Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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maginii Magrini, 1976, Duvalius 302

maginii Magrini & Vanni, 1990, Typhloreicheia 223

magistrettianus Schatzmayr, 1940, Duvalius 304

magistrettii De Monte, 1947, Bembidion 248

magistrettii Binaghi, 1939, Duvalius 306

magistrettii Focarile, 1949, Trechus 335

maglajensis Apfelbeck, 1908, Duvalius [Trechus] 306



maglajensis Apfelbeck, 1912, Duvalius [Anophthalmus] 306

maglianoi Giordan & Raffaldi, 1983, Duvalius 304

maglicensis Winkler, 1933, Duvalius 299

magniceps Hieke, 1993, Amara 562

magniceps Reitter, 1898, Trechus 335

magnicollis Csiki, 1930, Abax 470

magnicollis Tschitschérine, 1894, Amara 550

magnicollis Motschulsky, 1866, Leistus 83

magnicollis Motschulsky, 1865, Platynus [Limodromus] 464

magnifica W. Horn, 1905, Cicindela 107

magnificens Kleinfeld, 1999, Carabus 141

magnificus Kraatz, 1878, Carabus 173

magnificus Kurnakov, 1959, Jeannelius 312

magniventris Cavazzuti, 1999, Carabus 189

magnodentatus Schauberger, 1929, Harpalus 386

magnus Feng, 1936, Colymbetes 45

magnus Giordan & Raffaldi, 1989, Duvalius 302

magnus Motschulsky, 1860, Lesticus [Omaseus] 471

magnus Bates, 1873, Platynus [Anchomenus] 463

magnus Dejean, 1828, Pterostichus [Feronia] 488

magoides Straneo, 1937, Pterostichus [Feronia] 506



magrettii Gestro, 1923, Paussus 211

magrinii Vanni & Lanza, 1983, Duvalius 304



maguini Auvray, 1994, Carabus 157

magus Mannerheim, 1825, Pterostichus [Omaseus] 506

magyari Jedlika, 1960, Brachinus 216

magyarica Roeschke, 1891, Cicindela 105

mahoae Uéno, 1980, Trechiama 323

mahouxi Ledoux, 1972, Carabus 147



mahouxi Morvan, 1976, Trechus 329

maichensis Lafer, 1979, Pterostichus 495



maidantalensis Deuve & Dolin, 1997, Carabus 146

maifredii Ghidini, 1962, Duvalius 303

maillei Solier, 1835, Carabus 152



maillei Dejean, 1831, Chlaenius [Dinodes] 352

mainardii Straneo, 1934, Pterostichus 508

maindroni Bedel, 1907, Amara 562

maindroni Alluaud, 1919, Halocoryza 221

maindroni Régimbart, 1897, Laccophilus 76

maindroni W. Horn, 1897, Salpingophora [Cicindela] 114

maior Busulini, 1959, Orotrechus 317

mairei Peyerimhoff, 1922, Amara 552

mairei Antoine, 1925, Carabus 152

mairei Peyerimhoff, 1928, Harpalus 378

mairei Peyerimhoff, 1920, Laemostenus 534

maireyi Jeannel, 1953, Trichaphaenops 344

maironei Cavazzuti, 1994, Carabus 147

maisaicus Belousov & Kabak, 1994, Trechus 335

maiyasanus Bates, 1873, Carabus 177

majalis Mandl, 1935, Cicindela 106



majellae Schatzmayr, 1929, Pterostichus 503

majellensis Jeannel, 1937, Laemostenus [Cryptotrichus] 536

major Semenov, 1889, Acalathus 529

major Zamotajlov & Wrase, 1997, Chinapenetretus 281

major Giordan & Raffaldi, 1989, Duvalius 22, 304

major Knirsch, 1913, Duvalius [Anophthalmus] 299

major Motschulsky, 1850, Harpalus [Erpeinus] 379

major Régimbart, 1899, Hydaticus 53

major Sharp, 1882, Hydroglyphus [Bidessus] 56



major Sharp, 1882, Hyphydrus 74

major Bates, 1891, Kareya [Platymetopus] 387



major Brullé, 1834, Laemostenus [Pristonychus] 537

major L. Miller, 1868, Lionychus [Apristus] 432

major Nilsson, 1998, Platynectes 44



major Wasmann, 1904, Platyrhopalus 211

major Motschulsky, 1844, Poecilus [Argutor] 480

major Tschitschérine, 1895, Pseudotaphoxenus [Taphoxenus] 542

major Lindroth, 1956, Synuchus 545



majorbucephalus Kraatz, 1879, Carabus 185

majoricensis Schaufuss, 1882, Amblystomus 360

majoricensis Perez Arcas, 1869, Percus 478

majoricus Jeanne, 1988, Platyderus 523

majoroides Lassalle, 1999, Acalathus [Procalathus] 529

majus Gyllenhal, 1827, Bembidion 254

majuscula Chaudoir, 1850, Amara [Bradytus] 554



majusculus Semenov, 1897, Carabus 147

majusculus Uéno, 1999, Shenaphaenops 320

majusculus K. Daniel, 1902, Trechus 335

makaluensis Perrault, 1985, Leistus 80



makarovi Kabak, 1990, Carabus 145

makarovi O. Berlov, 1997, Cychrus 19, 202

maklini Munster, 1933, Nebria 89

makolskii Roubal, 1935, Agonum 450

makolskii Roubal, 1923, Amara 548

makolskii Netolitzky, 1929, Bembidion 260



makolskii J. Müller, 1934, Dyschirius 224

makolskii J. Müller, 1933, Pterostichus 493

malabaricum Fleutiaux & Maindron, 1903, Callytron [Cicindela] 101

malaccae Clark, 1863, Hydrovatus [Hydroporus] 70

malaccanus Csiki, 1932, Catascopus 435

malacensis Hieke, 1983, Amara 555

malacensis Nègre, 1966, Calathus 525

malacensis Mateu, 1957, Orthomus 477

malachiticolor Reitter, 1896, Carabus 135

malachiticus C. G. Thomson, 1875, Carabus 184

malaisei Andrewes, 1947, Parena [Bothynoptera] 434

malavoltii Moscardini, 1956, Duvalius 304



malawiensis Omer‑Cooper, 1971, Hyphydrus 74

maleki Deuve, 1991, Carabus 190

malhommei Antoine, 1954, Laemostenus [Ceuthosphodrus] 534

malickyi Wewalka, 1977, Scarodytes 68



malissorum Apfelbeck, 1919, Carabus 163

malissorum Apfelbeck, 1918, Deltomerus 283



malissorum Apfelbeck, 1918, Molops 474

malissorum J. Müller, 1914, Neotrechus [Trechus] 315

malissorum Winkler, 1933, Omphreus 446

malissorum Apfelbeck, 1905, Pterostichus 515

maljushenkoanus Breuning, 1926, Carabus 131

maljushenkoensis Breuning, 1967, Carabus 198



maljushenkoi Koenig, 1904, Carabus [Tribax] 201

malkovskyi Kabak, 1990, Carabus 150



mallarodensis A. Gaudin, 1947, Hydraphaenops 311

mallaszi Jeannel, 1929, Trechus 22, 335

mallaszianus P. Moravec & Lompe, nom. nov., Trechus 22, 335

mallaszii Csiki, 1901, Duvalius [Trechus] 299

malloryi Andrewes, 1930, Zabrus 571

malomvicensis Ganglbauer, 1900, Duvalius [Trechus] 23, 299

malomvizensis Csiki, 1902, Duvalius [Anophthalmus] 299

malshentianus Apfelbeck, 1918, Molops 473

malyi Obenberger, 1914, Anchomenus [Idiochroma] 455

mamillatum Netolitzky, 1943, Bembidion 245

mammalis Uéno, 1987, Trechiama 323



mamorensis Antoine, 1950, Cicindela 106

manap Bezina & Kabak, 1993, Carabus 150

manas Kataev, 1990, Harpalus 379


manasluensis Hieke, 1997, Amara 564

manasluensis J. Schmidt, 1999, Calathus 525

manasluensis Dvoák, 1994, Leistus 80

manca Jedlika, 1957, Amara 550

manca Ganglbauer, 1900, Cymindis 413

mancinii Schatzmayr, 1925, Stomis 518



mancinii Jeannel, 1921, Trechus 334

mancipium Duftschmid, 1812, Amara [Carabus] 565

mancus Bonelli, 1813, Scarites 232

mandarin Netolitzky, 1939, Bembidion 267



mandarina Baliani, 1932, Amara 557

mandarina Fleutiaux, 1886, Cylindera [Cicindela] 111

mandarinellus Bates, 1892, Craspedophorus 447



mandarinus Born, 1899, Carabus [Coptolabrus] 144

mandarinus Schaum, 1854, Craspedophorus [Isotarsus] 447

mandarinus Bänninger, 1928, Scarites 234

mandarinus Sciaky & Pavesi, 1995, Trechus 335

manderstjernae Ballion, 1871, Calosoma [Callisthenes] 118

manderstjernae Ballion, 1871, Lebia 427

manderstjerni Motschulsky, 1852, Carabus 157

mandibularis Uéno, 1978, Allotrechiama 290

mandibularis Fischer von Waldheim, 1828, Carabus 199

mandibularis Reiche, 1862, Carterus 364

mandibularis Jeannel, 1930, Duvalius 299

mandibularis Kurnakov, 1961, Lindrothius 529

mandibularis Bates, 1872, Nebria 86



mandibularis Minowa, 1932, Scarites 234

mandli Jedlika, 1933, Agonum 450



mandli Jedlika, 1957, Amara 551

mandli Jedlika, 1963, Amara [HN] 566

mandli Netolitzky, 1932, Bembidion 244



mandli Jedlika, 1953, Bradycellus [Tachycellus] 402

mandli Mandl, 1967, Calomera [Cicindela] 100

mandli Paulus, 1971, Cychropsis 202

mandli Jedlika, 1963, Cymbionotum [Graniger] 235

mandli Jedlika, 1963, Scarites 232

mandli Probst, 1986, Therates 116

mandlianum Fassati, 1957, Bembidion 246

mandriolae Ganglbauer, 1911, Orotrechus [Trechus] 317

mandschurensis Breuning, 1926, Carabus 199

mandschuricus Jedlika, 1942, Anisodactylus 361

mandschuricus Semenov, 1898, Carabus 144

mandschuricus Breit, 1912, Chlaenius 348

mandsuricus Guignot, 1956, Agabus [Ilybius] 37



manducus Schaufuss, 1882, Graptodytes [Hydroporus] 59

mandzhurica Lutshnik, 1935, Amara 548

mandzhuricus Lutshnik, 1916, Pterostichus [Platysma] 493

manensis Belousov & Kabak, 1994, Trechus 335

manevali Jeannel, 1942, Amara 548

manfredeggeri Kirschenhofer, 1997, Cymindis 411

manfredschmidi Kirschenhofer, 1985, Bembidion 256

mangartensis Schauberger, 1936, Trichotichnus 396

manhartensis Meschnigg, 1943, Anophthalmus 292

manicatum Andrewes, 1935, Bembidion 258

manifestus Kraatz, 1881, Carabus 183

maniganggo Deuve, 1992, Carabus 191

manii Vazirani, 1970, Nebrioporus [Potamonectes] 66

manlius Imura, 1995, Carabus 126



mannerheimi Dejean, 1826, Carabus 199

mannerheimi Gebler, 1843, Cymindis 416

mannerheimi Kolenati, 1845, Laemostenus [Pristonychus] 538

mannerheimii Chaudoir, 1842, Anthia 347

mannerheimii C. R. Sahlberg, 1827, Bembidion 264

mannerheimii Faldermann, 1836, Cylindera [Cicindela] 110

mannerheimii R. F. Sahlberg, 1844, Dicheirotrichus [Bradycellus] 402

mannerheimii Dejean, 1828, Platynus [Anchomenus] 463

mannerheimii Dejean, 1831, Pterostichus [Feronia] 493

mannheimsi Mandl, 1954, Neocollyris [Collyris] 116

manoianus Imura, 1989, Carabus [Damaster] 122

manriquianus Wollaston, 1862, Licinus 442

mansuetus Casey, 1914, Harpalus 379

manto Holdhaus, 1924, Typhloreicheia [Reicheia] 223

manzoniana Monzini, 1995, Boldoriella 296



maoershanensis Li, 1992, Carabus [Coptolabrus] 144

maoershanensis Li, 1992, Carabus 183

maoershanicus Cavazzuti, 1995, Carabus 126

maolanensis Imura, Zhou & Su, 1999, Carabus 126

maoniuensis Deuve, 1996, Cychropsis 201

maotaiophilus Deuve, 1996, Carabus 128

maowenensis Deuve, 1995, Trechus 335



maracandensis Solsky, 1874, Cicindela 108

maracandica Lutshnik, 1922, Amara [Pterostichus] 558

marani Knirsch, 1930, Duvalius [Paraduvalius] 308

marani Kult, 1949, Dyschirius 226

marani Kult, 1939, Lebia 429

marani Mlyná, 1977, Molops 472

marani Jedlika, 1935, Platynus [Colpodes] 465

marani Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 541

marani Jedlika, 1967, Pterostichus 491

marani Jedlika, 1932, Tachyura [Tachys] 279



marani V. Guéorguiev & B. Guéorguiev, 1999, Tapinopterus 520

marani Jedlika, 1968, Xestagonum [Agonum] 468

maraniana Kult, 1943, Lebia 429

marcellae Colas, 1943, Calathus 526



marcelliceps Roubal, 1928, Philorhizus [Dromius] 425

marcelloi Busulini, 1957, Molops 475



marcens Zoubkoff, 1833, Cephalota [Cicindela] 102

marchali Raynaud, 1971, Carabus 138

marchicus Herbst, 1784, Anchomenus [Carabus] 455

marcilhaci Deuve, 1992, Carabus 159

marcilhaci Deuve, 1992, Cychrus 204

marcilhaci Pham, 1987, Trechus 335

margaliti Wewalka, 1977, Scarodytes 68

margaritophorus G. Hauser, 1912, Carabus [Coptolabrus] 142

margelanicus Belousov & Kabak, 1998, Trechus 335

margellanicus Kraatz, 1884, Craspedonotus 236

margheritae Schatzmayr, 1930, Broscus 236

margianus Kryzhanovskij, 1962, Chilotomus 364

marginalis Belousov & Kabak, 1997, Aepiblemus 289

marginalis Sharp, 1882, Agabus 40

marginalis Fabricius, 1794, Carabus 198



marginalis Motschulsky, 1865, Cymindis [Tarus] 417

marginalis Linné, 1758, Dytiscus 52

marginalis Belousov, 1998, Nannotrechus 315

marginalis Busulini, 1959, Orotrechus 317



marginalis Dejean, 1828, Poecilus 483

marginalis Schilsky, 1889, Suphrodytes [Hydroporus] 70

marginalis Schaum, 1862, Trechus 335

marginalis Morvan, 1999, Xestagonum 468

marginata Heer, 1837, Amara [HN] 565

marginata Stephens, 1831, Amara [Bradytus] 565

marginata Kraatz, 1890, Cephalota [Cicindela] 102

marginata Dupuis, 1912, Coptodera 436



marginata Fischer von Waldheim, 1829, Cymindis 417

marginata Lucas, 1846, Cymindis 413

marginata Geoffroy, 1785, Lebia [Buprestis] 429

marginata Ledoux & Roux, 1995, Nebria 94

marginata Mannerheim, 1823, Pelophila 98

marginata Motschulsky, 1861, Pentagonica 448

marginatum Linné, 1758, Agonum [Carabus] 450

marginatum Gebler, 1829, Calosoma 119

marginatus Lucas, 1846, Acupalpus 398

marginatus Lallemant, 1868, Carabus 161

marginatus Voet, 1778, Carabus [Buprestis] 153

marginatus Sharp, 1873, Dineutus [Dineutes] 26


marginatus Germar, 1824, Gyrinus 27

marginatus Duftschmid, 1805, Hydroporus [Dytiscus] 62



marginatus Descourtilz, 1827, Laemostenus [Carabus] 539

marginatus Gené, 1839, Notiophilus 97

marginatus Matsumura, 1911, Pterostichus [ND] 515

marginatus Petri, 1912, Pterostichus [HN] 488

marginatus Dejean, 1829, Stenolophus 405

marginella Brullé, 1839, Cymindis 414

marginellus Bates, 1873, Badister 441



marginellus Sturm, 1826, Calathus 524

marginellus Régimbart, 1895, Cybister 50

marginellus Fabricius, 1794, Dromius [Carabus] 420

marginellus Gyllenhal, 1827, Harpalus 379



marginellus Stephens, 1828, Harpalus 381

marginellus Schmidt‑Göbel, 1846, Lionychus 432



marginepunctata Dejean, 1826, Calomera [Cicindela] 100

marginepunctata Gistel, 1856, Cicindela 103

marginepunctatus Panzer, 1793, Haliplus [Dytiscus] 31

marginepunctatus Dejean, 1831, Molops [Feronia] 475



marginepunctatus Reitter, 1894, Ophonus [Harpalus] 391

marginepunctatus Heer, 1837, Pterostichus 504

marginicolle Wollaston, 1864, Bembidion 254

marginicolle Reitter, 1889, Clinidium 78



marginicollis Reitter, 1891, Acupalpus 398

marginicollis Fairmaire, 1860, Agabus 40

marginicollis A. Morawitz, 1862, Amara 557

marginicollis Lutshnik, 1915, Amara 554

marginicollis Chaudoir, 1846, Calathus 527



marginicollis Kraatz, 1886, Carabus 153

marginicollis Bates, 1883, Demetrias 419



marginicollis Motschulsky, 1845, Dicheirotrichus [Bradycellus] 403

marginicollis Aubé, 1838, Nebrioporus [Hydroporus] 66

marginicollis Motschulsky, 1854, Pheropsophus 217



marginicollis Balkenohl, 1994, Reicheiodes [Dyschirius] 230

marginicollis Duftschmid, 1812, Synuchus [Carabus] 546

marginicollis Dejean, 1828, Zabrus 569

marginipenne Solsky, 1874, Bembidion 256

marginipennis Motschulsky, 1864, Brachinus 19, 216

marginipennis Laporte, 1835, Copelatus [Colymbetes] 48

marginipennis Fairmaire, 1891, Reflexisphodrus [Antisphodrus] 543



marginithorax Donckier, 1915, Carabus [Coptolabrus] 142

marginiventris Motschulsky, 1864, Brachinus 214

mariae Lutshnik, 1935, Amara 549

mariae Schatzmayr, 1904, Anophthalmus [Trechus] 22, 292

mariae Hummel, 1823, Blemus [Trechus] 296

mariae Kataev, 1997, Harpalus 386

mariae A. Gaudin, 1947, Hydraphaenops 311

mariae Millán & Rocchi, 1991, Nebrioporus [Potamonectes] 67

mariae Mulsant, 1855, Paussus 211

mariae Lutshnik, 1921, Pterostichus [Platysma] 487

mariaerosae Genest, 1983, Aphoenops 295

marianii Focarile, 1949, Trechus 335

mariannae Knirsch, 1924, Duvalius [Duvalites] 299

mariannae Wewalka, 1974, Hydroporus 62

mariettii Cristoforis & Jan, 1837, Carabus 153

marii Magrini & Vanni, 1989, Anillus 238

marii Vanni, Magrini & Pennisi, 1991, Duvalius 304

marilanensis Morvan, 1982, Carabus 162

marina Uéno, 1954, Sakagutia 273



marinae Maux, 1995, Carabus 165

marinae Lassalle & Marcilhac, 1999, Cephalosdrophus 532



marinae Marcilhac, 1993, Pterostichus 515

marinum Schiødte, 1841, Bembidion 263

marinus Strøm, 1783, Aepus [Cicindela] 288

marinus Gyllenhal, 1808, Gyrinus 27

maritima Jeannel, 1942, Amara [Leiromorpha] 563

maritima Schiødte, 1837, Amara 565

maritima Dejean, 1822, Cicindela 106

maritima Korge, 1958, Cymindis 412

maritima Jeanne, 1976, Nebria 93



maritimum Küster, 1847, Bembidion 258

maritimum Motschulsky, 1850, Bembidion 262

maritimum Stephens, 1839, Bembidion [Peryphus] 261



maritimus Schilsky, 1894, Calathus 528

maritimus Motschulsky, 1850, Carabus 199

maritimus Schaum, 1856, Carabus 180

maritimus Lafer, 1989, Dromius 420



maritimus Boheman, 1849, Dyschirius 229

maritimus Fairmaire & Laboulbène, 1855, Haliplus 30

maritimus O. Schneider, 1898, Harpalus 382

maritimus Fairmaire, 1862, Lionychus 432



maritimus Gaubil, 1844, Pterostichus [Feronia] 486

maritimus J. Daniel, 1903, Pterostichus 503

maritimus Jeannel, 1942, Pterostichus [Argutor] [HN] 508

maritimus Helliesen, 1890, Stictotarsus [Deronectes] 69

maritimus Sainte‑Claire Deville, 1907, Trechus 335

markakolensis Belousov & Kabak, 1999, Trechus 335

markamensis Lassalle, 1999, Acalathus 529

markamensis Deuve, 1998, Amerizus 242

markamensis Deuve, 1992, Carabus 175



markamensis Korell & Kleinfeld, 1999, Cychrus 204

markensis Gottwald, 1985, Carabus 131

marklini Gyllenhal, 1813, Hygrotus [Hyphydrus] 72

marmorae Gené, 1839, Cymindis 413

marmoratus Stephens, 1835, Graptodytes [Hydroporus] 59

marmoratus Geoffroy, 1785, Laccophilus [Dyticus] 76

marmoreus Olivier, 1795, Laccophilus [Dytiscus] 76

marocanus Coiffait, 1969, Pseudanillus [Parastylulus] 238

maroccana Fabricius, 1801, Cicindela 106

maroccana Reiche, 1872, Cymindis 414

maroccana Antoine, 1925, Nebria 93



maroccana Fairmaire, 1897, Singilis 438

maroccanum Antoine, 1956, Asaphidion 21, 242

maroccanum Antoine, 1923, Bembidion 259


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