Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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mercatii Breuning & Ruspoli, 1970, Carabus 158

merdelensis Antoine, 1963, Cymindis 413

merdeniki Cavazzuti & Korell, 1992, Carabus 200

merditana Apfelbeck, 1906, Nebria 86

merditanus Apfelbeck, 1918, Aptinus 212

merditanus Apfelbeck, 1918, Carabus 165

merditanus Apfelbeck, 1906, Molops 473

merditanus Apfelbeck, 1906, Trechus 336

meregallii Casale, 1983, Trechus 336

merenicus Belousov & Kabak, 1994, Trechus 336

mergus Ahrens, 1812, Gyrinus 27

meridians Deuve, 1992, Carabus 190

meridianus Linné, 1761, Acupalpus [Carabus] 398

meridianus Csiki, 1927, Carabus 130

meridianus Fedorenko, 1992, Dyschirius 226

meridianus Andrewes, 1923, Harpalus 385

meridianus Schauberger, 1926, Ophonus [Harpalus] 390

meridianus Uéno, 1994, Trechiama 323

meridies Habu, 1975, Nipponagonum [Platynus] 461


meridionale Ganglbauer, 1891, Bembidion 249

meridionale Kolenati, 1845, Bembidion [Notaphus] 250

meridionalis Lassalle, 1999, Acalathus 529

meridionalis Machado, 1992, Aepus 288

meridionalis Putzeys, 1867, Amara [Leiocnemis] 555

meridionalis L. Gaudin, 1925, Aphoenops 294

meridionalis Puel, 1925, Badister 440

meridionalis Born, 1905, Carabus 171

meridionalis Lapouge, 1908, Carabus 130

meridionalis J. Jacquet, 1938, Clivina 219

meridionalis Gené, 1836, Cybister [Trochalus] 50

meridionalis Chaudoir, 1861, Cychrus 204

meridionalis Dejean, 1825, Cymindis 412

meridionalis Dejean, 1825, Dromius 420

meridionalis Puel, 1937, Dyschirius 225

meridionalis Abdul‑Karim & Ali, 1986, Hydrovatus 70

meridionalis Aubé, 1837, Ilybius 42

meridionalis Aubé, 1838, Metaporus [Hydroporus] 65

meridionalis Mlyná, 1977, Molops 474

meridionalis Winkler, 1933, Omphreus 445

meridionalis Ali, 1967, Oodes 445

meridionalis Dejean, 1829, Ophonus [Harpalus] 389

meridionalis Binaghi & Sanfilippo, 1971, Oreodytes 67



meridionalis J. Müller, 1911, Patrobus 285

meridionalis Dejean, 1828, Pogonus 286

meridionalis Cabidoche, 1967, Pterostichus [HN!] 498

meridionalis Dejean, 1828, Pterostichus [Feronia] 499

meridionalis Casale, 1982, Taphoxenus 544

meridionalis Casale, 1985, Typhloreicheia 223

merisioi Casale, Giachino, Vailati & Vigna Taglianti, 1996, Duvalius 304

merkensis Kabak, 1992, Carabus 146

merkli Régimbart, 1884, Agabus 39

merkli Apfelbeck, 1904, Amphimasoreus 521

merkli Kraatz, 1880, Carabus 172

merkli J. Frivaldszky, 1880, Licinus 442

merkli Pawowski, 1973, Trechus 336

merkliana Apfelbeck, 1904, Nebria 91



merklianus Géhin, 1885, Carabus 172

merklii Hopffgarten, 1878, Carabus 197

merklii J. Frivaldszky, 1877, Duvalius [Anophthalmus] 307

merklii J. Frivaldszky, 1879, Pterostichus [Feronia] 512

merlini Schaum, 1861, Carabus 138



mertolaensis Breuning, 1975, Carabus 167

merum Jedlika, 1933, Bembidion 21, 255



merus Tschitschérine, 1895, Harpalus 372

merzbacheri G. Hauser, 1922, Carabus 160

merzbacherianus Breuning, 1932, Carabus 123

mesasiaticum Mikhailov, 1988, Bembidion 258



mesasiaticus Fedorenko, 1989, Dyschirius 230

mesasiaticus Uéno, 1976, Perileptus 289



mesatlantica Antoine, 1953, Amara [Celia] 556

meschniggi Kult, 1948, Amara 551

meschniggi Netolitzky, 1943, Bembidion 246

meschniggi Winkler, 1936, Caecoparvus [Scotodipnus] 239

meschniggi Meixner, 1928, Duvalius 299

meschniggi Schatzmayr, 1928, Tapinopterus [Pterostichus] 520

meschniggi Jeannel, 1930, Trechus 336



mescitliensis Staven & Heinz, 1998, Carabus 201

mesembrinus Tschitschérine, 1896, Poecilus [Feronia] 480

mesepotamicus Abdul‑Karim & Ali, 1986, Colymbetes 45

mesites Fairmaire, 1897, Carabus 142

meskheticus Belousov, 1991, Pterostichus 496

mesoepisternalis W. Horn, 1933, Cylindera [Cicindela] 110



mesolcinus Born, 1906, Carabus [Orinocarabus] 180

mesopotamica Baliani, 1937, Amara 568

mesopotamicum Csiki, 1929, Cymbionotum [Graniger] 235

mesopotamicus Ali, 1967, Brachinus 216



mesopotamicus Mandl, 1979, Chlaenius 354

mesopotamicus J. Müller, 1922, Dyschirius 226

mesopotamicus Pic, 1901, Phloeozetus [Singilis] 438

mesostictum Bates, 1889, Agonum [Anchomenus] 455

messae Baliani, 1924, Amara 557

messai J. Müller, 1913, Orotrechus [Trechus] 317



messai J. Müller, 1913, Orotrechus [Trechus] 317

messii Bates, 1873, Pristomachaerus 356

mestscherjakovi Lutshnik, 1924, Carabus 171



mestus Cristoforis & Jan, 1837, Carabus 158

metalkanus Apfelbeck, 1919, Carabus 163

metallaicus Kleinfeld, 1999, Carabus 126



metallescens Dejean, 1828, Agonum 452

metallescens C. Zimmermann, 1831, Amara [Zabrus] [jun. HN?] 547



metallescens Dejean, 1831, Amara [jun. HN?] 547

metallescens Dejean, 1829, Amblystomus [Acupalpus] 360



metallescens Rye, 1874, Harpalus 378

metallescens Tschitschérine, 1897, Lioholus 404

metallescens Régimbart, 1907, Orectochilus 29

metallica Fairmaire, 1889, Odacantha [Casnonia] 444



metallica Peyron, 1858, Tachyura [Bembidium] 279

metallicum Sturm, 1826, Bembidion 248

metallicus Bedel, 1898, Anisodactylus 361

metallicus Dejean, 1828, Calathus 527

metallicus Kleinfeld, 1998, Carabus 126

metallicus Ahrens, 1830, Dyschirius [Clivina] 226

metallicus G. R. Waterhouse, 1833, Notiophilus 96

metallicus O. G. Costa, 1839, Notiophilus [Elaphrus] [HN] 97

metallicus Régimbart, 1884, Orectochilus 29



metallicus Scopoli, 1763, Poecilus [Carabus] [NO] 484

metallicus Fabricius, 1792, Pterostichus [Carabus] 488

metallifer Gistel, 1857, Pterostichus [Poecilus] [ND] 515

metallinus Ménétriés, 1836, Harpalus 379

metallorum Jeannel, 1955, Geotrechus 310

metarsius Andrewes, 1930, Harpalus 379

metax Jedika, 1962, Anchomenus [Agonum] 455

metax Jedlika, 1938, Pterostichus 486

metohiensis Apfelbeck, 1904, Neotrechus [Trechus] 316

meurguesae Ledoux, 1980, Carabus 155

meurguesae Ledoux, 1984, Carabus [Relictocarabus] 190

meurguesae Ledoux, 1972, Derostichus 441

meurguesae Ledoux, 1984, Ledouxius [Penetretus] 284

meurguesae Ledoux, 1985, Nebria 90

meurguesae Morvan, 1996, Platynus [Batenus] 463

meurguesae Casale & Ledoux, 1996, Pseudotaphoxenus 541



meurguesae Morvan, 1976, Trechus 329

meurguesianus Ledoux, 1990, Carabus 190

meurguesianus Morvan, 1996, Platynus [Batenus] 463

meurguesianus Deuve, 1980, Trechus 336

meurzwarelensis Morvan, 1996, Platynus [Batenus] 463

meuseli Reitter, 1910, Trechus 338

mewaensis Deuve, 1996, Carabus 191

mewaensis Deuve, 1997, Cychrus 206

meyeri De Monte, 1957, Bembidion 261



meyeri Born, 1898, Carabus 166

meyeri Jedlika, 1934, Platynus [Colpodes] 465

meyeri Jedlika, 1934, Pterostichus 515

meyeri Jedlika, 1935, Stenolophus [Egadroma] 405



meyerianus Born, 1898, Carabus [Coptolabrus] 144

meziadis Csiki, 1913, Duvalius [Anophthalmus] 300

miandamensis Heinertz, 1979, Carabus 162

mianshanicus Cavazzuti, 1998, Carabus 128

mianyangensis Deuve & Li, 1998, Carabus 183

miaorum Lassalle & Prunier, 1993, Carabus 156

micans Nicolai, 1822, Agonum [Anchomenus] 453

micans Tschitschérine, 1894, Amara 564

micans Fabricius, 1792, Chlaenius [Carabus] 348



micans Gautier des Cottes, 1863, Dyschirius 227

micans Gozis, 1873, Lebia 427

micans Ledoux & Roux, 1998, Nebria 92

micans Wollaston, 1863, Nesacinopus [Cratognathus] 388

micans Chaudoir, 1842, Pogonus 287

micans Heer, 1838, Pterostichus 512

micans Schaum, 1860, Trechus 341

micantula Hieke, 1994, Amara 564



michaelae Darge, 1994, Carabus 193

michaelensis Vidal y López, 1916, Cylindera [Cicindela] 109

michaeli Fourès, 1954, Aphoenops 295

michaeli Pawowski, 1978, Trechus 336

michaili Kataev, 1990, Harpalus 379

michailovi Müller‑Motzfeld, 1986, Bembidion 256

michailovi Kabak, 1992, Carabus 171

michailovi Emetz, 1972, Cymindis 416

michailovi Kataev, 1987, Harpalus 379

michailovi Komarov & Kabak, 1995, Philorhizus 425

michailovi Wrase, 1992, Pterostichus 511

michajlovi Vereschagina, 1988, Pseudotaphoxenus [Taphoxenus] 541

michinoku Nakane, 1968, Pterostichus 494

micklitzi Ganglbauer, 1913, Anophthalmus [Trechus] 292



micklitzi Sokolá, 1910, Carabus 181

micrangulus Jeannel, 1930, Duvalius 299

micrangulus Reitter, 1913, Trechus 336

micraulax Andrewes, 1924, Tachyura [Tachys] 277

microcephala Motschulsky, 1844, Amara [Celia] 565

microcephala K. Daniel & D. Daniel, 1891, Oreonebria [Nebria] 96



microcephalus Stephens, 1835, Badister 440

microcephalus Dejean, 1828, Calathus 528

microcephalus Gistel, 1857, Carabus 152

microcephalus Rambur, 1837, Carterus 364

microcephalus Coiffait, 1958, Geotrechus 310

microcephalus Ledoux, 1984, Ledouxius [Penetretus] 284

microcephalus Faldermann, 1835, Neophygas [Acinopus] 387

microcephalus Motschulsky, 1860, Pterostichus [Argutor] 513



microchondrus Fischer von Waldheim, 1842, Carabus 172

microcratophyrtus Kaláb, 1996, Carabus 146

microdemas Schauberger, 1932, Harpalus 386

microdera Chaudoir, 1844, Amara [Bradytus] 552

microderus Chaudoir, 1867, Carabus 198

microderus Solsky, 1874, Dicheirotrichus 402

microgonus Bates, 1886, Loxoncus [Anoplogenius] 404

microgranum Cavazzuti & Ratti, 1999, Carabus 156



micromentum Deuve & Imura, 1993, Carabus 150

microphthalma Baliani, 1943, Amara 558

microphthalma Ledoux & Roux, 1998, Nebria 92

microphthalmum Chaudoir, 1876, Cymbionotum 235

microphthalmum Putzeys, 1874, Zuphium 574

microphthalmus Reitter, 1919, Brachinus 213

microphthalmus L. Miller, 1859, Duvalius [Trechus] 307

microphthalmus Delarouzée, 1857, Pterostichus 498

microphthalmus Heyden, 1870, Reicheiodes [Dyschirius] 230

microphthalmus Fairmaire, 1888, Robustopenetretus [Patrobus] 286

microphthalmus Uéno, 1955, Trechoblemus 324

microphthalmus Uéno, 1955, Trephionus 547

microporus Ledoux & Roux, 1998, Archastes 79



microps Bedel, 1893, Apotomus 237

microps Uéno & Yin, 1993, Junnanotrechus 313

microps Zamotajlov & Sciaky, 1996, Parapenetretus 284

microps Andrewes, 1935, Perileptus 289

microps Heyden, 1887, Pterostichus 501

micropterus Duftschmid, 1812, Calathus [Carabus] 527

microputus O. Berlov & Shilenkov, 1996, Carabus 174

micros Fischer von Waldheim, 1828, Dromius 421

micros Fischer von Waldheim, 1828, Tachys [Dromius] 21, 276

micros Herbst, 1784, Trechoblemus [Carabus] 324

microscopicus Bates, 1873, Polyderis [Tachyta] 274

microsticta Klug, 1862, Myriochila [Cicindela] 114

Microtachys Casey, 1918, Polyderis 274

microtatos Cavazzuti, 1997, Carabus 126

microtes Habu, 1978, Synuchus 545

microthorax Deuve, 1992, Cychrus 203



microthorax Semenov, 1900, Deronectes 57

microthorax Motschulsky, 1844, Dyschirius 226

microthorax Motschulsky, 1849, Harpalus [Anisodactylus] 379

microthorax Motschulsky, 1844, Nebria 89

microthorax Fischer von Waldheim, 1829, Platynus 464

microthorax Coiffait, 1952, Trechus 328

microtis R. T. Bell & J. R. Bell, 1982, Omoglymmius 78

midas Reitter, 1896, Carabus 148

midas Jeannel, 1927, Trechus 336

middendorffi J. R. Sahlberg, 1875, Pterostichus [Feronia] 491

middendorfi Ménétriés, 1851, Carabus 170

migliaccioi Casale, 1982, Laemostenus [Pristonychus] 538

migliaccioi Straneo, 1982, Pterostichus 515

migrans Cavazzuti & Ratti, 1999, Carabus 127

mihatschi Reitter, 1900, Harpalus 381

mihoki Csiki, 1912, Duvalius [Anophthalmus] 300

mihoki Jedlika, 1953, Pseudotaphoxenus [Taphoxenus] 541



mikado Bates, 1873, Calosoma 119

mikae Lafer, 1980, Amara 554

mikhaili Deuve & Mourzine, 1997, Carabus 154

mikhailovianus Deuve & Dolin, 1993, Carabus 153

mikii Habu, 1966, Trephionus 547

mikitjukovi Mikhailov, 1997, Bembidion 258

mikurana Nakane, 1968, Cylindera [Cicindela] 110

milalaensis Hieke, 1997, Amara 560

milenae Casale, 1983, Duvalius 304

miles Semenov, 1887, Carabus 134

miles Tschitschérine, 1889, Pterostichus 497

miliaris Fabricius, 1801, Cymindis [Carabus] 415

militaris Sciaky & Wrase, 1997, Aristochroa 471

militaris Tschitschérine, 1893, Pterostichus [Feronia] 497



millavensis Schaefer, 1969, Carabus [Autocarabus] 195

milleri Jacquelin du Val, 1852, Bembidion 259



milleri C. G. Thomson, 1875, Carabus 181

milleri J. Frivaldszky, 1862, Duvalius [Anophthalmus] 299

milleri Schaufuss, 1862, Pseudotaphoxenus [Sphodrus] 542

milleri L. Redtenbacher, 1858, Trechus 337

millerianum Heyden, 1883, Bembidion 268

milleti Antoine, 1932, Chlaenius 355

millingeni Bates, 1878, Hypaetha [Cicindela] 112

millingeni J. Balfour‑Browne, 1951, Nebrioporus [Potamonectes] 66



milyaevi O. Berlov, 1997, Cychrus 19, 202

mimethes Kraatz, 1878, Carabus 180

mimica Casey, 1918, Amara [Celia] 565

mimicus G. Ochs, 1936, Orectochilus 29



mimmi J. R. Sahlberg, 1875, Ilybius [Gaurodytes] 42

mimus H. Kolbe, 1913, Carabus 196

mimus Uéno, 1965, Ryugadous 319

minaicus Belousov & Kabak, 1994, Trechus 336

minamikawai Habu, 1959, Nipponagonum [Agonum] 461

minax Andrewes, 1935, Cymbionotum 235

minax Tschitschérine, 1900, Tapinopterus [Platysma] 518

minervae Coiffait, 1956, Anillus 238

ming Cavazzuti, 1995, Carabus 143

ming Sciaky & Wrase, 1997, Pterostichus 500



mingazzinii Magrini & Vanni, 1985, Duvalius 301

mingazzinii Magrini & Vanni, 1990, Typhloreicheia 223

mingens Quensel, 1806, Carabus 182

mingrel Novotný & Voíšek, 1988, Carabus 200

mingrelicum Belousov & Sokolov, 1994, Bembidion 246

mingrelicus Reitter, 1889, Carabus [Tribax] 200

mingrelicus Reitter, 1885, Trechus 328

mingrelicus Belousov, 1998, Troglocimmerites 344



miniangularis J. Schmidt, 1998, Deltomerodes 281

miniatulus Venet, 1930, Carabus 140

miniatus Schaufuss, 1865, Laemostenus [Pristonychus] 539

minima Jedlika, 1931, Diplocheila 439



minima Duftschmid, 1812, Tachyta [Elaphrus] 277

minima Binaghi, 1936, Typhloreicheia 223

minimum Fabricius, 1792, Bembidion [Carabus] 249

minimus Semenov & Znojko, 1932, Carabus 192

minimus Zaitzev, 1908, Colymbetes 45

minimus Ahrens, 1830, Dyschirius [Clivina] 229

minimus Motschulsky, 1845, Ophonus 389

minimus Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 541

minimus Csiki, 1930, Pterostichus 489

minimus Cerruti, 1977, Speluncarius 516

minimus Lindroth, 1956, Synuchus 545

minimus Curtis, 1828, Tachys [Cillenum] 275

minimus Vuillefroy, 1866, Trymosternus [Cymindis] 410

minioculatus Machado, 1987, Trechus 336

minkwitzii Dejean, 1828, Pterostichus 487

minobusanus Habu, 1978, Yukihikous 469

minoensis Ishikawa & Kubota, 1994, Carabus 177

minor G. Quedenfeldt, 1883, Acinopus 368

minor J. R. Sahlberg, 1875, Agabus 37

minor Jedlika, 1936, Agonum [Colpodes] 452

minor Coiffait, 1962, Aphoenops 294

minor Jedlika, 1958, Brachinus 213

minor Bates, 1889, Carabus 125

minor Escalera, 1944, Carabus 137

minor Gautier des Cottes, 1866, Carabus 186

minor Haury, 1881, Carabus 138

minor Kraatz, 1878, Carabus [Orinocarabus] 180

minor Kwon & Lee, 1984, Carabus [Leptocarabus] 159

minor Pirazzoli, 1872, Carabus 130

minor Nietner, 1856, Distichus [Scarites] 231

minor Reitter, 1884, Harpalus [Pseudophonus] 380

minor A. Costa, 1847, Hygrotus [Hydroporus] 73

minor Tschitschérine, 1900, Idiomelas 404

minor Ménétriés, 1848, Lebia 428

minor Reiche & Saulcy, 1855, Olisthopus 461

minor Tschitschérine, 1894, Oxylobus 231

minor Hrka & Pulpán, 1981, Parazuphium 574

minor Shiraki, 1907, Paussus 210

minor Gyllenhal, 1827, Pterostichus [Harpalus] [NP] 511

minor N. Ito, 1998, Trichotichnus 394

minoricensis J. Vives & E. Vives, 1994, Scybalicus 363



minorita Jedlika, 1957, Amara 547

minorita Gistel, 1857, Harpalus 386

minos Linder, 1860, Hydraphaenops [Anophthalmus] 311

minos Schauberger, 1934, Tschitscherinellus 367

minoum Huber & Marggi, 1997, Bembidion 264

minowai Habu, 1947, Scarites 234

minozzii Dodero, 1917, Duvalius [Trechus] 304

minshanensis Deuve, 1987, Carabus 154

minshanica Hieke, 1997, Amara 554

minshanicola Deuve, 1987, Cychrus 205

minshanicus Deuve, 1992, Cychrus 204

minshanus Jedlika, 1965, Pterostichus 513

minuens Casey, 1918, Patrobus 285

minusculus Semenov, 1903, Carabus 147

minusculus G. Ochs, 1936, Orectochilus 29



minusculus Poppius, 1906, Pterostichus [Cryobius] 490

minusculus Straneo, 1989, Speluncarius 516

minuta Motschulsky, 1844, Amara [Bradytus] 567

minuta Olivier, 1790, Cylindera [Cicindela] 111

minuta Kabak & Wrase, 1997, Cymindis 416

minutissimus Germar, 1824, Bidessus [Hydroporus] 55

minutissimus J. Balfour‑Browne, 1939, Copelatus 48

minutissimus Motschulsky, 1850, Polyderis [Tachys] 274

minutissimus Péringuey, 1908, Polyderis [Tachys] 274

minutissimus Stephens, 1828, Tachys 275

minutulus Goeze, 1777, Microlestes [Carabus] 423

minutulus J. Müller, 1937, Speluncarius 516

minutum Motschulsky, 1844, Agonum 453

minutum Audinet‑Serville, 1821, Bembidion 249

minutus Brullé, 1832, Acinopus 368

minutus Motschulsky, 1844, Acupalpus [Stenolophus] 398

minutus Gautier des Cottes, 1866, Calathus 525

minutus Motschulsky, 1850, Carabus 199

minutus Dejean, 1825, Dyschirius [Clivina] 226

minutus Dejean, 1822, Graphipterus 359

minutus Fabricius, 1798, Gyrinus 27

minutus Takizawa, 1931, Haliplus 31


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