Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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minutus Dejean, 1829, Harpalus 379

minutus Stephens, 1828, Hydroporus 65

minutus Linné, 1758, Laccophilus [Dytiscus] 76



minutus Geoffroy, 1785, Microlestes [Buprestis] 423

minutus Motschulsky, 1844, Microlestes [Dromius] 423

minutus Reiche, 1871, Orthomus 477

minutus Reiche & Saulcy, 1855, Platyderus [Feronia] 523



minutus Stichel, 1923, Pseudaristus 367

minutus Motschulsky, 1866, Pterostichus [Argutor] 507

minutus Uéno, 1997, Stevensius 320

minutus Uéno, 1971, Trechiama 323

minutus Fabricius, 1792, Trechus [Carabus] 339

minyops Wollaston, 1862, Trechus 336



miolansicus Tarrier, 1965, Carabus [Archicarabus] 130

mirabilis Casale, 1983, Aparupa 456

mirabilis Bates, 1892, Catascopus 435

mirabilis Zamotajlov, 1992, Deltomerus 283

mirabilis Jedlika, 1932, Loricera 98

mirabilis Jedlika, 1960, Marschneria 406

mirabilis Kurnakov, 1959, Meganophthalmus 314

mirabilis Ledoux & Roux, 1990, Nebria 86

mirabilis Belousov, 1998, Porocimmerites 318

mirabilis Jedlika, 1939, Pterostichus 498

mirabilis Uéno, 1958, Rakantrechus 319

mirabilis L. Miller, 1863, Spelaeodytes 223



mirabilis Bates, 1892, Tachyura [Tachys] 278

mirabilissimus Ishikawa & Deuve, 1982, Carabus [Damaster] 122

mirabilissimus Uéno & Wang, 1991, Sinaphaenops 320

miranda Dupuis, 1913, Tachyura [Tachys] 278

mirandior Deuve, 1993, Carabus 143

mirandus Jedlika, 1940, Colposphodrus [Colpodes] 457

mirandus Bates, 1873, Dischissus 447

mirei Nègre, 1966, Calathus 525

mirei Breuning, 1975, Carabus 161

mirei Rivalier, 1962, Myriochila [Myriochile] 113

miridita Apfelbeck, 1904, Tapinopterus [Pterostichus] 520

miriditus Apfelbeck, 1901, Carabus 136

mirifica Jedlika, 1956, Lebia 429

mirificum Casale, 1979, Calosoma [Callisthenes] 119

mirificus Kraatz, 1894, Carabus [Coptolabrus] 143

mirificus Jedlika, 1940, Platynus [Colpodes] 465

mirificus Bates, 1883, Pterostichus 498

miroshnikovi Zamotajlov, 1990, Carabus 132

miroshnikovi Zamotajlov, 1999, Deltomerodes 281

miroshnikovi Zamotajlov, 1994, Deltomerus 283

miroshnikovi Belousov & Zamotajlov, 1995, Duvalius 304

miroshnikovi Zamotajlov, 1991, Pterostichus [Aphaonus] 485

miroslavae Farka & Sciaky, 1998, Leistus 80

miroslavi Sciaky & Wrase, 1997, Pterostichus 500

mirus Schauberger, 1934, Proditomus 367

mirus Tschitschérine, 1894, Pterostichus [Feronia] 499

mirzayani Morvan, 1973, Bembidion [Peryphus] 257

mirzayani Morvan, 1973, Laemostenus [Antisphodrus] 534


mirzayani Morvan, 1974, Laemostenus [Pristonychus] 538

mirzayani Morvan, 1977, Psammodromius 426

mirzayani Morvan, 1974, Trechus 336

misella L. Miller, 1868, Amara 557

misellum Harold, 1877, Bembidion 271

misellus Lapouge, 1898, Carabus 195

misellus Schaum, 1857, Dyschirius 226

misellus Tschitschérine, 1897, Harpalus 374

miser Tschitschérine, 1897, Trichotichnus [Harpalus] 396

misera Tschitschérine, 1894, Amara 560

mishmicus Arrow, 1942, Rhyzodiastes [Clinidium] 78



mistellus Marseul, 1882, Hygrotus [Hydroporus] 72

mithrae Zaitzev, 1908, Gyrinus 27



mitis Andrewes, 1930, Amara 553

mitjaevi Belousov & Kabak, 1996, Trechus 336



mitodes Andrewes, 1929, Dyschirius 224

mitoyamanus Tanaka, 1971, Pterostichus 502

mitridati Pliginskiy, 1915, Harpalus 379

mitsumasai Imura, 1994, Carabus 164



mittrei Lucas, 1846, Carabus 161

miurai Uéno, 1957, Morimotoa 75



miwai Jedlika, 1946, Amara [Harpalus] 562

miwai Jedlika, 1946, Bembidion 271



miwai Kurosawa, 1959, Carabus 143

miwai Jedlika, 1946, Chydaeus 362

miwai Jedlika, 1940, Dromius [Lebidromius] 421

miwai Jedlika, 1951, Lebia 429

miwai Jedlika, 1940, Peliocypas [Risophilus] 434

miwai Jedlika, 1940, Platynus [Colpodes] 465

miwai Jedlika, 1940, Pristosia [Kanoldia] 531

miwai Jedlika, 1949, Trichotichnus 396

mixanigi Daffner, 1985, Anophthalmus 292

mixanigi Daffner, 1993, Duvalius 304

mixta Baliani, 1943, Amara 560

mixta Jedlika, 1957, Amara 547

mixta Ferrer y Dalmau, 1911, Cicindela 106

mixtum Schaum, 1863, Bembidion 258



mixtus Guignot, 1949, Agabus 40

mixtus Géhin, 1878, Carabus 166



mixtus Motschulsky, 1860, Hygrotus [Hydrocoptus] 72

mixtus Jedlika, 1940, Platynus [Colpodes] 465

mixtus Blanchard, 1843, Sandracottus [Hydaticus] 49

mixtus Herbst, 1784, Stenolophus [Carabus] 406

miyakei Habu, 1967, Apristus 431

miyakei Nakane, 1961, Carabus 159


miyakei Ohkura & Gotô, 1960, Carabus 153

miyakei Habu, 1974, Euplynes 458

miyakei Habu, 1978, Oxycentrus 392

miyakei Habu, 1980, Trichotichnus 395



miyakejimanum Nakane, 1961, Callytron [Cicindela] 101

miyamai Kasahara & Y. Itô, 1987, Pterostichus 514

miyamotoi Nakane, 1990, Leiodytes [Clypeodytes] 57

miyamotoi Nakane, 1959, Platambus [Gaurodytes] 44

miyamotoi Habu, 1943, Stomis 517

miyoshiorum Uéno, 1969, Stygiotrechus 320

mizunumai Ishikawa, 1972, Carabus 149

mizunumaianus Imura, 1991, Carabus 197



mizushimai Uéno, 1969, Ryugadous 319

mjobergi A. Zimmermann, 1927, Laccophilus 77

mladenovi V. B. Guéorguiev, 1964, Pheggomisetes 318

mlynari Kataev, 1990, Harpalus 379

mniszechi Piochard de la Brûlerie, 1873, Acinopus 368

mniszechi Chaudoir, 1852, Carabus 147



mniszechi W. Horn, 1891, Cephalota [Cicindela] 102

mniszechii Chaudoir, 1854, Nebria 91

moana Morvan, 1995, Skoeda 466

moanaatum Morvan, 1996, Xestagonum 469

moanaet Morvan, 1998, Tostkar 467

mochizukii Imura & Mizusawa, 1994, Carabus 177

mocsaryi Csiki, 1913, Duvalius [Anophthalmus] 300

moczarskii J. Müller, 1917, Duvalius [Trechus] 304

moczarskii Jeannel, 1937, Prioniomus 238

modeeri Marsham, 1802, Orectochilus [Gyrinus] 29

modesta Dejean, 1828, Amara 557

modestior Maindron, 1906, Carabus 128

modestior Deuve, 1991, Cychrus 204

modestium Bates, 1873, Agonum [Colpodes] 454

modestoides Casale, 1988, Laemostenus 536

modestulus Semenov, 1887, Carabus 191

modestum Sturm, 1824, Agonum 452

modestum Fabricius, 1801, Bembidion [Carabus] 255

modestus Schaum, 1863, Callistomimus [Callistus] 348

modestus Fischer von Waldheim, 1823, Carabus 199

modestus Kraatz, 1876, Carabus 185

modestus Oberthür, 1883, Carabus 128

modestus A. Zimmermann, 1924, Haliplus 31

modestus Dejean, 1829, Harpalus 379



modestus Sharp, 1882, Hydaticus 54

modestus Casale, 1981, Laemostenus 536

modestus Schaufuss, 1862, Laemostenus [Sphodrus] 534



modestus Reiche, 1871, Orthomus 476

modestus Schaum, 1858, Pseudaristus [Ditomus] 367

modestus Putzeys, 1874, Trechus 336

modestus Tschitschérine, 1906, Trichotichnus [Amaroschesis] 394

modicellus Tschitschérine, 1897, Pterostichus [Feronia] 513

modicus Coquerel, 1859, Orthomus [Feronia] 477

modron Morvan, 1995, Xestagonum [Colpodes] 468



moerea Jedlika, 1957, Amara 561

moerens Putzeys, 1873, Clivina 220



moesiacus Jeannel, 1921, Trechus 339

moesta Schmidt‑Göbel, 1846, Neocollyris [Collyris] 117



moestum Duftschmid, 1812, Agonum [Carabus] 450

moestus Curtis, 1835, Agabus [Colymbetes] 36



moestus Dejean, 1831, Carabus 196

moestus Fairmaire, 1858, Deronectes [Hydroporus] 58



moestus Walker, 1871, Nebrioporus [Hydroporus] 67

moestus Rey, 1887, Pterostichus [Feronia] 512

moestus Küster, 1847, Zabrus [Eutroctes] 569

mogadoricus Escalera, 1914, Carabus 152

mogadoricus Escalera, 1922, Laemostenus 536

mohrii Uéno, 1996, Phreatodytes 35

mohrii Uéno, 1982, Yamautidius 345

molchanovi Belousov, 1999, Caucasaphaenops 297

moldaviensis Born, 1903, Carabus 170

moldoensis Vereschagina & Kabak, 1997, Laemostenus 537



molestum Motschulsky, 1844, Agonum 450

mollicellus C. A. Dohrn, 1880, Carabidomemnus [Paussus] 209



mollii Dejean, 1826, Carabus 179

mollis Marsham, 1802, Calathus [Carabus] 528



mollis Lapouge, 1925, Carabus 130

mollis Strøm, 1768, Cymindis [Carabus] 418

mollis Motschulsky, 1866, Nebria 88

moloch Piochard de la Brûlerie, 1873, Dixus [Aristus] 24, 365

molopiformis Kryzhanovskij, 1964, Amara 558

molopiformis Lutshnik, 1922, Tapinopterus [Platysma] 518

molopinus Chaudoir, 1868, Tapinopterus [Feronia] 518

molopsoides Jedlika, 1934, Pterostichus 501

molossulus Deuve, 1993, Carabus 145

mona Jedlika, 1963, Coptodera 436

monachum Duftschmid, 1812, Agonum [Carabus] 450

monalisa W. Horn, 1927, Callytron [Cicindela] 101

monasteriensis Westhoff, 1881, Cicindela 105

monastica Zaballos & Wrase, 1998, Typhlocharis 241

monastirensis Reitter, 1913, Tapinopterus 520

monastiriensis Lapouge, 1922, Carabus 187


monaulacus Drapiez, 1820, Hydroglyphus [Hydroporus] 56

moncayensis Jeanne, 1985, Platyderus 523

mondaensis Deuve, 1998, Trechus 330

mondalaensis Hieke, 1997, Amara 553



mondanus Jedlika, 1958, Pterostichus 508

monfalconensis J. Müller, 1926, Typhlotrechus 345

mongolensis Mandl, 1981, Calomera [Cicindela] 100



mongolica Jedlika, 1966, Amara [Harpalus] [HN] 556

mongolica Motschulsky, 1844, Amara [NO] 548

mongolica Faldermann, 1835, Cylindera [Cicindela] 111



mongolica Jedlika, 1966, Cymindis 415

mongolica Motschulsky, 1850, Lionedya [Homalops] 430



mongolica Jedlika, 1969, Nebria 89

mongolicum Shilenkov, 1993, Agonum 451



mongolicum Jedlika, 1967, Bembidion 255

mongolicum Lapouge, 1930, Calosoma [Callistrata] 120

mongolicum Semenov, 1922, Omophron 208

mongolicus V. B. Guéorguiev, 1968, Agabus 37

mongolicus Lapouge, 1905, Carabus 154

mongolicus Kult, 1949, Dyschirius 228

mongolicus A. I. Jakovlev, 1899, Hygrotus [Coelambus] 72

mongolicus Reitter, 1900, Licinus 442

mongolicus Brinck, 1943, Oreodytes [Deronectes] 68

mongolicus Jedlika, 1953, Pseudotaphoxenus [Taphoxenus] 541

mongolicus Motschulsky, 1844, Pterostichus [Poecilus] 506

mongolicus Motschulsky, 1844, Syntomus [Dromius] 433

mongolicus P. Moravec, 1992, Trechus 336

mongoliensis Jedlika, 1962, Poecilus [Pterostichus] 483



mongoliensis Jedlika, 1967, Pterostichus 509

mongolorum Csiki, 1927, Carabus 170

mongolorum Belousov & Kabak, 1994, Trechus 336

monguzzii Bucciarelli, 1978, Boldoriella 296

monguzzii Casale, 1988, Laemostenus 534

monilicornis J. R. Sahlberg, 1875, Hydroporus 63

monilicornis Habu, 1979, Nepalorthogonius 446

monilifer Tatum, 1847, Carabus 144

monilis Fabricius, 1792, Carabus 171

monizii Wollaston, 1860, Zargus 440

monochroa Chaudoir, 1850, Cymindis 417

monogramma Chaudoir, 1859, Orthomus [Feronia] 477

monostigma Jedlika, 1957, Amara 559

monostigma J. Müller, 1918, Bembidion 260

monostigma Samouelle, 1819, Demetrias 419

monostigma Andrewes, 1923, Lebia 429

monostigma Bates, 1873, Parena [Crossoglossa] 434



monostigma Tschitschérine, 1898, Pterostichus [Feronia] 508

monostriatus Geoffroy, 1785, Bidessus [Dyticus] 55

mons Habu, 1975, Bradycellus 402

monspessulani Schatzmayr, 1942, Poecilus [Pterostichus] 482

monspessulantus Lapouge, 1898, Carabus 195

monstruosus Stark, 1890, Carabus 133

montadai Mateu, 1950, Pterostichus [Haptoderus] 489



montalii Luigioni, 1920, Cylindera [Cicindela] 111

montana Chaudoir, 1837, Amara [Leirus] 563

montana Dejean, 1828, Amara 557



montana Charpentier, 1825, Cicindela 108

montana Sirguey, 1926, Cicindela 106

montandoni Born, 1902, Carabus 187

montanellus Heinz, 2000, Calathus 528



montanellus Heinz, 1967, Carabus 130

montanellus Mateu, 1953, Harpalus 381

montanellus Graëlls, 1851, Platyderus 523

montanellus Gemminger & Harold, 1868, Trechus 336

montaneus Werner, 1992, Therates 116

montanheirorum Oromí & Borges, 1991, Trechus 336



montanum Heer, 1837, Agonum 451

montanum Rambur, 1838, Bembidion 269



montanus Heinz, 1970, Calathus 528

montanus Baba, 1939, Carabus 148

montanus G. Hauser, 1913, Carabus [Coptolabrus] 143

montanus Géhin, 1882, Carabus 139

montanus Lucas, 1859, Chlaenius 355

montanus Winkler, 1939, Cychrus 205



montanus Helliesen, 1890, Hydroporus 62

montanus Stephens, 1828, Ilybius [Colymbetes] 42

montanus Stephens, 1827, Leistus 83

montanus Heer, 1837, Molops 474

montanus Absolon & Maan, 1947, Neotrechus 315



montanus A. Zimmermann, 1915, Oreodytes [Hydroporus] 67

montanus Motschulsky, 1844, Pterostichus [Steropus] 506

montanus Bedel, 1913, Syntomus [Metabletus] 433

montanus Lindroth, 1956, Synuchus 545



montanus Casale, 1988, Taphoxenus [Pseudotaphoxenus] 544

montanus Uéno, 1958, Trechiama 324

montanus Motschulsky, 1844, Trechus 336

montanus Putzeys, 1847, Trechus 336

montanus Sturm, 1818, Trichotichnus [Harpalus] 396

montarerae Bernau, 1914, Carabus 180

montei Fassati, 1959, Bembidion 269

montei Kult, 1959, Clivina 220


montellensis Agazzi, 1956, Orotrechus 317

montemurroi Vanni & Magrini, 1986, Duvalius 303

montenegrina Apfelbeck, 1904, Nebria 85

montenegrinus Kraatz, 1876, Carabus 138

montenegrinus Apfelbeck, 1904, Cychrus 205



montenegrinus Schaufuss, 1882, Graptodytes [Hydroporus] [NO] [add, syn of veterator] 60

montenegrinus L. Miller, 1866, Molops [Pterostichus] 475



montenegrinus Küster, 1847, Syntomus [Dromius] 432

montesamanus Mandl, 1967, Carabus 188

montichares W. Hansen & Gaskowiak, 1973, Carabus [Morphocarabus] 171

monticola Dejean, 1831, Amara 565

monticola Sturm, 1825, Bembidion 260



monticola Motschulsky, 1864, Brachinus 214

monticola Habu, 1973, Broscosoma 235

monticola Dejean, 1826, Carabus 129

monticola Ménétriés, 1832, Cicindela 106

monticola Chevrolat, 1866, Cymindis 412

monticola Casey, 1914, Dicheirotrichus [Trichocellus] 403

monticola Sharp, 1869, Hydroporus 62

monticola Dejean, 1829, Ophonus [Harpalus] 392

monticola Shibata, 1987, Parena 434



monticola Guignot, 1957, Pseuduvarus [Uvarus] 57

monticola Crotch, 1871, Pterostichus [Feronia] [HN] 493

monticola Gebler, 1848, Pterostichus [Platysma] 506

monticola Nicolai, 1822, Pterostichus 485

monticola Kasahara, 1994, Trichotichnus 396

monticola Holdhaus, 1924, Typhloreicheia [Reicheia] 223

monticoloides Shilenkov, 1995, Pterostichus 506



monticusta Iablokoff‑Khnzorian, 1957, Amara 557

montigena Tschitschérine, 1898, Pterostichus [Feronia] 501

montigradus G. Hauser, 1920, Carabus [Coptolabrus] 143

montis Jedlika, 1965, Skoeda [Agonum] 466

montisabanti Cavazzuti, 1989, Carabus 185

montisageli Jeannel, 1947, Duvalius 304

montisarerae Focarile, 1950, Trechus 336

montisauthionis Jeannel, 1941, Duvalius 302

montisblidarii Jeannel, 1928, Duvalius 300

montiscanini Jeannel, 1927, Trechus 335

montiscetii Jeannel, 1928, Duvalius 299

montiscrystalli Casale, 1979, Trechus 336

montiscusnae Focarile, 1952, Trechus 336

montisdeorum Schweiger, 1967, Pterostichus 511



montisgrappae Agazzi, 1960, Duvalius 302

montismaiellettae Ghidini, 1932, Trechus 341

montismatajuris Colla, 1986, Anophthalmus 292


montisneronis Binaghi, 1942, Typhloreicheia 223

montisoetae Casale, 1987, Duvalius 304

montispennae Jeannel, 1927, Trechus 336

montisragognae E. Porro & G. Porro, 1971, Orotrechus 317

montisrosae Jeannel, 1921, Trechus 336

montissalini A. Gaudin, 1947, Geotrechus 310

montissiruolis Jeannel, 1941, Duvalius 302

montistartari Jeannel, 1928, Duvalius 300

montiszedae Binaghi, 1945, Trechus 340

montivaga Sturm, 1825, Amara 550

montivagus Dejean, 1831, Calathus 528

montivagus Palliardi, 1825, Carabus 129



montivagus Reitter, 1900, Harpalus 378

montivagus Ménétriés, 1832, Pterostichus [Feronia] 501

montivagus Poppius, 1906, Pterostichus 506

montrouzieri Lucas, 1860, Rhantus [Colymbetes] 47

montsenicus Jeanne, 1985, Pterostichus 512



montserratensis Xaxars, 1915, Abax 470

montuosella J. Schmidt, 1998, Xestagonum 468

montuosum Habu, 1973, Xestagonum [Agonum] 468

moosbruggeri Born, 1913, Carabus 179

mopsucrenae Peyron, 1858, Carabus 187

moralesi Nègre, 1966, Calathus 525

moralesi Mateu, 1979, Cymindis 414

morawitzi Csiki, 1928, Bembidion 262

morawitzi Ganglbauer, 1887, Carabus 188



morawitzi Kraatz, 1887, Carabus 173

morawitzi Géhin, 1885, Cychrus 205



morawitzi Jedlika, 1953, Pseudotaphoxenus [Taphoxenus] 540

morawitzianus Semenov, 1887, Carabus 190

morawitzianus Lutshnik, 1922, Pterostichus [Platysma] 508

morbillosus Fabricius, 1792, Carabus 161

mordkovitschi Shilenkov, 1982, Trechus 336

moreana Apfelbeck, 1904, Cymindis 412

moreanus Reitter, 1896, Carabus 198

moreanus J. Müller, 1932, Licinus 443



moreaui Morvan, 1976, Trechus 329

moreli Lapouge, 1908, Carabus 130

moreoticus Maan, 1937, Tapinopterus 520

morettoi Deuve, 1997, Carabus 147

morgani Lapouge, 1907, Carabus 164

morholt Morvan, 1996, Xestagonum 468

moricei Pic, 1904, Bembidion 264

morii Ashida, 1999, Trechiama 323

morimotoi Habu, 1954, Diacanthostylus [Agonum] 458


morimotoi Uéno, 1984, Epaphiopsis 309

morimotoi Uéno, 1996, Morimotoa 75

morimotoi Uéno, 1955, Perileptus 289

morimotoi Uéno, 1973, Stygiotrechus 320



morimotoi Habu, 1955, Synuchus [Calathus] 545

morio Ménétriés, 1832, Amara 550

morio Mannerheim, 1830, Carabus 182

morio Putzeys, 1866, Dyschirius 226


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