Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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maroccanum Lapouge, 1924, Calosoma [Campalita] 121

maroccanum Antoine, 1963, Parazuphium 574



maroccanus Schauberger, 1932, Acinopus 368

maroccanus Reitter, 1884, Acupalpus 398

maroccanus Antoine, 1941, Anchomenus [Agonum] 455

maroccanus Antoine, 1954, Apotomus 237

maroccanus Bedel, 1895, Carabus 161



maroccanus Antoine, 1932, Chlaenius 354

maroccanus Antoine, 1934, Dyschirius 224

maroccanus Schatzmayr, 1943, Mesolestes [Metabletus] 421

maroccanus Chaudoir, 1873, Orthomus 477



maroccanus Schauberger, 1932, Parophonus [Tachyophonus] 393

maroccanus Antoine, 1952, Scarites [Parallelomorphus] 232

maroccanus Antoine, 1928, Trechus 331

maroccanus Reiche, 1864, Zabrus 572

maroccanus Schaum, 1864, Zabrus 572

marovighii Fairmaire, 1876, Pedius [Feronia] 478

marquardti Netolitzky, 1920, Bembidion 269

marquardti Reitter, 1898, Carabus 160

marquettensis Casey, 1918, Amara 550

marschallii Dejean, 1826, Nebria 85

marshallana Stephens, 1827, Nebria [Helobia] 89

martensi Hieke, 1981, Amara 564

martensi J. Schmidt, 1999, Calathus 527

martensi Paulus, 1971, Calosoma 120

martensi Heinz, 1994, Cychropsis 202

martensi Casale, 1983, Duvalius 304

martensi Casale, 1985, Gunvorita 573

martensi Wewalka, 1972, Hydaticus 53

martensi Brancucci, 1981, Hydroporus 62



martensi Casale, 1980, Laemostenus [Antisphodrus] 534

martensi Casale, 1981, Laemostenus [Pristonychus] 538

martensi Straneo, 1977, Pterostichus 509

martensi Deuve & Hodebert, 1991, Trechus 335

martensi Freude, 1986, Zabrus 571

martensianus Casale, 1988, Laemostenus 538

marthae Reitter, 1902, Bembidion 266



marthae Reitter, 1908, Bembidion 259

marthae Reitter, 1896, Carabus 198

marthae Barthe, 1908, Cephalota [Cicindela] 102

martinae P. Machard, 1977, Carabus 157

martinae Jeanne, 1996, Duvalius 304

martinellii Daffner, 1987, Orotrechus 317



martinezi Bolívar y Pieltain, 1923, Carabus 140

martinezi Ganglbauer, 1891, Chlaenius 353

martinezi Vuillefroy, 1868, Styracoderus [Feronia] 518

martinezi Jeannel, 1927, Trechus 335

martini Machado, 1984, Orthomus [Eutrichopus] 476



martini Sciaky, 1988, Orthomus [Nesorthomus] [HN] 476

martini Bedel, 1896, Poecilus [Pterostichus] 480

martinii Fairmaire, 1858, Nebrioporus [Hydroporus] 67



martjanovi Lutshnik, 1922, Pterostichus [Platysma] 486

martjanovianus Obydov, 1999, Carabus 172



martjanowi Lutshnik, 1924, Carabus 171

martorelli Kraatz, 1890, Cylindera [Cicindela] 109

martviliensis Retezár & Djavelidze, 1992, Carabus 175

martynovi Semenov & Znojko, 1932, Carabus 147

maruokai Habu, 1973, Agonum [Platynus] 452



marusii Bielz, 1887, Carabus 173

marusiki Obydov, 1997, Carabus 170

marussii De Monte, 1956, Bembidion 261

marysae Letellier, 1979, Carabus 155

masaakii Uéno & Sone, 1994, Trechiama 323

masahiroi Kasahara, 1988, Pterostichus 502

masatakai Uéno, 1983, Trechiama 323

mascarauxi Lapouge, 1918, Carabus 140

mascarauxi Jeannel, 1928, Pterostichus 498

mascarauxi Jeannel, 1947, Pterostichus [Haptoderus] [HN!] 489

mascatensis Régimbart, 1897, Nebrioporus [Hydroporus] 66

mashunensis Péringuey, 1904, Metallica 434

masidai Ishida, 1959, Pterostichus 512

masidai Uéno, 1956, Thalassoduvalius 321

masoreoides Bates, 1878, Harpalus 379

masrina Andrewes, 1924, Nebria 90

massaensis Freude, 1986, Zabrus 570

massageta Emetz, 1972, Cymindis 416



massageta Silfverberg, 1977, Harpalus 375

massagetus Motschulsky, 1844, Carabus 149

massaicum Antoine, 1962, Bembidion [Ocydromus] 261

massanae Jeannel, 1957, Reicheia 222



massaniensis Dokhtouroff, 1887, Calomera [Cicindela] 99

massati Coiffait, 1962, Geotrechus 310



massiliensis Fairmaire, 1850, Somotrichus [Coptodera] 439

massinissa Dieck, 1869, Geocharis [Anillus] 239

masudai Nakane & Ishida, 1959, Gnathaphanus [Iwosiopelus] 362

masumotoi Morita, 1990, Eobroscus 237

masumotoi Tanaka, Morita & Suga, 1987, Pterostichus 498

masuzoi Imura & Satô, 1989, Carabus [Apotomopterus] 126

masuzoi Kasahara, 1989, Paropisthius 98

masuzoi Uéno, 1989, Suzuka 321

mataiurensis J. Müller, 1935, Anophthalmus 293

matalabai Jedlika, 1965, Bembidion 256

matchae Breit, 1917, Neotrechus [Trechus] 315



matchai Jedlika, 1924, Brachinus 212

matchai Jedlika, 1929, Duvalius [Duvalites] 307

matchai Roubal, 1917, Molops 473

matchai Jedlika, 1930, Pterostichus [Haptoderotapinus] 496

matchai Jeannel, 1927, Trechus 331

matejkai Maan, 1939, Duvalius [to leonhardi] 305

matejkai Maan, 1939, Nebria 86

matejkai Všeteka, 1938, Trechus 335

matejkai Maan, 1940, Zabrus 570

mateui Antoine, 1953, Bembidion [Emphanes] 249

mateui Nègre, 1969, Calathus 524

mateui Deuve & Quéinnec, 1994, Ifridytes 532

mateui Omer‑Cooper, 1970, Laccophilus 76

mateui Machado, 1992, Philorhizus 425

mateui Coiffait, 1952, Pterostichus [Haptoderus] 489

mateui Deuve & Quéinnec, 1985, Trechus 335

mateui Novoa, 1980, Zabrus 568



matheyi Born, 1917, Carabus 197

matheyi Pater, 1938, Ophonus [Metophonus] 390

matisi Lafer, 1989, Dyschirius 223



matitsi Roubal, 1917, Amara 568

matocqi Giordan, 1987, Duvalius 304

matrismeae Pawowski, 1972, Trechus 335

matshensis Belousov & Kabak, 1996, Trechus 329

matsudai Habu, 1952, Dromius 420

matsudai Uéno, 1962, Epaphiopsis 309



matsuii Nakane, 1990, Hydrovatus 70

matsuii Uéno, 1999, Kurasawatrechus 313



matsumotoi Satô & Nilsson, 1990, Agabus 36

matsumotoi Uéno, 1984, Trechus 325



matsumurae Csiki, 1929, Amara 554

matsumurai Habu, 1973, Aparupa 456



matsumurai Habu, 1973, Bembidion 20, 271

matsumurai Csiki, 1927, Carabus 148

matsumurai Habu, 1973, Platynus [Agonum] 465

matsumurai Habu, 1973, Pterostichus 510

mattheyi Brancucci, 1980, Hydronebrius 41



matthiesseni Reitter, 1913, Amara [Zabrus] 565

matthiesseni Lutshnik, 1930, Poecilus [Platysma] 482



mattosi Paulino de Oliveira, 1882, Agonum [Platynus] 451

matveichuki O. Berlov, 1996, Pterostichus 487

maublanci Colas, 1942, Brachinus 214

maueri Jeannel, 1955, Trichaphaenops 344

maura Linné, 1758, Cassolaia [Cicindela] 101

maurai Gridelli, 1944, Dicheirotrichus 402

mauretaniae Hrka, 1982, Parazuphium 573

mauretanica Peyerimhoff, 1922, Hydroscapha 25

mauretanicus Bedel, 1914, Brachinus 212



mauriennensis Carret, 1904, Carabus 180

mauritanicum Lucas, 1846, Bembidion 252

mauritanicus Lucas, 1846, Acinopus 368

mauritanicus Dejean, 1829, Amblystomus [Acupalpus] 360

mauritanicus Gaubil, 1844, Harpalus 379

mauritanicus Dejean, 1828, Laemostenus [Pristonychus] 535

mauritanicus Lucas, 1846, Lonchosternus [Oodes] 445

mauritanicus Lucas, 1846, Microlestes [Dromius] 423

mauritanicus Lucas, 1846, Phloeozetus 438

mauritanicus Dejean, 1831, Platytarus [Cymindis] 410

mauritanicus Dejean, 1828, Poecilus [Feronia] 481

mauritanicus Jeannel, 1909, Trechus 335

maurulus Mandl, 1955, Carabus 169

maurum Netolitzky, 1918, Bembidion 257

maurus A. Zimmermann, 1919, Agabus [Gaurodytes] 39

maurus M. F. Adams, 1817, Carabus [Calosoma] 169



maurus Schaufuss, 1883, Dytiscus 52

maurus Tschitschérine, 1897, Harpalus 374

maurus Sharp, 1882, Hydroporus 61

maurus Sturm, 1827, Microlestes [Dromius] 423

maurus Mandl, 1986, Pristomachaerus 356

maurus Fabricius, 1798, Pterostichus [Carabus] 510

maurus Putzeys, 1847, Trechus 336

maurusiacus Mannerheim, 1825, Pterostichus [Steropus] 493

mauxi Deuve, 1991, Cychrus 204


maxillosus Geoffroy, 1785, Dromius [Buprestis] 420

maxillosus Dejean, 1829, Harpalus 373

maxima Motschulsky, 1844, Lebia [Omalomorpha] 428

maximiliani Gistel, 1837, Cicindela 105

maximns Uyttenboogaart, 1937, Syntomus [Metabletus] 432

maximoviczi A. Morawitz, 1863, Calosoma 119



maximus Haury, 1880, Carabus 196

maximus Tschitschérine, 1889, Pterostichus [Haptoderus] 500



mayeri Schatzmayr, 1909, Acupalpus 398

mayeri J. Müller, 1909, Anophthalmus [Trechus] 292

mayeti Brisout de Barneville, 1878, Anillus 238

mayeti Bedel, 1907, Calodromius [Dromius] 419



mayeti Géhin, 1885, Carabus 168

mayeti Verdier, Quézel & Rioux, 1951, Harpalus 379

mayeti Sainte‑Claire Deville, 1935, Pterostichus 488

mayeti Abeille de Perrin, 1892, Scotodipnus 240

mayeti Abeille de Perrin, 1875, Speotrechus [Anophthalmus] 320

mayumiae Deuve & Imura, 1992, Carabus 190

mazametensis Prunier, 1999, Carabus 195

mazandarana Hieke, 1981, Amara 566

mazenderanus Vereschagina, 1987, Pseudotaphoxenus 541

mazurai A. Fleischer, 1910, Carabus 196

mazzii Cavazzuti, 1992, Carabus 182

mazzoldii Fery & Brancucci, 1997, Deronectes 58

meaillensis J. Ochs, 1949, Laemostenus 533

mecynodes Bates, 1890, Carabus 126

medea Kleinfeld, 1999, Carabus 124

medenikensis Knirsch, 1924, Pheggomisetes 318

medialis Sharp, 1882, Laccophilus 76



mediana Horvatovich, 1973, Nebria 87

mediatlantis Antoine, 1941, Amara [Celia] 554

mediatlantis Antoine, 1939, Cymindis 413



mediatlantis Antoine, 1938, Ophonus 390

mediatlantis Antoine, 1951, Poecilus [Parapedius] 481

mediberica Jeanne, 1985, Cymindis 413

medicimons Imura, 1999, Cychrus 204

medicirex Uéno, 1989, Trechiama 323

medicus Kataev, 1993, Harpalus 382



medieuropaeus Schweiger, 1983, Ophonus 390

mediocris Dvoák, 1995, Pterostichus 505



medioinsularis Deuve, 1991, Carabus 149

mediotuberculatus Schaufuss, 1882, Carabus 167

meditabundus Deuve, 1992, Carabus 189

mediterraneum Csiki, 1928, Bembidion 249

mediterraneus Csiki, 1932, Acupalpus 399



mediterraneus Apfelbeck, 1919, Carabus 163

mediterraneus Born, 1906, Carabus 187



medius Reiche, 1872, Acinopus 368

medius Lapouge, 1908, Carabus 129

medius Chaudoir, 1868, Molops 473



medius Dejean, 1828, Poecilus 484

medius Meixner, 1939, Trechus 338



medodiyensis Breuning, 1969, Cychrus 202

medvedevi Kryzhanovskij, 1974, Amara [Curtonotus] 560

medvedevi Kryzhanovskij, 1968, Carabus 146

medvedevi Kryzhanovskij & Emetz, 1973, Cymindis 417

medvedevi Kabak & Kataev, 1993, Dicheirotrichus [Trichocellus] 402

medvedevi Gramma, 1980, Haliplus 31

medvedevi Shilenkov, 1982, Nebria 88



medvedevi Jeannel, 1962, Oroblemites [Trechus] 316

medvedevi Vereschagina, 1988, Pseudotaphoxenus [Taphoxenus] 541

medvedevi Kryzhanovskij & Mikhailov, 1972, Pterostichus 486

megacephala Gebler, 1829, Amara 552

megacephalus P. Rossi, 1794, Acinopus [Carabus] 369

megacephalus Waltl, 1838, Dixus [Ditomus] 365

megacephalus Gschwendtner, 1931, Liodessus [Bidessus] 57



megaloderus Chaudoir, 1867, Leistus 83

megaloderus Sciaky, 1994, Pterostichus 500



megalops Peyerimhoff, 1931, Tachyura 278

megasomus Tarrier, 1972, Carabus [Archicarabus] 129

megaspilota C. A. Dohrn, 1884, Cicindela 105

megaspilum Walker, 1871, Bembidion [Peryphus] 254

meggiolaroi P. Moravec & Lompe, nom. nov., Anophthalmus 22, 292

megrel Belousov, 1992, Duvalius 304

megrelica Shilenkov, 1983, Nebria 87

megrelicus Shilenkov, 1999, Leistus 82



megrelicus Kryzhanovskij, 1989, Pterostichus 495

mehadiensis J. Müller, 1918, Molops 474



mehelyanus Csiki, 1906, Carabus 173

mehelyi Ganglbauer, 1896, Carabus 167

mehmeti S. Battoni & Blumenthal, 1973, Carabus 187

meiguensis Imura, 1998, Carabus 128

meihuanae Imura, 1998, Cychrus 204

meissonnieri Deuve & Mourzine, 1997, Carabus 189

meisteri Reitter, 1885, Pterostichus 515

meixneri Kreissl, 1993, Duvalius 304

meixneri J. Müller, 1912, Neotrechus [Trechus] 315



meixneri Jeannel, 1924, Trechus 22, 335

meixnerianus P. Moravec & Lompe, nom. nov., Trechus 22, 335



melaenus Lutshnik, 1922, Ophonus [Harpalus] 388

melambaphus Fischer von Waldheim, 1832, Carabus 197

melampus Ménétriés, 1849, Chlaenius 350



melampus Duftschmid, 1812, Harpalus [Carabus] 375

melanaria Putzeys, 1846, Clivina 219

melanarius Aubé, 1837, Agabus 40



melanarius Boheman, 1858, Gnathaphanus [Platymetopus] 362

melanarius Sturm, 1835, Hydroporus 62



melanarius Chaudoir, 1871, Orthogonius 446

melanarius Illiger, 1798, Pterostichus [Carabus] 499

melanauges Andrewes, 1922, Dioryche 369

melanchlorus Fischer von Waldheim, 1828, Carabus 164

melancholica Schiødte, 1837, Amara 557

melancholica Fabricius, 1798, Myriochila [Cicindela] 114

melancholicus Fabricius, 1798, Carabus 146

melancholicus Gistel, 1857, Carabus 186

melancholicus Putzeys, 1866, Dyschirius 226

melancholicus Dejean, 1829, Harpalus 370

melancholicus Motschulsky, 1860, Hydroporus 63

melancholicus Schaum, 1864, Zabrus 571

melandetos Imura & Mizusawa, 1996, Carabus 124

melanelytrius G. Hauser, 1921, Carabus [Coptolabrus] 143

melanensis Giordan & Raffaldi, 1999, Duvalius 303

melaneus Bates, 1878, Harpalus 379

melanipes Lienhart, 1936, Carabus 139

melanocephala Dejean, 1825, Cymindis 412



melanocephalum Dejean, 1828, Agonum [Anchomenus] 453

melanocephalus Dejean, 1829, Anthracus [Acupalpus] 400

melanocephalus Linné, 1758, Calathus [Carabus] 527



melanocephalus Stephens, 1828, Haliplus 33

melanocephalus Stephens, 1828, Ocys 272

melanocephalus Dejean, 1825, Philorhizus [Dromius] 425



melanocephalus Dejean, 1829, Stenolophus 406

melanocephalus Kolenati, 1845, Trechus 335



melanocera Tschitschérine, 1899, Amara 550

melanochlora Lutshnik, 1933, Amara 550

melanochrus A. Morawitz, 1886, Carabus 134

melanochrus Tschitschérine, 1901, Poecilus [Pseudoderus] 484

melanocornis Dejean, 1826, Chlaenius 350

melanocornis A. Zimmermann, 1915, Ilybius [Agabus] 42

melanodes Chaudoir, 1878, Pterostichus [Feronia] 495



melanogastrica Dejean, 1828, Amara 561

melanogrammus Régimbart, 1899, Nebrioporus [Hydroporus] 66



melanoleuca Dokhtouroff, 1885, Cicindela 106

melanophtalmos Geoffroy, 1785, Laccophilus [Dyticus] 77

melanophthalmus Šlosser‑Klekovski, 1877, Notiophilus 96

melanopoda Schmidt‑Göbel, 1846, Neocollyris [Collyris] 117

melanopterus Zetterstedt, 1828, Rhantus [Dyticus] 46

melanopus Andrewes, 1923, Callistoides [Chlaenius] 348

melanoscelis Marseul, 1880, Pterostichus [Feronia] 487

melanostoma Schenkling, 1889, Cicindela 105

melanotus Putzeys, 1873, Calathus 527

melanoxantha Kryzhanovskij, 1994, Klapperichella 459



melanoxanthus Gryuntal, 1984, Dyschirius 228

melantho Bates, 1883, Synuchus [Crepidactyla] 545

melanura Linné, 1767, Odacantha [Attelabus] 444

melas Aubé, 1837, Agabus 38

melas Oberthür, 1885, Carabus 139

melas Dejean, 1828, Molops 473

melas Kraatz, 1875, Molops 473

melas Sturm, 1818, Molops 473

melas Schmidt‑Göbel, 1846, Oxycentrus [Agreuter] 392



melas Gebler, 1848, Platynus [Anchomenus] 463

melas Creutzer, 1799, Pterostichus [Carabus] 495

meliki Csiki, 1912, Duvaliopsis [Anophthalmus] 298

melillensis Escalera, 1922, Laemostenus 539

melillensis Escalera, 1914, Singilis 438

melitensis Fairmaire, 1855, Laemostenus [Sphodrus] 539

melitops Acciavatti & Pearson, 1989, Cylindera [Cicindela] 111

melittus Bates, 1889, Platynus [Colpodes] 465

melleii Westwood, 1833, Platyrhopalopsis [Platyrhopalus] 211

melletii Heer, 1837, Agonum 451

melletii Heer, 1837, Ophonus [Harpalus] 390

melleus Lapouge, 1909, Carabus 172

melli Born, 1923, Carabus [Apotomopterus] 126

melli G. Ochs, 1925, Orectochilus 29

mellianus H. Kolbe, 1914, Carabus 142

mellyi Chaudoir, 1846, Carabus 175



mellyi Chaudoir, 1850, Chlaenius [Diaphoropsophus] 355

mellyi Régimbart, 1882, Dineutus [Dineutes] 26

mellyi Gebler, 1847, Nebria 89

mellyi Gebler, 1843, Pterostichus [Omaseus] 506



mellyi Gestro, 1893, Tricondyla 118

melyachi Shilenkov & Menstshikov, 1996, Carabus 170

memnonia Dejean, 1831, Pseudoclivina [Clivina] 221

memnonius Nicolai, 1822, Hydroporus 62



memnonius Nicolai, 1822, Platynus [Anchomenus] 463

memorabilis Deuve, 1992, Deltomerodes 281

memorabilis Ledoux & Roux, 1992, Nebria 94

menander Andrewes, 1935, Bembidion 256



mendacissimum J. R. Sahlberg, 1913, Bembidion 254

mendax Csiki, 1906, Carabus 173

mendax Zamotajlov, 1988, Carabus 132

mendax Apfelbeck, 1902, Molops 473



mendax Schauberger, 1926, Ophonus [Harpalus] 388

mendax P. Rossi, 1790, Parophonus [Carabus] 393



mendelssohni Schweiger, 1970, Carabus 198

mendizabali Mateu & Colas, 1954, Philorhizus 425

mendizabali Jeannel, 1921, Pterostichus [Troglorites] 514

menetriesi Falderman [in Hummel], 1827, Carabus 136

menetriesi Ballion, 1869, Lebia 429

menetriesii Faldermann, 1839, Agonum 450

menetriesii Kolenati, 1845, Bembidion [Peryphus] 253

meneziadum Morvan, 1996, Xestagonum 468



mengenensis Breuning, 1975, Cychrus 202

mengenicola Schweiger, 1969, Carabus 131

mengtzei Jedlika, 1931, Steropanus [Pterostichus] 517

menorquensis Coiffait, 1961, Microtyphlus 240

menozzii Schatzmayr, 1936, Corcyranillus [Scotodipnus] 238

mensioni Lagar Mascaró, 1976, Aphoenops 293

mentitus Vereschagina, 1989, Pseudotaphoxenus [Taphoxenus] 541

mentor Blumenthal & Breuning, 1967, Carabus 185

menviellei A. Gaudin, 1947, Geotrechus 310

mephistopheles Deuve, 1990, Carabus 189

mequignoni Colas, 1939, Bembidion 253



mera Jedlika, 1934, Amara 554

merangensis Mollard, 1999, Carabus 141


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