hovorka & Sciaky, nom. nov., Pristosia 24, 531

jedlickai Maan, 1932, Tapinopterus [Pterostichus] 519

jedlickai Schauberger, 1932, Trichotichnus [Amaroschesis] 394



jedlickana Fassati, 1952, Amara 568

jedlickanum Toledano, 1999, Bembidion 267

jedlitshkaiana Lutshnik, 1935, Amara 548

jedompansis Deuve, 1990, Carabus 189



jeholensis Kamiya, 1935, Agabus 39

jejunus Dawson, 1854, Dyschirius 227

jelepa Andrewes, 1930, Lepcha 459

jelepus Andrewes, 1932, Pterostichus 515

jemelianovi Lafer, 1992, Agonum 454

jenisseense J. R. Sahlberg, 1880, Bembidion 247

jenisseicus Csiki, 1930, Pterostichus 491



jenisseiensis J. Müller, 1924, Dyschirius 226

jenissoni Dejean, 1831, Carabus 172

jenissoni Dejean, 1826, Siagona 218



jenseni Netolitzky, 1943, Bembidion 20, 265

jerdani Westwood, 1847, Paussus 210



jermolowi Stark, 1894, Carabus 200

jessoensis A. Morawitz, 1862, Pheropsophus 217



jessoensis Tschistchérine, 1897, Pterostichus [Feronia] 493

jessoensis Tschitschérine, 1899, Pterostichus 494

jezercensis Horvatovich, 1974, Nebria 85

jezerensis Apfelbeck, 1908, Trechus 333

jezerensis Apfelbeck, 1912, Trechus 333

jiajiensis Deuve, 1998, Carabus 133

jianensis Li & Chen, 1990, Ishidatrechus [DA] 312

jianyangensis Huang, Lei, Yan & Hu, 1996, Harpalus 380

jiddaica Ali, 1978, Cephalota [Cicindela] 102

jigongshanicus Kleinfeld, 1998, Carabus 124

jilanzhui Mazzoldi, 1998, Orectochilus 29

jilanzhui Wewalka & Brancucci, 1995, Platambus 43

jilinensis Deuve & Imura, 1990, Carabus 174

jilinensis Li, 1992, Carabus [Acoptolabrus] 122

jilinicus Deuve, 1992, Carabus 122

jindrai Farka, 1999, Leistus 80

jinfoshanus Cavazzuti & Ratti, 1999, Carabus 125

jingdongensis Deuve & Pan, 1999, Carabus 188

jinxiuensis Imura, 1995, Carabus 156

jirisanensis Ishikawa & Kim, 1983, Carabus [Eurycarabus] 149

jitkae Deuve & Macek, 1990, Carabus 158

jiudingensis Deuve, 1994, Carabus 175

jiulongensis Deuve, 1994, Carabus 190

jiulongensis Deuve, 1994, Cychrus 205

jiuyiensis Deuve, 1995, Carabus 127

jiuzhaiensis Deuve, 1992, Carabus 189

jiuzhaiensis Deuve, 1998, Trechus 333

jizuensis N. Ito, 1998, Trichotichnus 394

joannidisi Casale, Giachino & M. Etonti, 1990, Duvalius 308



joaquini Breuning & Toulgoët, 1974, Carabus 167

jockischii Duftschmid, 1812, Nebria [Carabus] 88

jockischii Sturm, 1815, Nebria [HN!] 90



joenius Patti, 1844, Brachinus 213

joergeri Born, 1927, Carabus 173

joerini Stierlin, 1890, Carabus 139

joffrei Sainte‑Claire Deville, 1925, Anillus 238

joffrei J. Ochs, 1926, Duvalius [Duvalius] 305

joffrei J. Ochs, 1938, Hydraphaenops [Aphoenops] 312

jogaesanensis Lafer, Paik & Park, 1996, Pterostichus 502



johannis Reitter, 1898, Carabus 178

johannis Peyron, 1856, Tapinopterus [Feronia] 520

johnbeckeri Bänninger, 1927, Archaeocindis [Cicindis] 98


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