Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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neboti Español, 1966, Ildobates 573

nebrioides Kryzhanovskij, 1974, Amara [Curtonotus] 560

nebrioides Antoine, 1961, Chlaenius 355

nebrioides Hoppe & Hornschuch, 1825, Licinus 442

nebrioides Vuillefroy, 1866, Penetretus [Patrobus] 285

nebulosum P. Rossi, 1792, Asaphidion [Elaphrus] 242

nebulosum Stephens, 1828, Bembidion [Notaphus] 254

nebulosus Forster, 1771, Agabus [Dytiscus] 40



nebulosus Schiødte, 1841, Agabus 39

nebulosus Chaudoir, 1876, Pheropsophus 217

necessarius Tschitschérine, 1894, Pterostichus [Feronia] 502

nechaevi Lafer & Kuznetsov, 1996, Pterostichus [add to subg. Cryobius] 493

necincertus Deuve, 1998, Carabus 174

necopinatus Basquin & Darge, 1988, Carabus 154

necopinatus Fery, 1999, Hydroporus 63

nedhalensis Ali, 1978, Cylindera [Cicindela] 111

neelgheriensis Guérin‑Méneville, 1840, Chlaenius 354

neerworti Reitter, 1893, Carabus 132


neesi Hoppe & Hornschuch, 1825, Carabus 166

negevensis Schweiger, 1970, Carabus 188



neglecta Landin, 1955, Amara 562

neglectoides Jedlika, 1960, Harpalus 375

neglectum Dawson, 1849, Bembidion [Peryphus] 250

neglectus Kraatz, 1887, Carabus 137

neglectus Lapouge, 1913, Carabus 170

neglectus Wrase, 1994, Carterus 364

neglectus Audinet‑Serville, 1821, Harpalus 379

neglectus Schaum, 1845, Hydroporus 63

neglectus Erichson, 1837, Ilybius [Agabus] 42

neglectus G. Ochs, 1925, Orectochilus 29

neglectus A. Morawitz, 1862, Pterostichus 509

neglectus Ghidini, 1935, Trechus 341

neglectus Schaum, 1864, Zabrus 569



negligens Dejean, 1828, Pedius [Feronia] 478

negligens Sturm, 1824, Pterostichus [Platysma] 491



negotinensis Reitter, 1896, Carabus 173

negrei Hieke, 1976, Amara 564

negrei Jeannel, 1953, Apoduvalius 295

negrei Habu, 1958, Bembidion 255

negrei Breuning, 1966, Carabus 179

negrei A. Gaudin, 1947, Geotrechus 311

negrei Perrault, 1985, Leistus 81

negrei Ledoux & Roux, 1992, Nebria 94

negrei Mateu, 1950, Trechus 330

negrei Mateu, 1952, Trymosternus 410



negreianus Deuve, 1994, Carabus 148

negrita Wollaston, 1854, Microlestes [Dromius] 423

nemethi Guignot, 1950, Nebrioporus [Potamonectes] 67

nemoralis Olivier, 1790, Calomera [Cicindela] 100

nemoralis O. F. Müller, 1764, Carabus 130

nemoralis Chaudoir, 1866, Platyderus 523

nemoralis Graëlls, 1851, Pterostichus [Argutor] [NP] 491



nemoralis Heer, 1837, Pterostichus [NO] 500

nemorensis Lapouge, 1925, Carabus 154

nemorivagus Duftschmid, 1812, Anisodactylus [Carabus] 361

nemotoi Straneo, 1989, Poecilus [Pterostichus] 483

nenkaoshanensis N. Ito, 1997, Trichotichnus 396

neochinensis Deuve & Li, 1998, Carabus 149

neoguineensis Régimbart, 1892, Limbodessus [Bidessus] 57

neoguineensis V. B. Guéorguiev & Rocchi, 1993, Rhantus 47

neomixtus Tarrier, 1975, Carabus 166

Neotachys Kult, 1961, Polyderis 274

neowinkleri Deuve, 1991, Carabus 182


neoziegleri Deuve, 1991, Carabus 196

nepalense Morvan, 1998, Nipponagonum 461

nepalense Jedlika, 1965, Xestagonum [Agonum] 468

nepalensis Hieke, 1981, Amara 553

nepalensis Kirschenhofer, 1994, Anchista 408

nepalensis Casale, 1980, Aparupa 456



nepalensis Motschulsky, 1858, Brachyodes 444

nepalensis Jaeger & N. Ito, 1995, Bradycellus 402



nepalensis Hope, 1831, Broscus [Percus] 236

nepalensis Kirschenhofer, 1990, Calathus 528

nepalensis Habu, 1978, Callistomimus 348



nepalensis Mandl, 1965, Carabus 162

nepalensis Hope, 1831, Chlaenius 355

nepalensis Mandl, 1965, Cychropsis 202

nepalensis Morvan, 1999, Deliaesianum 457

nepalensis Morvan, 1992, Deltocolpodes 457

nepalensis J. Schmidt, 1994, Deltomerodes 281

nepalensis Hope, 1831, Desera 358

nepalensis G. Ochs, 1929, Dineutus 26

nepalensis Kirschenhofer, 1996, Dischissus 447

nepalensis Jedlika, 1966, Dromius 421

nepalensis Jedlika, 1965, Ebertius [delete] 237

nepalensis Baehr, 1998, Gunvorita 573



nepalensis Jedlika, 1966, Harpalus 386

nepalensis Satô, 1961, Hydaticus 54

nepalensis Brancucci, 1980, Hydronebrius 41

nepalensis Löbl, 1994, Hydroscapha 25

nepalensis Morvan, 1998, Kar 459

nepalensis Brancucci, 1989, Lacconectus 48

nepalensis Morvan, 1979, Laemostenus [Pristonychus] 538

nepalensis Jedlika, 1965, Leistus 81

nepalensis Morvan, 1972, Lesticus 471

nepalensis Morvan, 1977, Mastax 217

nepalensis Habu, 1973, Megalonychus 460

nepalensis Sciaky & Facchini, 1997, Microzargus 439

nepalensis Jedlika, 1965, Mimocolliuris [Colliuris] 444



nepalensis Mandl, 1965, Myriochila [Cicindela] 113

nepalensis Naviaux, 1994, Neocollyris 117

nepalensis Dostal, 1986, Notiophilus 97

nepalensis Habu, 1979, Orthogonius 446

nepalensis Kirschenhofer, 1994, Parena 434

nepalensis Kirschenhofer, 1994, Phloeozetus 438

nepalensis V. B. Guéorguiev, 1968, Platambus [Stictogabus] 44

nepalensis Morvan, 1996, Platynus [Colpodes] 465

nepalensis Kirschenhofer, 1999, Praepristus 437


nepalensis Straneo, 1977, Pterostichus 510

nepalensis Morvan, 1999, Skorlagad 466

nepalensis Jedlika, 1965, Stenolophus [Egadroma] 405

nepalensis Jedlika, 1964, Syntomus [Metabletus] 433

nepalensis Morvan, 1994, Synuchus 545

nepalensis Probst & Wiesner, 1994, Therates 116

nepalensis Morvan, 1998, Tostkar 467

nepalensis Morvan, 1972, Trechus 328

nepalensis Morvan, 1994, Trigonotoma 520

nepalensis Morvan, 1982, Xestopus 530

nepta Ishikawa, 1984, Carabus [Leptocarabus] 159



nereensis A. Nicolas, 1898, Carabus 140

nereensis Heyden, 1889, Zabrus 569

neresheimeri J. Müller, 1929, Bembidion 264

neresheimeri Wagner, 1915, Dyschirius 226

nescium Lutshnik, 1938, Bembidion 261

nestor Breuning, 1934, Carabus [Apotomopterus] 128

netolitzkyanum Schatzmayr, 1940, Bembidion 249

netolitzkyi Krausse, 1910, Bembidion 264

netolitzkyi J. Müller, 1908, Duvalius [Trechus] 306

netuschili Reitter, 1905, Philorhizus [Dromius] 426

netuschili Reitter, 1903, Tachys 275

neuhoffii Cederhjelm, 1798, Nebrioporus [Dytiscus] 66

neumanni J. Müller, 1911, Duvalius [Trechus] 307



neumanni Jeannel, 1921, Trechus 340

neumeyeri Schaum, 1856, Carabus 179

neustria Rivalier, 1962, Cylindera 111

nevadense Jeanne, 1974, Bembidion [Ocydromus] 19, 245

nevadensis Håkan Lindberg, 1939, Agabus 40



nevadensis Jeanne, 1967, Elaphrus 207

nevadensis K. Daniel & J. Daniel, 1898, Harpalus 379

nevadensis Sharp, 1882, Hydroporus 63

nevadensis Jeannel, 1947, Pterostichus [Haptoderus] 492

nevadensis Jeanne, 1967, Trechus 331

nevelskii Shilenkov, 1996, Carabus 170

nevesi Jeannel, 1957, Typhloreicheia 222

newar Deuve, 1988, Trechus 336



newmanni G. R. Waterhouse, 1833, Notiophilus 97

ngi Deuve, 1994, Carabus 126



ngyukmingi Deuve, 1991, Carabus 142

nianus Fedorenko, 1993, Dyschirius 229

niba Uéno, 1998, Epaphiopsis 309

nicaeense Jeannel, 1941, Bembidion [Peryphus] 269

nicaeense Schuler, 1962, Bembidion [Testediolum] 269

nicaeensis A. Villa & G. B. Villa, 1835, Pterostichus [Steropus] 511



nicanor Haury, 1878, Carabus 196

nicanus Motschulsky, 1865, Chlaenius 351

nichollsii Wollaston, 1864, Paranchus [Anchomenus] 462

nicobaricus L. Redtenbacher, 1867, Leiodytes [Hydroporus] 57

nicolae Auvray, 1995, Carabus 131

nicolae Jeanne, 1972, Trechus 341

nicolaiensis Lapouge, 1921, Carabus 133

nicolasi Schuler, 1965, Calosoma [Campalita] 120

nicolasi Chobaut, 1897, Carabus 195

nicolasi Lapouge, 1925, Carabus 145

nicolasi Reitter, 1888, Carabus 197

nicolasi Venet, 1932, Carabus 168

nicolasi Brancucci, 1986, Lacconectus 48



nicolasi Heyden, 1889, Pterostichus 504

nicolasiellus Breuning, 1934, Carabus [Mesocarabus] 168

nicoleae Moncoutier, 1986, Trechus 334



nicolianus Moncoutier, 1990, Trechus 334

niedli Jedlika, 1965, Bembidion 256

nietneri Putzeys, 1866, Coryza 221

nietneri Schaum, 1863, Tachyta [Tachys] 277

nigellatus Kirschenhofer, 1991, Pterostichus 509



nigellus Bleuse, 1914, Carabus 195

nigellus Mannerheim, 1853, Hydroporus 63

nigellus A. Morawitz, 1862, Pterostichus 506

niger Heer, 1841, Amblystomus [Stenolophus] 360



niger A. Nicolas, 1898, Carabus 168

niger Kraatz, 1877, Carabus 175

niger Kraatz, 1886, Carabus [Eutelocarabus] 183

niger Schilsky, 1888, Carabus 180

niger Semenov, 1887, Carabus 153

niger Ahrens, 1830, Dyschirius [Clivina] 227

niger Seidlitz, 1887, Haliplus 32

niger Andrewes, 1937, Holcoderus 436



niger Sturm, 1835, Hydroporus 62

niger Gebler, 1847, Leistus 82



niger Heyden, 1880, Perileptus 288

niger Jedlika, 1953, Pseudotaphoxenus [Taphoxenus] 541

niger Schaller, 1783, Pterostichus [Carabus] [NP] 509

niger Schilsky, 1888, Pterostichus [Calopterus] [HN] 488

niger Lacordaire, 1835, Rhantus 45

niger Jedlika, 1962, Trichotichnus 396

nigeriensis Omer‑Cooper, 1974, Bidessus 55

nigeriensis Omer‑Cooper, 1971, Hyphydrus 74

nigerrima Chaudoir, 1846, Nebria 91



nigerrimus Heyden, 1892, Acupalpus 398

nigerrimus Jedlika, 1962, Brachinus 215

nigerrimus Kraatz, 1886, Carabus 135

nigerrimus Chaudoir, 1837, Platynus [Anchomenus] 464

nigerrimus Habu, 1942, Platynus [Agonum] [HN] 463

nigerrimus Dejean, 1828, Pterostichus [Feronia] [HN!] 499



nigerrimus Sturm, 1824, Pterostichus [Platysma] 500

nigra Chaudoir, 1837, Amara [Bradytus] 553

nigra Motschulsky, 1850, Cephalota [Cicindela] 102

nigra Beuthin, 1898, Cicindela 107

nigra Dokhtouroff, 1885, Cicindela 104

nigra Lallemant, 1868, Cicindela 106

nigra W. Horn, 1900, Megacephala 118

nigraelabris Dokhtouroff, 1885, Cicindela 104

nigrans Barthe, 1912, Bembidion 247

nigrans A. Morawitz, 1862, Harpalus 379

nigrans Bates, 1889, Scalidion [Lebia] 430

nigrescens Ledoux & Roux, 1995, Archastes 79



nigrescens Friederichs, 1903, Bembidion 264

nigrescens Schilsky, 1888, Bembidion 257

nigrescens Westhoff, 1881, Carabus 135

nigrescens Heer, 1838, Cicindela 103

nigrescens Casale & Ledoux, 1996, Laemostenus 536



nigrescens Favre, 1890, Stictotarsus [Hydroporus] 69

nigricans Wiedemann, 1821, Chlaenius 353



nigricans Bedel, 1906, Cymindis 413

nigricans Schilsky, 1888, Harpalus 376

nigricans Schrank, 1781, Hydroporus [Dytiscus] 64

nigricans Newman, 1833, Leistus 84

nigricans Ledoux & Roux, 1999, Nebria [add] 89



nigricans Schilsky, 1888, Pterostichus [Calopterus] 487

nigriceps Dejean, 1829, Acupalpus 398

nigriceps LeConte, 1848, Agonum 454

nigriceps A. Morawitz, 1863, Badister 441

nigriceps Preller, 1862, Hydroporus 64

nigriceps Schaum, 1864, Hydroporus 62

nigriceps Schilsky, 1888, Nebria 90

nigriceps Dejean, 1831, Perigona [Bembidium] 449



nigriceps Sturm, 1824, Pterostichus [Platysma] 508

nigricolle L. Redtenbacher, 1849, Bembidion 250

nigricollis Zoubkoff, 1833, Agabus [Colymbetes] 38

nigricollis Desbrochers des Loges, 1871, Hydroporus 65

nigricollis Gené, 1839, Lebia 428

nigricollis Andrewes, 1925, Nebria 91

nigricollis Bielz, 1850, Stenolophus 406

nigricollis Paulino de Oliveira, 1876, Stenolophus 406

nigricolor Reitter, 1923, Cymindis 411

nigricorne Panzer, 1793, Agonum [Carabus] 452

nigricorne Gyllenhal, 1827, Bembidion 252

nigricornis C. G. Thomson, 1857, Amara 550

nigricornis Stephens, 1835, Anisodactylus [Harpalus] 361

nigricornis Gebler, 1829, Brachinus 216



nigricornis Krynicki, 1832, Brachinus 19, 215

nigricornis Brullé, 1834, Calodromius [Dromius] 419

nigricornis Dejean, 1826, Carabus 196

nigricornis Fabricius, 1787, Chlaenius [Carabus] 350

nigricornis Chaudoir, 1877, Demetrias 419

nigricornis Motschulsky, 1844, Dyschirius 226



nigricornis Schaufuss, 1865, Laemostenus [Pristonychus] 536

nigricornis A. Villa & G. B. Villa, 1833, Nebria 90

nigricornis Motschulsky, 1864, Neocollyris [Collyris] 117

nigricornis Motschulsky, 1844, Trechus 325

nigricoxis Motschulsky, 1865, Chlaenius 353

nigridorsis Nakane, 1962, Brachinus 216



nigrifemoris Habu & Inouye, 1963, Cymindis 415

nigrifrons Fauvel, 1863, Tachys 276

nigrinopomeranus Rade, 1895, Carabus 135

nigrinus Motschulsky, 1866, Carabus 157



nigrinus Schilsky, 1888, Carabus 184

nigrinus Westhoff, 1881, Carabus 135

nigrinus Andrewes, 1925, Elaphropus [Tachys] [DA] 273



nigrinus Schilsky, 1888, Harpalus 372

nigrinus Bates, 1873, Peronomerus 448

nigrinus Putzeys, 1847, Trechus 336

nigripalpis J. R. Sahlberg, 1880, Agabus 36

nigripalpis Stephens, 1835, Harpalus 386

nigripalpis Jedlika, 1963, Poecilus [Pterostichus] 483

nigripalpis Poppius, 1906, Pterostichus [Cryobius] 491



nigripennis A. Fleischer, 1898, Carabus 196

nigripennis G. Hauser, 1921, Carabus [Coptolabrus] [HN] 141

nigripennis Imura, 1996, Carabus 124

nigripennis Chaudoir, 1856, Chlaenius 354



nigripennis Gistel, 1857, Dolichus [Matulus] 530

nigripennis Chaudoir, 1877, Drypta 358

nigripennis Chaudoir, 1861, Galerita 359

nigripennis Motschulsky, 1844, Harpalus 380

nigripennis Wojas, 1993, Leistus 83

nigripennis Ferrante, 1908, Megacephala 118

nigripennis J. R. Sahlberg, 1903, Ophonus 390

nigripennis Bates, 1873, Pentagonica 448

nigripennis Tanaka, 1958, Pterostichus [HN] 501


nigripes A. Costa, 1847, Agabus [Colymbetes] 40

nigripes Gersdorf, 1953, Anisodactylus 361

nigripes Motschulsky, 1850, Asaphidion [Tachypus] 242

nigripes Kirby, 1837, Bembidion [Notaphus] 254



nigripes G. Hauser, 1922, Carabus [HN] 150

nigripes Heyden, 1875, Carabus 139

nigripes Dejean, 1826, Chlaenius 352

nigripes Faldermann, 1836, Chlaenius 352

nigripes Chaudoir, 1876, Drypta 358

nigripes J. R. Sahlberg, 1913, Harpalus 372

nigripes Letzner, 1885, Harpalus [HN] 378

nigripes Sturm, 1818, Harpalus 373

nigripes Reitter, 1894, Idiomelas [Stenolophus] 404



nigripes Dejean, 1825, Lebia 428

nigripes Pic, 1925, Nebria 86

nigrita Chaudoir, 1844, Amara 548

nigrita Dejean, 1825, Cicindela 104



nigrita Krynicki, 1832, Cicindela 104

nigrita Chaudoir, 1848, Galerita 359

nigrita Van de Poll, 1889, Hexagonia 407

nigrita Fabricius, 1792, Hydroporus [Dytiscus] 63

nigrita Paykull, 1790, Pterostichus [Carabus] 511



nigritarsis J. R. Sahlberg, 1882, Graptodytes [Hydroporus] 59

nigritarsis C. R. Sahlberg, 1827, Harpalus 379



nigritarsis Stephens, 1827, Lebia [Lamprias] 427

nigritulus Reitter, 1894, Bradycellus 401

nigritulus Kraatz, 1876, Carabus 186

nigritulus Kraatz, 1881, Carabus 171

nigritulus Kraatz, 1882, Carabus 172

nigritulus Gschwendtner, 1936, Laccoporus [Laccophilus] 77

nigritulus Wollaston, 1863, Perileptus 289

nigritus Reiche & Saulcy, 1855, Laemostenus [Sphodrus] 539

nigriventris Lutshnik, 1927, Amara 551

nigriventris C. G. Thomson, 1857, Philorhizus [Dromius] 426

nigriventris Motschulsky, 1860, Rhantus 46

nigriventris Newbery & Sharp, 1915, Rhantus 46

nigriventris A. Zimmermann, 1919, Scarodytes [Deronectes] 69



nigroaeneum Polentz, 1937, Calosoma 120

nigroaeneus Marsham, 1802, Ilybius [Dytiscus] 42

nigroaeneus Régimbart, 1907, Orectochilus 28



nigrocaeruleus Stephens, 1828, Harpalus 381

nigrocruciatus Wollaston, 1854, Trechus 336



nigrocyanescens Carret, 1904, Carabus 180

nigrocyaneus Kraatz, 1877, Carabus 175

nigrofasciata Schmidt‑Göbel, 1846, Ophionea 444



nigrolimbata Péringuey, 1908, Tachyura [Tachys] 21, 277

nigrolineata Chaudoir, 1852, Parena [Phloeodromus] 434

nigrolineatus Steven, 1808, Hygrotus [Hyphydrus] 72

nigromaculatus Goeze, 1777, Agabus [Dytiscus] 40

nigromarmoratus Clark, 1864, Hydaticus 53

nigrominimus Deuve & Mourzine, 1997, Carabus 129



nigromontana Lafer, 1978, Amara 567

nigronotatus Clark, 1863, Hyphydrus 74

nigroparvula Mandl, 1967, Calomera [Cicindela] 100

nigropiceum Marsham, 1802, Bembidion [Carabus] 251



nigropolitulus Mandl, 1971, Carabus 157

nigropunctatostriatus Mandl, 1986, Callistomimus 348



nigropunctatus Joseph, 1870, Laemostenus [Sphodrus] 535

nigropunctatus Motschulsky, 1860, Rhantus 46

nigrosericans Heller, 1923, Sericoda [Anchomenus] 466

nigrosericeus Goeze, 1777, Harpalus [Carabus] 385

nigrosignata Régimbart, 1895, Yola 57



nigrostriatus DeGeer, 1774, Pterostichus [Carabus] [NO] 509

nigrosuturatus Mandl, 1978, Chlaenius 354



nigrothoracicum Goeze, 1777, Bembidion [Carabus] 261

nigrotibialis N. Ito, 1999, Trichotichnus 394

nigrotibitarsis Mandl, 1978, Chlaenius 353

nigrotinctum Schatzmayr, 1941, Bembidion 255

nigroviolaceus Morvan, 1978, Pterostichus 510

nigrovirescens Reitter, 1889, Carabus 175

nigrovittatus Clark, 1864, Hydaticus 53

nigrum Dejean, 1828, Agonum 451



nigrum Stephens, 1828, Bembidion [Lopha] 266

nigrum W. Kolbe, 1918, Bembidion 245

nigrum Parry, 1845, Calosoma 121

nigrum J. Schmidt, 1996, Xestagonum 468


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