Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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andreae Fabricius, 1787, Bembidion [Carabus] 260

andreae Ménétriés, 1832, Cymindis 411

andreae Jeannel, 1920, Geotrechus 309

andreae Jeannel, 1937, Ocys [Oreocys] 272

andreae Tschitschérine, 1897, Pterostichus [Feronia] 495

andreii Imura, 1998, Carabus 133

andreinii Jeannel, 1937, Anillus 238

andreinii Gestro, 1907, Duvalius [Anophthalmus] 301

andreinii Gestro, 1909, Paussus 211

andreinii Mainardi, 1914, Percus 478

andreinii Dodero, 1922, Pterostichus 511

andreinii Jeannel, 1921, Trechus 338

andreinii Dodero, 1916, Typhloreicheia 223

andreuccii Magrini & Vanni, 1984, Duvalius 301

andrewesi Jeannel, 1923, Agonotrechus 289

andrewesi Baliani, 1932, Amara 547

andrewesi Casale, 1980, Aparupa 456

andrewesi Zamotajlov, 1990, Apatrobus [Apenetretus] 281

andrewesi Jedlika, 1932, Bembidion 263

andrewesi Schauberger, 1932, Chydaeus 361

andrewesi Schauberger, 1932, Harpalus 372

andrewesi Jedlika, 1932, Horniulus 436

andrewesi Jedlika, 1933, Lebia 428

andrewesi Perrault, 1985, Leistus 80

andrewesi Jedlika, 1932, Myas [Trigonognatha] 475

andrewesi Jeannel, 1923, Neoblemus 288

andrewesi Jedlika, 1932, Peliocypas [Risophilus] 434

andrewesi Jedlika, 1931, Platynus [Colpodes] 464

andrewesi Lutshnik, 1934, Pogonus 287

andrewesi Jaeger, 1997, Psychristus 404

andrewesi Jedlika, 1931, Pterostichus 497

andrewesi Landin, 1955, Stenolophus 404

andrewesi Habu, 1955, Synuchus 544

andrewesi Jedlika, 1932, Tachyura [Tachys] 278

andrewesianum Fassati, 1955, Bembidion 270

andrewesianus Schauberger, 1932, Harpalus 385

andrewesianus Jedlika, 1932, Platynus [Colpodes] 464

andrzejuscii Fischer von Waldheim, 1823, Carabus 165

angelae Magrini & Vanni, 1989, Anillus 238

angelae Vanni & Magrini, 1984, Duvalius 303

angelae Magrini, 1984, Trechus 341



angelescui Marcu, 1937, Syntomus [Metabletus] 433

angelicae Reitter, 1892, Trechus 326

angelieri Dajoz, 1971, Pterostichus 498

angelinii Fery & Brancucci, 1997, Deronectes 58



angelinus Reitter, 1899, Carabus [Diocarabus] 149

angenensis Deuve, 1996, Carabus 150



angladensis Jeanne, 1969, Carabus 139

angladensis Coiffait, 1953, Pterostichus [Haptoderus] 490

anglicanum Sharp, 1869, Bembidion 260

anglicus Motschulsky, 1866, Carabus 136

anglicus Schiødte, 1861, Stenolophus 406

angoarnigi Morvan, 1994, Pterostichus 509

angolensis Peschet, 1925, Neohydrocoptus [Hydrocoptus] [delete, act. jaechi] 34

angularis Brullé, 1839, Calathus 526



angularis Chevrolat, 1866, Calathus 526

angularis Putzeys, 1866, Clivina 220

angularis Gyllenhal, 1810, Cymindis 24, 411

angularis Reitter, 1899, Dicheirotrichus [Trichocellus] 402

angularis Rey, 1886, Harpalus 373

angularis Klug, 1834, Hydroglyphus [Hydroporus] 56



angulata Fabricius, 1801, Batoscelis [Carabus] 400

angulata Fabricius, 1798, Calomera [Cicindela] 99

angulata Bänninger, 1949, Nebria 94

angulatoides Puel, 1935, Harpalus 381

angulatus G. Hauser, 1921, Carabus [Damaster] 148

angulatus Fabricius, 1781, Craspedophorus [Carabus] 446



angulatus Motschulsky, 1844, Demetrias 419

angulatus Putzeys, 1878, Harpalus 372

angulatus Piochard de la Brûlerie, 1872, Leistus 81

angulatus Schmidt‑Göbel, 1846, Mochtherus 437

angulatus Belousov, 1998, Troglocimmerites 344

anguleus Morvan, 1972, Pterostichus 509

angulicolle A. Morawitz, 1862, Asaphidion [Tachypus] 242



angulicolle Stierlin, 1878, Asaphidion [Tachypus] 243

angulicollis Jaeger, 1995, Bradycellus 401

angulicollis G. Hauser, 1911, Carabus [Coptolabrus] 142

angulicollis Sella, 1874, Cychrus 202

angulicollis Fairmaire, 1886, Leistus 81

angulicollis J. Müller, 1936, Molops 472

angulicollis Régimbart, 1899, Platambus [Agabus] 43

angulicollis Jeannel, 1954, Trechiama 321

angulifer Belousov & Kabak, 1992, Trechus 326

angulihabitus Kleinfeld, 1998, Carabus 123

angulimaculata Mandl, 1955, Cylindera [Cicindela] 111

angulipennis Meixner, 1925, Arctaphaenops [Trechus] 295



angulipennis Nakane & Ohkura, 1956, Coptodera [Coptoderina] 435

angulipennis Peyron, 1858, Deronectes [Hydroporus] 58

anguliscutis H. Kolbe, 1886, Tenomerga [Cupes] 25

angulosa Bates, 1883, Pentagonica 448



angulosus Reiche, 1872, Platyderus 523

angusi Nilsson, 1994, Agabus 35

angusi Fery & Brancucci, 1990, Deronectes 58

angusi Vondel, 1991, Haliplus 31

angusi Nilsson, 1990, Hydroporus 60

angustata J. R. Sahlberg, 1887, Amara 558

angustata Fischer von Waldheim, 1823, Cylindera [Cicindela] 109

angustata Motschulsky, 1844, Cylindera [Cicindela] 109

angustata A. Costa, 1858, Cymindis 411



angustata Chaudoir, 1842, Drypta 358

angustata Fabricius, 1787, Odacantha [Carabus] 444

angustata Dejean, 1831, Oreonebria [Nebria] 95



angustata Dejean, 1831, Tachyura [Bembidium] 279

angustata Lucas, 1846, Tachyura [Tachys] 277

angustatostriatus N. Ito, 1999, Trichotichnus 394



angustatum Dejean, 1828, Agonum 450

angustatus A. Fiori, 1896, Abax 470

angustatus Dejean, 1831, Brachinus [Aptinus] 212



angustatus L. Redtenbacher, 1844, Calathus 527

angustatus Rambur, 1838, Calathus 524

angustatus Ballion, 1878, Carabus 178

angustatus L. Redtenbacher, 1849, Carabus 181

angustatus Panzer, 1810, Carabus 179

angustatus Petri, 1912, Carabus 179

angustatus Dejean, 1831, Chlaenius 351



angustatus Fischer von Waldheim, 1844, Chlaenius 355

angustatus Heer, 1837, Cychrus [HN] 203

angustatus Hoppe & Hornschuch, 1825, Cychrus 202

angustatus Jeannel, 1928, Duvalius 300

angustatus Ahrens, 1830, Dyschirius [Clivina] 227



angustatus Curtis, 1826, Dytiscus 51

angustatus Stephens, 1828, Dytiscus [HN] 51

angustatus Chaudoir, 1850, Elaphrus 206

angustatus Aubé, 1838, Gyrinus 27

angustatus Sturm, 1835, Hydroporus 60

angustatus Dejean, 1828, Laemostenus [Pristonychus] 532

angustatus Faldermann, 1838, Laemostenus [Pristonychus] [HN] 538

angustatus Stephens, 1839, Microlestes [Dromius] 423

angustatus Motschulsky, 1865, Olisthopus 461

angustatus Duftschmid, 1812, Pterostichus [Carabus] 487

angustatus Hampe, 1850, Trechus 332


angustatus Rambur, 1838, Zabrus 569

angusteforcipatus Antoine, 1941, Microlestes 422

angusticeps Scheibel, 1937, Anophthalmus 292

angusticeps Baehr, 1998, Gunvorita 573



angusticeps Fairmaire, 1897, Lymnastis [Zuphiolum] 274

angusticeps Tanaka, 1962, Synuchus 544

angusticeps Apfelbeck, 1904, Trechus 326

angusticolle Dejean, 1831, Bembidion 245

angusticolle Motschulsky, 1844, Bembidion [Omala] 249

angusticolle Motschulsky, 1850, Bembidion [Metallina] 252

angusticolle Motschulsky, 1860, Bembidion [Peryphus] 255

angusticollis J. R. Sahlberg, 1871, Agabus [Gaurodytes] 38

angusticollis Motschulsky, 1844, Amara [Bradytus] 552

angusticollis J. Müller, 1943, Apotomus 237



angusticollis Dejean, 1831, Calathus 528

angusticollis Boheman, 1848, Calleida 409



angusticollis G. Hauser, 1911, Carabus [Coptolabrus] 141

angusticollis Haller, 1885, Carabus [Procerus] 185

angusticollis Motschulsky, 1859, Carabus 187

angusticollis Chaudoir, 1842, Chlaenius [Dinodes] 352

angusticollis Mannerheim, 1844, Chlaenius 355

angusticollis J. R. Sahlberg, 1880, Dromius 419

angusticollis Putzeys, 1866, Dyschirius 224

angusticollis R. F. Sahlberg, 1844, Elaphrus 206

angusticollis Dejean, 1826, Leistus 81

angusticollis J. Müller, 1921, Ophonus 390

angusticollis Bonelli, 1810, Oreonebria [Alpaeus] 95



angusticollis Schmidt‑Göbel, 1846, Orthogonius 446

angusticollis Mannerheim, 1853, Patrobus 285

angusticollis Motschulsky, 1860, Pelophila 98

angusticollis Fabricius, 1801, Platynus [Carabus] 464

angusticollis Poppius, 1906, Poecilus 483

angusticollis Poppius, 1906, Pogonus 288

angusticollis Fischer von Waldheim, 1823, Pseudotaphoxenus [Sphodrus] 540



angusticollis Reitter, 1894, Tachyura [Tachys] 279

angusticollis Kiesenwetter, 1850, Trechus 326



angusticollis Ganglbauer, 1915, Zabrus 571

angustifrons Régimbart, 1892, Haliplus 31

angustilimbus G. Ochs, 1925, Orectochilus 29

angustioculum Hrka, 1982, Parazuphium 573



angustior Born, 1895, Carabus 168

angustior Kraatz, 1884, Cymindis 416



angustior J. R. Sahlberg, 1875, Harpalus 383

angustior Gyllenhal, 1808, Ilybius [Dytiscus] 41



angustior Baudi di Selve, 1864, Perileptus 23, 288

angustior Poppius, 1904, Poecilus [Pterostichus] 480

angustioricollis Mandl, 1981, Cicindela 105

angustipennis Mandl, 1970, Carabus 162

angustipennis Chaudoir, 1852, Carterus [Ditomus] 364



angustipennis Gestro, 1895, Triaenogenius [Meladroma] 407

angustistylis Tanaka, 1958, Pterostichus 502

angustitarsalis Deuve, 1991, Cychrus 202

angustitarsis Reitter, 1887, Harpalus 373



angustithorax Deuve, 1991, Cychrus 203

angustula Machado, 1987, Licinopsis 540

angustula Motschulsky, 1866, Nebria 94

angustulus Semenov, 1891, Broscus 236

angustulus Wollaston, 1862, Calathus 526

angustulus Haury, 1878, Carabus 138

angustulus J. R. Sahlberg, 1880, Dicheirotrichus 403



angustulus Régimbart, 1899, Ilybius 42

angustulus Zaitzev, 1953, Noterus 34

angustulus Reitter, 1899, Tachys 276

angustus Roeschke, 1898, Carabus 183

angustus Ménétriés, 1832, Carterus [Ditomus] 364

angustus Brullé, 1834, Dromius 420

angustus Reitter, 1883, Leistus 81

angustus Chevrolat, 1840, Licinus 442



angustus Bates, 1876, Oxycentrus 392

angustus Westwood, 1838, Platyrhopalus 211

angustus Gebler, 1829, Pogonistes [Pogonus] 286

angustus Belousov, 1998, Porocimmerites 318

angustus Chaudoir, 1855, Scarites [Taeniolobus] 233

angustus Habu, 1978, Synuchus 545

angustus Uéno, 1985, Trechiama 321



angustus J. R. Sahlberg, 1908, Trechus 326

anhuinus Imura, 1996, Carabus 128



anhweiensis G. Hauser, 1932, Carabus [Coptolabrus] 141

animosa Shilenkov, 1983, Nebria 87

animosus Jeannel, 1962, Trechus 326

anisodactyliformis Solsky, 1874, Harpalus 373

anitae Vazirani, 1969, Hyphoporus 73

anjae Nilsson, 1999, Ilybius 41



anjiensis Li, 1993, Carabus [Coptolabrus] 143

anjuensis Deuve, 1998, Trechus 326

ankarensis Jedlika, 1962, Brachinus 213

ankarensis Breuning, 1961, Carabus 156

annae Briganti, 1976, Duvalius 303

annae Jeannel, 1949, Trichaphaenops 344

annamariae Vanni & Magrini, 1989, Duvalius 301


annamensis Bates, 1889, Anthracus [Acupalpus] 399

annamensis Tschitschérine, 1903, Chlaeminus 347

annamita Régimbart, 1889, Hydroglyphus [Bidessus] 56

annamita Fauvel, 1908, Perigona 448

annamita Régimbart, 1899, Rhantus 46

annandalei W. Horn, 1908, Therates 115

annapurna J. Schmidt, 1998, Rupa 466

annapurnae J. Schmidt, 1999, Calathus 528

annettae Korell, 1982, Carabus 162

annikae Kleinfeld, 1996, Carabus 124



annulata Brullé, 1834, Lebia 427

annulata Andrewes, 1924, Mastax 216



annulatus Zoubkoff, 1833, Agabus [Colymbetes] 38

annulatus Chaudoir, 1837, Ophonus 388

annulicornis Chaudoir, 1842, Brachinus 215



annulicornis Stephens, 1828, Harpalus 381

annulicornis Schmidt‑Göbel, 1846, Tricondyla 117



annulosum Schuler, 1962, Bembidion [Metallina] 251

anodon Chaudoir, 1868, Poecilus [Feronia] 481

anomala Kolenati, 1845, Tachyura [Tachys] 278

anomalum Jedlika, 1958, Bembidion 244

anomalus Perris, 1866, Badister [Olisthopus] 440

anomalus Géhin, 1880, Carabus 171

anomalus Chaudoir, 1878, Chaetobroscus [Broscus] 236

anomostriatus Antoine, 1954, Laemostenus [Pristonychus] 539

anomostriatus Sciaky, 1995, Pterostichus 485

anonymus Ledoux & Roux, 1995, Archastes 79

anophthalmus Reitter, 1886, Speluncarius 516

antaeus F. Hauser & G. Hauser, 1914, Carabus [Coptolabrus] 142

antaiensis Deuve, 1994, Carabus 125



antakyae Korell, 1985, Carabus 188

antecedens Netolitzky, 1922, Bembidion 267

antennarium Duftschmid, 1812, Agonum [Carabus] 452

antennarium Morvan, 1972, Bembidion [Peryphus] 256

antennarius Daffner, 1991, Orotrechus 316



antennata Rosenhauer, 1847, Amara 564

antennatus Régimbart, 1892, Hydroglyphus [Bidessus] 56

antennatus Gautier des Cottes, 1860, Paranchus [Anchomenus] 462

antennatus Poppius, 1906, Pterostichus [Cryobius] 492

anthobia A. Villa & G. B. Villa, 1833, Amara 547



anthophora L. Dufour, 1820, Lebia 429

anthracina Klug, 1834, Cylindera [Cicindela] 109

anthracina A. Morawitz, 1862, Nebria 89

anthracinum Lapouge, 1924, Calosoma [Callipara] 120

anthracinus Solsky, 1874, Agatus [Glycia] 408



anthracinus Sturm, 1836, Gyrinus 27

anthracinus Fairmaire & Laboulbène, 1854, Harpalus 380

anthracinus Illiger, 1798, Pterostichus [Carabus] 510

anthrax Semenov, 1900, Calosoma 118

anticatus Sharp, 1890, Laccophilus 76



antidotus Sharp, 1882, Hydroporus 64

antipai Panin, 1943, Carabus 197

antiquum Geoffroy, 1785, Calosoma [Buprestis] 119

antiquus Dejean, 1826, Carabus 167

antiquus Motschulsky, 1860, Pterostichus [Steroderus] 493

antoinei Straneo, 1951, Abacetus 346

antoinei Puel, 1934, Acinopus 367

antoinei Puel, 1931, Anisodactylus 360



antoinei Puel, 1925, Badister 440

antoinei Puel, 1935, Bembidion 263

antoinei Puel, 1939, Calathus 527

antoinei Schatzmayr, 1944, Cymindis 414

antoinei Paulian & Villers, 1939, Deltomerus 282

antoinei Puel, 1925, Dyschirius 228

antoinei Mateu, 1984, Masoreus 357

antoinei Schauberger, 1926, Ophonus [Harpalus] 389

antoinei Mateu, 1955, Orthomus 477

antoinei Schauberger, 1932, Parophonus [Tachyophonus] 393

antoinei Puel, 1938, Scarites 233

antoinei Mateu, 1952, Trymosternus 410



antoineianus Schauberger, 1929, Ophonus [Harpalus] 391

antonellae Casale, Giachino, Vailati & Vigna Taglianti, 1996, Duvalius 301



antonellii Luigioni, 1921, Carabus 160

antoniae Reitter, 1892, Duvalius [Anophthalmus] 301

antoniettae Magrini & Vanni, 1986, Italodytes 221

antonii Jeannel, 1936, Trechus 326



antonini Deuve, 1997, Carabus 176

antonini Deuve, 1998, Trechus 326

antonowi Semenov, 1891, Cymindis 24, 416

antonowi Tschitschérine, 1898, Harpalus [Brachypangus] 373



antonowi Tschitschérine, 1901, Ophonus [Penthophonus] 389

antonrichteri Heinz, 1983, Carabus 155



antonrichteri Kirschenhofer, 1992, Harpalus 377

antonrichteri Casale, 1988, Laemostenus 536



antracinus J. Jacquet, 1938, Acupalpus 398

antroherponomimus Noesske, 1928, Adriaphaenops [Trechus] 289

antropovi Shilenkov, 1996, Carabus 172

antungensis G. Hauser, 1921, Carabus [Coptolabrus] 144

anubis Bokor, 1913, Duvalius [Anophthalmus] 300



anurus Semenov, 1898, Carabus 148

anxia Tschitschérine, 1898, Amara 547

anxiensis Kleinfeld, 1998, Carabus 123

anxioides Kataev, 1991, Harpalus 373

anxius Lapouge, 1914, Carabus 188

anxius Mannerheim, 1843, Dytiscus 51

anxius Duftschmid, 1812, Harpalus [Carabus] 373

aogashimensis Habu, 1957, Harpalus [Ophonus] 384

aomorii Jedlika, 1956, Agonum [Colpodes] 454

aparupa Andrewes, 1920, Loricera 99

apennina Binaghi, 1942, Loricera 98

apenninicus A. Fiori, 1896, Pterostichus 503

apenninum De Monte, 1943, Bembidion 269



apenninus Depoli, 1927, Carabus 197

apenninus Gestro, 1885, Duvalius [Anophthalmus] 302

apenninus Dejean, 1831, Pterostichus [Feronia] 489

aper Sharp, 1882, Hyphoporus 73

apex Jedlika, 1940, Agonum [Colpodes] 452

apfelbecki Ganglbauer, 1891, Aphaenopsis [Trechus] 293

apfelbecki Müller‑Motzfeld, 1986, Bembidion 249

apfelbecki Jakobson, 1908, Brachinus 214

apfelbecki Born, 1904, Carabus 196

apfelbecki Ganglbauer, 1891, Hydroporus 61

apfelbecki Ganglbauer, 1891, Leistus 83

apfelbecki Holdhaus, 1904, Microlestes 423

apfelbecki Ganglbauer, 1891, Molops 472

apfelbecki Ganglbauer, 1891, Nebria 85

apfelbecki Reitter, 1893, Omphreus 445



apfelbecki Schauberger, 1926, Ophonus [Harpalus] 390

apfelbecki Csiki, 1908, Pterostichus 507

apfelbecki Ganglbauer, 1915, Zabrus 570

apfelbeckianus Jakobson, 1906, Trechus 328

aphaenopsianus Español & E. Vives, 1983, Apoduvalius 295

aphrodisia Baudi di Selve, 1864, Calomera [Cicindela] 101

apicale Ménétriés, 1832, Bembidion 251



apicale Motschulsky, 1844, Bembidion [Notaphus] 253

apicalis Jedlika, 1956, Agatus [Singilis] 408



apicalis Newman, 1833, Calathus 527

apicalis Kraatz, 1890, Cassolaia [Cicindela] 101

apicalis Wiedemann, 1819, Chlaenius [Carabus] 351



apicalis Krausse, 1910, Cicindela 104

apicalis Mannerheim, 1837, Cymindis 417

apicalis Motschulsky, 1865, Cymindis [Tarus] 417

apicalis Motschulsky, 1844, Demetrias 419

apicalis Putzeys, 1846, Dyschirius 224

apicalis C. G. Thomson, 1868, Haliplus 30


apicalis Schmidt‑Göbel, 1846, Hexagonia 407

apicalis Schilsky, 1888, Hydroporus 63

apicalis Sharp, 1873, Ilybius 41



apicalis Brullé, 1834, Miscelus [Leptodactyla] 436

apicalis Breit, 1914, Nebria 91

apicalis Della Beffa, 1909, Perileptus 288

apicalis Küster, 1852, Pogonus 287

apicalis Motschulsky, 1864, Stenolophus [Egadroma] 405

apicalis Wasmann, 1922, Stenorhopalus [Platyrhopalus] 211



apicalis Normand, 1933, Syntomus [Metabletus] 432

apicalis Motschulsky, 1845, Trechus 326

apicedentatus Uéno, 1979, Trechiama 321


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