Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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lucasseni Régimbart, 1893, Laccophilus 76

lucens Baliani, 1943, Amara 567

lucens Schaum, 1856, Carabus 184

lucens Bates, 1889, Lampetes [Lamprophonus] 387

lucensis Scheibel, 1935, Orotrechus [Gspaniella] 317

luceornatus Cavazzuti & Mazzi, 1994, Carabus 149

lucescens Bates, 1873, Lebia 430

luchuensis Brouerius van Nidek, 1957, Cylindera [Cicindela] 111

lucida Duftschmid, 1812, Amara [Carabus] 549

lucidens Putzeys, 1846, Dyschirius 224


lucidipennis Jedlika, 1958, Harpalus [HN] 374

lucidipennis Schauberger, 1934, Harpalus 383

lucidissima Baliani, 1932, Amara 567

lucidissima Sciaky & Pavesi, 1994, Nebria 89

lucidulum Schaum, 1857, Agonum 450

lucidulus Chaudoir, 1880, Distichus [Scarites] 231

lucidum Fairmaire & Laboulbène, 1854, Agonum [Anchomenus] 450

lucidum Faldermann, 1836, Bembidion [Peryphus] 271

lucidum LeConte, 1848, Bembidion [Peryphus] 262

lucidum Mannerheim, 1853, Bembidion 262

lucidus Alluaud, 1930, Acinopus 368

lucidus Andrewes, 1932, Apristus 431

lucidus Reitter, 1900, Dixus [Aristus] 365

lucidus J. Müller, 1903, Duvalius [Anophthalmus] 306



lucidus Putzeys, 1866, Dyschirius 224

lucidus Motschulsky, 1844, Pterostichus [Argutor] 491

lucidus Dejean, 1829, Stenolophus 405

lucidus Jeannel, 1960, Trechus 335

lucidus A. Morawitz, 1863, Trichotichnus [Harpalus] 396

lucifer Mandl, 1955, Carabus 131

lucifer Ledoux & Roux, 1998, Nebria 92

lucifera Mandl, 1971, Cicindela 104

lucifera Cavazzuti, 1996, Cychropsis 202

lucifuga Saulcy, 1862, Reicheia 222

lucifugum Neri & Pavesi, 1989, Bembidion [Ocydromus] 260

lucilla Reitter, 1898, Lebia 428

lucillum Bates, 1883, Bembidion 266



lucorum Geoffroy, 1785, Agonum [Buprestis] 450

luctifer Deuve, 1991, Cychrus 204



luctisonus Csiki, 1927, Carabus 136

luctuosa Ragusa, 1904, Calomera [Cicindela] 101

luctuosa Dejean, 1831, Cephalota [Cicindela] 101

luctuosior Deuve, 1994, Carabus 188

luctuosus Geoffroy, 1785, Agabus [Dyticus] 39

luctuosus Dejean, 1828, Calathus 525

luctuosus Latreille, 1804, Calathus 525

luctuosus Aubé, 1838, Canthydrus [Hydrocanthus] 34

luctuosus Géhin, 1885, Carabus 136

luctuosus Lapouge, 1925, Carabus 136

luctuosus Zoubkoff, 1837, Carabus 188

luctuosus Dejean, 1825, Graphipterus 359

luctuosus Régimbart, 1883, Gyrinus 27

luctuosus Holdhaus, 1904, Microlestes 423

luctuosus Putzeys, 1875, Mochtherus 437

luctuosus Aubé, 1838, Nebrioporus [Hydroporus] 66



luctuosus Desbrochers des Loges, 1906, Poecilus [Pterostichus] 482

luczonicus Aubé, 1838, Hydaticus 53



luczotii Laporte, 1833, Carabus 167

ludia Dejean, 1831, Myriochila [Cicindela] 114

ludingensis Deuve & Vigna Taglianti, 1992, Carabus 128

ludivinae Deuve, 1996, Carabus 125

ludmilae Deuve, 1992, Carabus 175

ludmilae Imura, 1999, Cychrus 204

ludmilae Casale & Sciaky, 1994, Dimorphopatrobus 283

ludmilae Dvoák, 1994, Leistus 82

ludovici Baliani, 1932, Amara 564

ludovici A. Gaudin, 1935, Aphoenops 295

ludovici Schatzmayr, 1929, Pterostichus 512

ludovicus Dietl, 1896, Carabus 179

luedersi Wagner, 1915, Dyschirius 227

luetgensi Beuthin, 1886, Carabus 179

lugdunensis Dejean, 1825, Cylindera [Cicindela] 110

lugdunensis Chaudoir, 1837, Oreonebria [Nebria] 96

lugens Duftschmid, 1812, Agonum [Carabus] 450

lugens Vuillefroy, 1866, Calathus 525

lugens Dejean, 1831, Calomera [Cicindela] 101



lugens Grandi, 1906, Calomera [Cicindela] 100

lugens Ragusa, 1881, Calomera [Cicindela] 101

lugens Chaudoir, 1869, Calosoma 121



lugensis Breuning, 1972, Carabus 165

lugi Morvan, 1982, Pterostichus 510



lugubre Duftschmid, 1812, Agonum [Carabus] 450

lugubre Motschulsky, 1844, Calosoma 120

lugubris Schaum, 1862, Aptinus 212



lugubris Géhin, 1881, Carabus 139

lugubris Fairmaire, 1894, Chlaenius 352

lugubris Motschulsky, 1845, Hydroporus 62

lugubris Preudhomme de Borre, 1871, Hyphydrus 75

lugubris Ragusa, 1887, Nebrioporus [Hydroporus] 66

lugubris Dejean, 1828, Poecilus [Feronia] 480

lugubris Faldermann, 1836, Zabrus 569

lugubriveste Netolitzky, 1943, Bembidion 249

luguense Toledano, 1999, Bembidion 245

luhuo Deuve, 1994, Cychrus 204

luhuoense Toledano, 1998, Bembidion 252

luhuoensis Lassalle, 1999, Acalathus 529

luhuoensis Deuve, 1994, Carabus 189

luigionii J. Müller, 1931, Bembidion 264

luigionii Leoni, 1908, Calathus 524

luigionii J. Müller, 1934, Cymindis 412



luigionii Dodero, 1899, Lymnastis 274

luigionii Straneo, 1934, Pterostichus 503

lukjanovitshi Lutshnik, 1927, Pterostichus [Platysma] 515

lulinense Habu, 1973, Bembidion 259

lulinensis Habu, 1979, Trichotichnus 396

lumbaris Mannerheim, 1825, Harpalus 378

lumensis Apfelbeck, 1905, Pterostichus 515

lunaris Duftschmid, 1812, Cymindis [Lebia] 411

lunatum Duftschmid, 1812, Bembidion [Elaphrus] 243

lunatus Fabricius, 1775, Callistus [Carabus] 347

lunatus Régimbart, 1899, Hydaticus 54

lundbergi Jeanne, 1996, Laemostenus [Antisphodrus] 534

lundbladi Colas, 1938, Calathus 527

lundbladi Falkenström, 1939, Haliplus 33

lundbladi Falkenström, 1939, Hydroporus 60

lundbladi Falkenström, 1938, Hydrotarsus 65

lundbladi Jeannel, 1938, Orthomus 476

lundbladi Jeannel, 1938, Syntomus [Metabletus] 433

lundbladi Jeannel, 1938, Trechus 335

lungtschuanensis G. Hauser, 1919, Carabus [Coptolabrus] 143

lunicollis Schiødte, 1837, Amara 550

luniferum Andrewes, 1924, Bembidion 251

luniferum Andrewes, 1935, Cymbionotum 235



luniger Kolenati, 1845, Agabus 39

luntaka Andrewes, 1924, Bembidion 261

lunulata Fabricius, 1781, Calomera [Cicindela] 101

lunulatum Geoffroy, 1785, Bembidion [Buprestis] 264



lunulatus Polentz, 1932, Badister 440

lunulatus Fischer von Waldheim, 1829, Platambus [Colymbetes] 44

luojicus Cavazzuti, 1998, Carabus 128

luojiensis Cavazzuti, 1998, Cychrus 204

lupisylvaticus Imura, 1993, Carabus 183

lupoglavensis Knirsch, 1927, Neotrechus 315

luppovae Kryzhanovskij, 1962, Amara [Zabrus] 566

luquensis Deuve, 1990, Carabus 191



lurida Bruneau de Miré, 1952, Tachyura [Tachys] 279

luridicorne Solsky, 1874, Bembidion 247



luridipenne Schaum, 1860, Bembidion 251

luridipennis Germar, 1823, Pogonus [Harpalus] 287



luridipes Gautier des Cottes, 1866, Bembidion 259

luridipes Reiche & Saulcy, 1855, Bembidion 248

luridum Duftschmid, 1812, Bembidion [Elaphrus] 245

luridum Suffrian, 1843, Bembidion 255

luridus Dejean, 1829, Acupalpus 398

luridus Schmidt‑Göbel, 1846, Peliocypas 434



luridus Schaum, 1864, Philodytes [Laccophilus] 78

luridus Rey, 1882, Tachys 276

luristanus Casale, 1988, Laemostenus 536

luschanensis G. Hauser, 1919, Carabus [Apotomopterus] 126

lusciosus Antoine, 1944, Tachys 275



lushanensis Born, 1918, Carabus [Coptolabrus] 141

lushanicus Kleinfeld, 1996, Carabus 127

lusitanica Mandl, 1935, Cicindela 106

lusitanicum Jacquelin du Val, 1852, Bembidion 260

lusitanicum Putzeys, 1845, Bembidion 261

lusitanicus Dejean, 1829, Bradycellus [Acupalpus] 401

lusitanicus Fabricius, 1801, Carabus 167

lusitanicus Wagner, 1932, Chlaenius 352

lusitanicus Sharp, 1882, Cybister 51

lusitanicus Schatzmayr, 1943, Harpalus 381



lusitanicus Scholz, 1916, Metaporus [Hydroporus] 65

lusitanicus Dejean, 1828, Platyderus [Feronia] 523

lusitanicus Jeannel, 1921, Trechus 335

lusitanus Reitter, 1884, Acupalpus 398

lustralis Casey, 1884, Harpalus 372

lutarius Gistel, 1857, Harpalus 386

lutea Andrewes, 1925, Tachyura [Tachys] 277

luteaster Zaitzev, 1906, Agabus 37

luteatus Duftschmid, 1812, Acupalpus [Carabus] 398

luteicollis Gebler, 1848, Rhantus 46

luteicornis Duftschmid, 1812, Harpalus [Carabus] 378

luteicornoides Breit, 1913, Harpalus 386

luteidorsis Roubal, 1917, Badister 441

luteipes Andrewes, 1942, Abacetus 347



luteipes Andrewes, 1935, Asaphidion 242

luteipes Motschulsky, 1844, Bembidion [Trachypachus] 247

luteipes Chaudoir, 1850, Nebria 91

luteolus Régimbart, 1899, Agabus 39

luteolus Jeannel, 1960, Trechus 335



lutescens Panzer, 1796, Agonum [Carabus] 453

lutescens Reitter, 1888, Amara [Harpalodema] 562

lutescens Jedlika, 1940, Platynus [Colpodes] 465

lutevanus Jeannel, 1919, Duvalius [Trechus] 305

luticola Chaudoir, 1850, Dyschirius 226

lutosus Crotch, 1873, Ilybius [Gaurodytes] 42

lutschniki Baliani, 1937, Amara 559

lutschniki Jedlika, 1962, Pterostichus 509

lutshniki Zamotajlov, 1988, Carterus 364

lutshniki Matalin, 1999, Cicindela 107

lutshniki Roubal, 1928, Eobroscus [Broscus] 237


lutshniki Schauberger, 1932, Harpalus 378

lutshniki Arnoldi & Kryzhanovskij, 1964, Lymnastis 274



lutshniki Roubal, 1928, Patrobus 285

lutshniki Kryzhanovskij, 1990, Pogonus 287

lutshniki Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 541

lutshniki Belousov, 1987, Trechus 335



lutshnikianus Basilewsky, 1937, Carabus 134

lutshnikianus Kataev, 1987, Harpalus 378

lutshnikianus Bogachev & Kurnakov, 1958, Pterostichus 500



lutzi Reitter, 1908, Hygrotus [Hydroporus] 72

luxatus Audinet‑Serville, 1821, Masoreus [Harpalus] 357

luxuriosus Motschulsky, 1844, Carabus 157

lyauteyi Alluaud, 1923, Laemostenus 535



lycaon Linder, 1860, Harpalus 384

lycicum Jeanne, 1996, Bembidion [Philochtus] 264

lycicus Breuning, 1932, Carabus 186

lycius Kataev & Wrase, 1997, Harpalus 372

lycius Ganglbauer, 1915, Zabrus 571

lydius Kataev & Wrase, 1997, Harpalus 372



lyellii Wollaston, 1857, Nebrioporus [Hydroporus] 67

lynx Chaudoir, 1856, Chlaenius 353



lyoni Gory, 1838, Neolaphyra [Cicindela] 114

lyonii Vigors, 1825, Cephalota [Cicindela] 102



lyperoides Gautier des Cottes, 1869, Pterostichus [Omaseus] 500

lyrata Hieke, 1981, Amara 553



lyratus Klug, 1832, Chlaenius 355

lyratus Motschulsky, 1844, Harpalus 384

lyratus Swartz, 1808, Hyphydrus 74



lyrodera Motschulsky, 1860, Nebria 94

lyroderus Chaudoir, 1846, Pterostichus [Feronia] 498

lysander Andrewes, 1935, Bembidion 20, 256

lysholmi Munster, 1930, Bembidion 261

maa Erwin & Kavanaugh, 1999, Tachyta 277

maacki A. Morawitz, 1862, Carabus 160

macairei Dejean, 1826, Carabus 180



macedo Schaufuss, 1882, Amblystomus 360

macedo Jeannel, 1919, Duvalius [Trechus] 308

macedonica Maan, 1938, Nebria 84

macedonicus Maan, 1935, Calathus 525

macedonicus J. Müller, 1917, Duvalius [Trechus] 308



macedonicus Apfelbeck, 1918, Molops 473

macedonicus V. Guéorguiev & B. Guéorguiev, 1997, Pachycarus 367

macedonicus Apfelbeck, 1918, Pterostichus 491

macedonicus Jedlika, 1923, Pterostichus 491

macedonicus V. B. Guéorguiev, 1959, Stictotarsus [Potamonectes] 69



macedoniens Jureek, 1928, Carabus 138

maceki Deuve, 1992, Trechus 335

macellus K. Daniel, 1903, Reicheiodes [Dyschirius] 230

macer Jedlika, 1940, Platynus [Colpodes] 465

macer Jedlika, 1957, Pseudotaphoxenus [Taphoxenus] 542

macer Marsham, 1802, Pterostichus [Carabus] 485

macer Uéno, 1987, Rakantrechus 319

macer Habu, 1978, Synuchus 545

macerellus Lorenz, 1998, Platynus [Colpodes] 465

machadoensis Franz, 1984, Trechus 335



machadoi Jeanne, 1970, Calathus 525

machadoi Jeanne, 1971, Harpalus 381

machadoi Jeannel, 1941, Trechus 335

machaon Breuning, 1950, Carabus 124

machardi Jeanne, 1982, Speluncarius 516

machardi Jeanne, 1976, Trechus 335

machardianus P. Machard, 1986, Carabus 193

machardiellus Deuve, 1991, Carabus 157

machiko Uéno, 1962, Epaphiopsis 309



machulkai Roubal, 1929, Duvalius 306

machulkai Jedlika, 1931, Pterostichus 515

macilentior Deuve, 1996, Carabus 189

macilentus Lapouge, 1899, Carabus [HN] 161



macilentus Petri, 1912, Carabus 167

mackaensis Cavazzuti & Korell, 1991, Carabus 197

macleayi Dejean, 1826, Carabus 153



macleayi Andrewes, 1919, Chlaenius 354

macleayi Sloane, 1907, Dolichoctis [Mochtherus] 436

macleayi Brullé, 1834, Neocollyris [Collyris] 117

macra Bates, 1883, Amara [Bradytus] 554

macra Andrewes, 1934, Pristosia [Calathus] 531

macretus Kraatz, 1877, Carabus 163

macrocephala Breit, 1914, Nebria 94

macrocephala Motschulsky, 1844, Nebria 95



macrocephaloides Jeanne, 1972, Carabus 167

macrocephalus Dejean, 1826, Carabus 167



macrocephalus Bokor, 1921, Duvalius [Trechus] 300

macrocephalus Gschwendtner, 1923, Hydroporus 61

macrocephalus Habu, 1955, Pterostichus 498



macrocephalus Heer, 1837, Trechus 331

macrocerus Chaudoir, 1876, Brachinus 216



macrocerus Chaudoir, 1856, Chlaenius 355

macrodera K. Daniel, 1903, Oreonebria [Nebria] 96

macrodera Chaudoir, 1860, Tricondyla 118

macroderus Chaudoir, 1850, Dyschirius 226



macroderus Jeannel, 1928, Trechus 332

macrogenys Bates, 1883, Pterostichus 498

macrogona Bates, 1873, Nebria 94

macrogonus Chaudoir, 1847, Carabus 193

macrograptina Acciavatti & Pearson, 1989, Calomera [Cicindela] 101

macromandibularis Habu & Tanaka, 1956, Diplocheila [Submera] 439

macromelus Jeannel, 1926, Anophthalmus 292

macronota Solsky, 1875, Amara [Curtonotus] 560

macronotus Tschitschérine, 1893, Harpalus 378

macrophthalmum Reitter, 1890, Bembidion 246

macrophthalmus Székessy, 1938, Lymnastis 274

macrophthalmus Poppius, 1906, Pterostichus [Cryobius] 490

macrops Antoine, 1951, Laemostenus [Sphodroides] 539

macrops Jeannel, 1927, Trechus 335

macropterum J. R. Sahlberg, 1880, Bembidion 243

macropus Chaudoir, 1877, Carabus 200

macropus Chaudoir, 1861, Laemostenus [Sphodrus] 533



Macrotachys Uéno, 1953, Porotachys 274

macrothorax Poppius, 1906, Pterostichus [Cryobius] 491



maculala DeGeer, 1774, Cicindela 105

macularis Fischer von Waldheim, 1824, Cymindis 417



macularis Stephens, 1827, Cymindis [Tarus] 412

maculata Nietner, 1856, Coryza [Clivina] 221

maculata Klug, 1832, Trichis 439

maculatipenne Pic, 1901, Omophron 208

maculatum Dejean, 1831, Bembidion 270

maculatus Schaum, 1860, Acupalpus [Stenolophus] 398

maculatus Klug, 1832, Brachinus 212

maculatus Motschulsky, 1860, Haliplus 32



maculatus Sharp, 1882, Hydrovatus 70

maculatus Babington, 1841, Hyphydrus 74

maculatus Linné, 1758, Platambus [Dytiscus] 44

maculicollis Reitter, 1894, Dicheirotrichus [Trichocellus] 403



maculicornis Maan, 1935, Calathus 528

maculicornis Stephens, 1835, Harpalus 381

maculicornis Chaudoir, 1850, Neocollyris [Collyris] 117

maculicornis Duftschmid, 1812, Parophonus [Carabus] 393

maculicornis Chaudoir, 1846, Trechus 335

maculifrons Ménétriés, 1849, Anisodactylus 361

maculipennis Habu, 1975, Dyschirius 229

maculipennis Schaum, 1864, Haliplus 32

maculipennis Bedel, 1897, Trechus 329

maculipes Grimmer, 1841, Amara 547

maculosus A. Zimmermann, 1919, Hydrovatus 70

maculosus Gschwendtner, 1931, Liodessus [Bidessus] 57


macurebus Géhin, 1885, Carabus 161

madagascariense Chaudoir, 1876, Bembidion 258

madagascariensis Putzeys, 1846, Clivina 220



madagascariensis Aubé, 1838, Cybister 51

madara Habu, 1957, Coptodera 435

madefactus Cavazzuti, 1997, Carabus 126



madeirensis Serrano & Borges, 1995, Trechus 331

maderae Fabricius, 1775, Calosoma [Carabus] 121

maderae Wollaston, 1857, Cymindis 414

maderae Schauberger, 1932, Nesarpalus 24, 388



maderae Franz, 1981, Trechus 331

maderensis Wollaston, 1854, Agabus 40

maderensis Mateu, 1958, Bradycellus 400

maderensis Wollaston, 1854, Olisthopus 461

maderensis Csiki, 1928, Trechus 335

maderi Winkler, 1914, Anophthalmus [Trechus] 292



maderi Born, 1915, Carabus 196

maderi J. Müller, 1933, Molops 473

maderi Jedlika, 1938, Pterostichus 501

maderianus Mandl, 1957, Carabus 173

madidus Paykull, 1790, Molops [Carabus] 474

madidus Fabricius, 1775, Pterostichus [Carabus] 514

maeander Fischer von Waldheim, 1820, Carabus 154

maeander Uéno, 1959, Rakantrechus 319



maeklini Poppius, 1906, Pterostichus [Cryobius] 491

maeklini Zaitzev, 1910, Suphrodytes [Hydroporus] 70

maeoticum Kolenati, 1845, Bembidion [Leja] 249

maeoticus Kolenati, 1845, Badister 441

maeoticus Fischer von Waldheim, 1823, Carabus 182

maeoticus Motschulsky, 1850, Pterostichus [Argutor] 507

maestroae Fresneda & Hernando, 1987, Ilybius [Agabus] 41

maetai Ishikawa & Kubota, 1994, Carabus 178



magadanensis Deuve & Dolin, 1991, Carabus 134

magadanicus Obydov, 1999, Carabus 172

magar J. Schmidt, 1998, Aparupa 456

magdalenae Abeille de Perrin, 1869, Duvalius [Anophthalmus] 301

magdalenae Abeille de Perrin, 1894, Pseudanillus [Anillus] 238

magdalenai Pierre Meyer & Mollard, 1996, Carabus 167

magdelainei Bleuse, 1914, Carabus 195

magdelainei Jeannel, 1914, Duvalius [Trechus] 304



magdeleinei Lapouge, 1922, Carabus 179

magellense Schauberger, 1922, Bembidion 269

magellensis Leoni, 1907, Laemostenus 536

magellensis A. Fiori, 1899, Zabrus 570

maghrebica Antoine, 1941, Amara 551


maghrebicum Antoine, 1941, Bembidion 267

maghrebinus Mazzoldi & Toledo, 1998, Stictotarsus 69

maginianus Magrini & Vanni, 1984, Duvalius 303


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