Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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johnnyi Curti, 1981, Duvalius 306


johnsahlbergi Csiki, 1929, Amara 561

jollyi Deuve & Mourzine, 1997, Carabus 182

jollyi Deuve & Mourzine, 1997, Cychrus 206

jonescoi Jeannel, 1919, Duvalius [Trechus] 299

jonicus L. Miller, 1862, Hydroporus 62

joniensis Breuning, 1932, Carabus 187

jordai Reitter, 1914, Henrotius [Hypogeobium] 471

jordanense Piochard de la Brûlerie, 1876, Bembidion 254

jordani Beuthin, 1892, Carabus 129

jordanicus Schweiger, 1970, Carabus 188

jordanicus Fedorenko, 1996, Dyschirius [Dyschiriodes] 226

jordanis Reiche & Saulcy, 1855, Cybister 51

jordanus Jedlika, 1964, Harpalus 377

jorgensis Oromí & Borges, 1991, Trechus 333

josephi Bonavita, 2001, Bembidion 246

josephi Csiki, 1930, Pterostichus 491

jousselinii Guérin‑Méneville, 1836, Paussus 210

joyceae Nilsson, 2001, Hydrovatus 70

jozei Daffner, 1996, Anophthalmus 291

juccii Pomini, 1940, Orotrechus 317

jucunda Boheman, 1848, Drypta 358

jucunda Reitter, 1885, Tachyura [Tachys] 21, 280

jucunda Holdhaus, 1924, Typhloreicheia [Reicheia] 223



jucundus Csiki, 1906, Carabus 173

jucundus Reiche, 1850, Hydaticus 53

jucundus Guignot, 1954, Platambus [Agabus] 44

jucundus Perris, 1869, Rhithrodytes [Hydroporus] 68

jucundus Csiki, 1912, Trechus 334

judaeus Puel, 1936, Chlaenius 355

judaeus Piochard de la Brûlerie, 1876, Ophonus 390

judaicum J. R. Sahlberg, 1908, Bembidion 264

judaicus Géhin, 1885, Carabus 157

judicariae Jeannel, 1926, Duvalius 306

juengerianus Kleinfeld, 1995, Carabus 135

juenthneri Reitter, 1899, Carabus [Plectes] 132



jugicola Barthe, 1909, Carabus 168

jugicola Fairmaire & Laboulbène, 1854, Platyderus 522

jugicola Lutshnik, 1933, Pterostichus [Platysma] 496



jugicola Lorenz, 1998, Trechus 333

jugivagus Tschitschérine, 1898, Pterostichus [Feronia] 510



jugivagus Jeannel, 1962, Trechus 333

jugivagus Lutshnik, 1930, Trechus 333

jugosa Ledoux & Roux, 1998, Nebria 92

jugularis Babington, 1832, Hydroporus 62

juldusanus Breuning, 1933, Carabus 178


juldusanus Reitter, 1913, Leistus 82

juldusiensis Hieke, 1988, Amara 560



juliae Ballion, 1871, Cylindera [Cicindela] 109

julianae Zaballos, 1989, Geocharis 239

julianum De Monte, 1943, Bembidion 269

julianus Jeannel, 1927, Trechus 335

julieni Lesne, 1896, Clivina 219

juliettae Fourès, 1948, Aphoenops 294

julonshanensis Zamotajlov, 1993, Diplous 283

jumlana Hieke, 1981, Amara 553

jumlanus Straneo, 1982, Pterostichus 510

junceus Andrewes, 1923, Chlaenius 354

junceus Scopoli, 1763, Platynus [Carabus] 464

juncoi Jeanne, 1996, Platyderus 522



juncta Kraatz, 1887, Cicindela 104

jungens Tschitschérine, 1893, Pterostichus [Feronia] 495



jurassicus Born, 1911, Carabus 184

jurassicus David & Marchal, 1968, Carabus 166

jurassicus Sermet, 1960, Trichaphaenops 344



jureceki Jedlika, 1940, Agonum 454

jureceki Jedlika, 1957, Amara 560

jureceki Maan, 1939, Calathus 527

jureceki Jedlika, 1935, Chlaenius 350

jureceki Jedlika, 1935, Dromius [Lebidromius] 421

jureceki Dodero, 1917, Duvalius [Trechus] 303



jureceki Jedlika, 1928, Dyschirius 228

jureceki Jedlika, 1928, Harpalus [Pseudophonus] 385

jureceki Jedlika, 1931, Pristosia [Kanoldia] 531

jureceki Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 541

jureceki Jedlika, 1936, Pterostichus [Neohaptoderus] 501

jureceki Reitter, 1908, Tachyura [Tachys] 278

jureceki Maan, 1947, Typhlotrechus 344

jurecekianum Jedlika, 1952, Agonum 453

jurecekianus Maan, 1940, Pterostichus 506



jureckovae Maan, 1943, Zabrus 570

jurgae Obydov, 1995, Carabus 170



jurgitae Deuve, 1993, Carabus 135

jurine Panzer, 1803, Pterostichus [Carabus] 503



jurinei Born, 1901, Carabus 184

jurjurae Peyerimhoff, 1909, Duvalius [Trechus] 303

jurjurae Peyerimhoff, 1908, Zabrus 569

jurjurensis Régimbart, 1895, Hydroporus 62

jusmeti Español, 1971, Speleotyphlus [Catalanotyphlus] 240

justinae Meschnigg, 1947, Bembidion 245



justinae Reitter, 1888, Carabus [Tribax] 200

justusii W. Redtenbacher, 1842, Pterostichus 507

juvencus Ballion, 1871, Pseudotaphoxenus [Sphodrus] 541



juvenilis K. Daniel & J. Daniel, 1898, Platynus 463

juvenilis Schaum, 1858, Pterostichus 488

juxtata Acciavatti & Pearson, 1989, Cosmodela [Cicindela] 108



juzai Deuve, 1992, Carabus 160

juzetensis Coiffait, 1958, Geotrechus 310



kaadeni Schaufuss, 1879, Carabus 138

kabacaensis P. Machard, 1992, Carabus 201

kabaki Vereschagina, 1988, Pseudotaphoxenus [Taphoxenus] 542

kabakianus Kataev, 1988, Harpalus 377

kabakovi Hieke, 1976, Amara 566

kabakovi Mikhailov, 1984, Bembidion 20, 256

kabakovi Lafer, 1989, Carabus 134

kabakovi Fedorenko, 1996, Dyschirius [Dyschiriodes] 230

kabakovi Kataev, 1987, Harpalus 377

kabakovi Shilenkov, 1982, Nebria 90

kabakovi Casale & Vereschagina, 1986, Pseudotaphoxenus 541

kabakovi Dostal, 1997, Scarites [Scallophorites] 233

kabakovi Pawowski, 1978, Trechus 333

kabakovianus Gottwald, 1991, Carabus 155

kabardensis Kurnakov, 1972, Carabus 131

kabardensis Zamotajlov, 1996, Deltomerus 282



kaboulensis Coiffait, 1962, Laemostenus [Pristonychus] 538

kabulensis Jedlika, 1956, Lebia 427

kabulensis Jedlika, 1956, Poecilus [Pterostichus] 479

kabyliana Bruneau de Miré, 1955, Reicheia 222

kabylianus Reiche, 1862, Scybalicus 363

kabylianus Puel, 1935, Tachys [DA] 275

kabylicus Bruneau de Miré, 1958, Laemostenus [Antisphodrus] 534

kabylicus Casale, 1983, Trechus 333

kacheticus Lutshnik, 1928, Pterostichus [Platysma] 501



kachovskyi Lutshnik, 1935, Amara 558

kackardagi Pawowski, 1978, Trechus 333

kadjaranze Jedlika, 1947, Pterostichus [Haptoderus] 491

kadleci Novotný & Voíšek, 1988, Carabus 200

kadleci Kataev & Wrase, 1995, Harpalus 377

kadleci Dvoák, 1995, Pterostichus 504

kadmos Wewalka, 1989, Bidessus 55

kadoudali Morvan, 1982, Carabus 162

kadoudali Morvan, 1982, Pterostichus 510

kadyrbekovi Kabak, 1991, Amara [Curtonotus] 560

kadyrbekovi Kabak, 1994, Carabus 146

kadyrbekovi Kataev, 1988, Harpalus 377

kaempferi C. G. Thomson, 1875, Carabus 177


kaesongensis Imura, 1993, Carabus 174

kafekus Morvan, 1976, Trechus 329

kafkai Deuve, 1994, Carabus 145

kaganensis Heinz & Ledoux, 1987, Ledouxius [Penetretus] 284

kaghanensis Heinertz, 1978, Carabus 155



kagoshimaensis Iga, 1955, Cicindela 106

kagyzmanicus Kataev, 1984, Harpalus 378

kahleni Daffner, 1998, Anophthalmus 291

kaii Morvan, 1982, Aparupa 456

kaikanica Kabak, 1999, Cymindis 416

kaikanicus Belousov & Kabak, 1994, Trechus 333

kailasensis Hieke, 1997, Amara 557

kailiensis Huang, 1992, Harpalus 385

kaimakcalanensis Horvatovich, 1974, Nebria 86

kaiyaensis Kanô & Chô, 1933, Calomera [Cicindela] 99

kajimotoi Uéno, 1975, Ryugadous 319

kajimurai Habu & Tanaka, 1957, Pterostichus 494

kajmakcalensis Jedlika, 1939, Pterostichus 508

kalabellus Deuve, 1993, Carabus 178

kalabi Deuve, 1993, Broscosoma 235

kalabi Deuve, 1990, Carabus 146

kalabi Deuve, 1991, Cychrus 204

kalabi Farka, 1995, Leistus 80

kalabi Sciaky, 1994, Pterostichus 500

kalabi Deuve & Quéinnec, 1993, Trechus 333

kalabianus Deuve, 1993, Trechus 333

kalalae Maan, 1935, Dyschirius 225

kalalovae Roubal, 1932, Brachinus 215



kalardachti Jedlika, 1968, Laemostenus [Pterostichus] 536

kalavrytana Reitter, 1884, Cymindis 413

kalavrytanus Maan, 1939, Tapinopterus 519

kalchhigenn Morvan, 1999, Henvelik 458



kalchledanum Morvan, 1996, Xestagonum 467

kalchmoanum Morvan, 1996, Xestagonum 468

kaleea Bates, 1866, Cylindera [Cicindela] 111

kalganus Jedlika, 1953, Pseudotaphoxenus [Taphoxenus] 541

kalhys Motschulsky, 1866, Pterostichus [Logarus] 513

kaliki Kult, 1949, Dyschirius 230

kalisi J. Müller, 1931, Orotrechus 316

kaljuzhnyji Zamotajlov, 1988, Carabus 133



kalmyk Gryuntal, 1984, Dyschirius 226

kalmyka Lutshnik, 1927, Amara 552

kalofericus Mlyná, 1977, Molops 472



kalofirensis Apfelbeck, 1904, Carabus 129

kalofirensis Maan, 1933, Tapinopterus 519



kamareti Morvan, 1995, Platynus [Colpodes] 465

kamaretianum Morvan, 1995, Xestagonum [Colpodes] 468



kamberskyi Reitter, 1889, Carabus [Tribax] 200

kamberskyi Reitter, 1889, Poecilus 482

kamberskyi Reitter, 1888, Thermoscelis 529

kamensis Semenov, 1903, Carabus 175

kamenskii Kryzhanovskij, 1964, Amara 558

kamicola Deuve, 1995, Carabus 191

kamikochii Jedlika, 1965, Bembidion 271

kamikochii Jedlika, 1955, Pterostichus 502

kamiyai Nakane, 1963, Haliplus 31

kammuriensis Tamu & Tsukamoto, 1955, Copelatus 47

kamnikensis Drovenik, 1987, Aphaenopidius 293

kampalaensis Hieke, 1997, Amara 564



kamtchaticus Motschulsky, 1844, Carabus 135

kamtschatensis Breuning, 1942, Carabus 172

kamtschaticus Motschulsky, 1860, Pterostichus 499

kamtschatkensis Putzeys, 1847, Trechus 326

kana Imura, 1989, Carabus [Damaster] 122

kanabei Csiki, 1940, Duvalius 299

kanalense Fauvel, 1882, Omophron 208

kanalensis Perroud & Montrousier, 1864, Tachyura [Bembidium] 278

kandaharensis Jedlika, 1956, Abacetus 347

kandaharensis Jedlika, 1956, Harpalus 378

kandshutica Glasunov, 1902, Nebria 91

kanekiyo Uéno, 1958, Trechiama 323



kanganensis Deuve, 1982, Carabus 155

kangdingensis Hieke, 1997, Amara 560



kangdingensis Deuve, 1989, Carabus 190

kangdingensis Deuve, 1991, Cychrus 203

kangdingensis Farka, 1995, Leistus 80

kangdingi Korell, Kleinfeld & Görgner, 1992, Carabus 147

kangwonensis Kwon & Lee, 1984, Carabus [Damaster] 142

kaninensis Poppius, 1906, Pterostichus [Cryobius] 491

kanoi Kamiya, 1938, Oreodytes [Deronectes] 67

kansaiensis Nakane, 1961, Carabus 152

kansou Feng, 1936, Cybister 50

kansouis Feng, 1936, Platambus 43

kanssuensis Schilder, 1953, Cicindela 107



kanssuensis Tschitschérine, 1889, Pterostichus [Bothriopterus] 487

kansuensis Mandl, 1986, Callistomimus 348

kansuensis Schauberger, 1932, Gnathaphanus 362

kansuensis Jedlika, 1965, Pseudotaphoxenus [Pterostichus] 541



kantaikensis Géhin, 1885, Carabus 163

kantegiricus Belousov & Kabak, 1994, Trechus 333



kantinensis Zamotajlov, 1992, Archipatrobus 281

kantoonus Habu, 1961, Trichotichnus 396

kantous Tanaka, 1958, Pterostichus 502

kaonastennus Morvan, 1995, Platynus [Colpodes] 465

kaorutanakai Habu, 1977, Tachys 275

kapparicola Ganglbauer, 1889, Tapinopterus 519

kaqiensis Deuve, 1998, Trechus 338

kara Andrewes, 1921, Bembidion 270

karaalmensis Obydov, 1999, Calosoma [Callisthenes] 119

karaalmicus Gottwald, 1994, Carabus 182

karadaghensis S. Battoni & Blumenthal, 1973, Carabus 157

karadagi Zamotajlov, 1996, Deltomerus 282

karadenizus Pawowski, 1976, Trechus 333

karafutensis Csiki, 1927, Carabus 136

karagaicum Lapouge, 1924, Calosoma [Callisthenes] 119



karagaicus Lapouge, 1921, Carabus 192

karageorgii Lutshnik, 1915, Pterostichus [Platysma] 495

karakorum Jedlika, 1958, Harpalus 378

karakuldzhensis Kabak, 1994, Poecilus 484

karalangana Hieke, 1996, Amara 551



karamani Mandl, 1989, Cychrus 205

karamani J. Müller, 1922, Dyschirius 224

karamani Apfelbeck, 1902, Harpalus 378

karasawai Ishikawa, 1966, Carabus 149

karasawai Tanaka, 1958, Pterostichus 493



karasawai Tanaka, 1960, Sericoda 466

karasawai Tanaka, 1962, Stenolophus 405



karascharensis Eidam, 1931, Carabus [Zoocarabus] 192

karasibensis Belousov & Kabak, 1994, Trechus 333

karasudominus Cavazzuti, 1991, Carabus 194

karatavensis Kryzhanovskij, 1953, Carabus 147

karateghinicus Mikhailov, 1972, Pterostichus 486

karaterekensis Kaláb, 1996, Carabus 147

karatogani Belousov & Kabak, 1994, Trechus 327

karavaensis Horvatovich, 1974, Nebria 84

karavaievi Lutshnik, 1922, Cicindela 108

karavshinensis Kabak, 1998, Carabus 182

karelhurkai Farka, 1990, Duvalius 308

karelini Zoubkoff, 1837, Broscus 236

karelini Fischer von Waldheim, 1830, Calosoma [Callisthenes] 119

karelini Fischer von Waldheim, 1830, Carabus 199



karelini Fischer von Waldheim, 1832, Cicindela 107

karelini Motschulsky, 1844, Glycia [Glycia] 409

karelinii Chaudoir, 1842, Chlaenius [Dinodes] 352

karelinii Mannerheim, 1844, Chlaenius [Epomis] 353

karelinii Chaudoir, 1842, Poecilus 484

karenia Bates, 1892, Lebia 429

karennius Bates, 1892, Anisodactylus [Harpalus] 361

karianus Straneo, 1935, Tapinopterus [Pterostichus] 520

karibanus Ishikawa, 1992, Carabus [Leptocarabus] 159

karkarensis Kabak & Ovchinnikov, 1994, Carabus 150

karkarensis Belousov & Kabak, 1992, Trechus 331

karli J. Schmidt, 1998, Xestagonum 468

karluk Kabak, 1997, Poecilus 484

karlukovensis Genest, 1978, Pheggomisetes 318

karokhense Marggi, nom. nov., Bembidion 19, 256

karounense Jedlika, 1959, Bembidion 248

karpinskii Kryzhanovskij & Matveev, 1993, Carabus 171

kartalinicum Lutshnik, 1938, Bembidion 245

karumicus Bolívar y Pieltain, 1914, Broscodes [Broscus] 235

karvanekum Morvan, 1996, Xestagonum 468

karzhantavensis Kabak, 1991, Amara [Curtonotus] 560

kasaharai Habu, 1978, Cillenus 272



kasaharai Lindroth, 1980, Cillenus [Bembidion] 272

kasaharai Uéno, 1989, Epaphiopsis 309

kasaharai Habu, 1978, Planetes 359

kasaharai Habu, 1983, Trichotichnus 396

kasakh Kryzhanovskij & Emetz, 1973, Cymindis 416

kasakhstanicus Gryuntal, 1984, Dyschirius 226

kasakorum Semenov, 1897, Carabus 169

kasantsevi Kabak, 1993, Carabus 146

kasantsevi Zamotajlov & Sciaky, 1999, Robustopenetretus [Chinapenetretus] 286

kasbekianus Kraatz, 1877, Carabus 200

kasbekianus Kolenati, 1845, Pterostichus [Platysma] 495

kaschgaricus Jedlika, 1957, Poecilus [Pterostichus] 479

kaschkarowi Semenov, 1898, Carabus 175

kaschmirense Netolitzky, 1920, Bembidion 268



kaschmirensis Reitter, 1922, Yamatosa [Rhysodes] 78

kashgarensis Deuve, 1992, Trechus 333



kashgarica Tschitschérine, 1903, Cylindera [Cicindela] 110

kashgaricus Jedlika, 1957, Poecilus [Pterostichus] 481

kashmirana Straneo, 1984, Pterostichus [Amolopsa] 494

kashmiranus J. Balfour‑Browne, 1944, Platynectes 44

kashmirense Breuning, 1927, Calosoma 120

kashmirensis Schauberger, 1927, Acinopus 368

kashmirensis Bates, 1889, Harpalus 377

kashmirensis Vazirani, 1970, Herophydrus [Hydroporus] 71

kashmirensis Vazirani, 1964, Hydronebrius [Amphizoa] 41

kashmirensis Bates, 1889, Laemostenus [Pristonychus] 538

kashmirensis Andrewes, 1927, Leistus 80



kashmirensis Morvan, 1980, Molops 474

kashmirensis Régimbart, 1899, Nebrioporus [Hydroporus] 65

kashmirica Jakobson, 1908, Archicolliuris [Colliuris] 443

kashmiricus Jedlika, 1938, Amerizus [Tiruka] 241

kashmiricus Grundmann, 1955, Chlaenius [Stenochlaenius] 355

kassandra Kleinfeld, 1999, Carabus 125

kassbergensis Schauberger, 1921, Pterostichus [Calopterus] 488

kastschenkoi Brancsik, 1899, Carabus 146

kasyi Mandl, 1967, Cosmodela [Cicindela] 108

kasyi Wewalka, 1984, Hydroporus 62



kasyi Jedlika, 1967, Lebia 429

kaszabi V. B. Guéorguiev, 1972, Agabus 36



kaszabi Fassati, 1952, Amara 568

kaszabi Jedlika, 1953, Amara 559

kaszabi Jedlika, 1961, Bembidion 257

kaszabi Jedlika, 1967, Blethisa 206

kaszabi Jedlika, 1958, Calathus 528

kaszabi Mandl, 1965, Cicindela 104



kaszabi Jedlika, 1964, Cymindis 415

kaszabi Székessy, 1943, Deltomerus 282

kaszabi Jedlika, 1952, Dianella 437

kaszabi Jedlika, 1968, Dyschirius 227

kaszabi V. B. Guéorguiev, 1970, Hygrotus [Coelambus] 72

kaszabi Horvatovich, 1972, Leistus 18, 82

kaszabi Shilenkov, 1982, Nebria 88



kaszabi Jedlika, 1956, Nesacinopus [Harpalus] 388

kaszabi Jedlika, 1968, Notiophilus 97

kaszabi Luna de Carvalho, 1967, Protopaussus 211

kaszabi Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 542

kaszabi Jedlika, 1959, Pterostichus 499

kaszabianus Casale, 1988, Pseudotaphoxenus 541

kaszabiella Hieke, 1983, Amara 552

kaszonensis Csiki, 1946, Scarodytes 68

kataevi Jaeger & Wrase, 1994, Bradycellus 401

kataevi Zamotajlov, 1988, Deltomerus 282

kataevi Kryzhanovskij, 1989, Pterostichus 494

kataevi Belousov, 1987, Trechus 333



katajevi Gottwald, 1989, Carabus 146

katashinensis Habu, 1958, Pterostichus 502



katavensis Jedlika, 1957, Anisodactylus [Harpalus] 361

katharinae Reitter, 1898, Carabus 181

kathmanduense Kirschenhofer, 1994, Lachnoderma 438

katkae Deuve, 1994, Carabus 175

katoi Uéno, 1959, Kurasawatrechus 313

katoi Kasahara, 1992, Pterostichus 502



katomelas G. Hauser, 1921, Carabus [Acoptolabrus] 122

katorum Uéno, 1983, Oroblemus 316

katranicus Belousov & Kabak, 1996, Trechus 333

katsumai Imura & Mizusawa, 1995, Carabus 177

katsumii Deuve & Ohshima, 1992, Carabus 143

kaufmanni Ganglbauer, 1900, Anophthalmus [Trechus] 291

kaufmanni Solsky, 1874, Carabus 145

kaufmanni Ganglbauer, 1897 (as 1896), Tapinopterus 520



kaulbacki Baliani, 1938, Amara 550

kautianus Apfelbeck, 1907, Duvalius [Anophthalmus] 301



kautianus Apfelbeck, 1912, Duvalius [Anophthalmus] 301

kavani Jedlika, 1951, Lebia 430

kavani Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 541

kawaguchii Uéno, 1973, Kurasawatrechus 313

kawanoi Kamiyoshi & Mizoguchi, 1960, Carabus [Apotomopterus] 177

kawanoi Ishida, 1961, Pterostichus 506

kawanoi Uéno, 1975, Trechiama 322

kawasawai Uéno, 1951, Rakantrechus 319

kaynashensis Schweiger, 1967, Pterostichus 491


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