Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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obliquelunatum Motschulsky, 1844, Bembidion [Peryphus] 255

obliquesignatus Solsky, 1871, Metablus [Metabletus] 431

obliquesignatus Bielz, 1852, Porhydrus [Hydroporus] 68

obliquetruncatus Perris, 1875, Brachinus 212

obliquevittata Fleutiaux, 1898, Cicindela 108

obliquulus Belousov & Kabak, 1998, Trechus 335

obliquum Sturm, 1825, Bembidion 254

obliquus C. G. Thomson, 1875, Carabus 166

obliquus Fabricius, 1787, Haliplus [Dytiscus] 30



obliquus Motschulsky, 1850, Stenolophus 406

obliquus Uéno, 1985, Trechiama 323



obliterata Sumakov, 1909, Clivina 219

obliterata Pic, 1925, Eurynebria 79

obliterata Wollaston, 1865, Licinopsis [Pristonychus] 540



obliterata Jacquemet, 1882, Lophyra [Cicindela] 113

obliteratus Wollaston, 1865, Calathus 528

obliteratus Fischer von Waldheim, 1828, Carabus 199

obliteratus Andrewes, 1929, Oxylobus 231

obliteratus Gebler, 1848, Patrobus 285



oblitus Sharp, 1914, Gyrinus 28

oblitus Dejean, 1829, Harpalus 379



oblitus Boieldieu, 1859, Philorhizus [Dromius] 426

oblonga Dejean, 1831, Batoscelis [Agonoderus] 400



oblonga Motschulsky, 1865, Cymindis [Tarus] 414

oblonga Motschulsky, 1850, Nebria 87

oblongicollis Joseph, 1871, Typhlotrechus [Anophthalmus] 344

oblongior Deuve, 1992, Carabus 128

oblongiusculus Régimbart, 1886, Orectochilus 29

oblongiusculus Motschulsky, 1850, Pterostichus [Pseudocryobius] [DA] 491

oblongiusculus Dejean, 1829, Scybalicus [Harpalus] 363

oblongoparallelus Maan, 1932, Pterostichus 496

oblongopunctatus Fabricius, 1787, Pterostichus [Carabus] 487

oblongulus Motschulsky, 1866, Carabus 197

oblongum Dejean, 1831, Agonum 453

oblongum Dejean, 1831, Bembidion 250

oblongus Dejean, 1831, Abax [Feronia] 470

oblongus Dejean, 1825, Brachinus 216



oblongus Sturm, 1815, Carabus 196

oblongus Illiger, 1801, Copelatus [Dytiscus] 47

oblongus Putzeys, 1846, Dyschirius 225

oblongus Casey, 1914, Harpalus 377

oblongus Stephens, 1835, Laccornis [Hydroporus] 75

oblongus Dejean, 1828, Laemostenus [Pristonychus] 533

oblongus Ledoux, 1984, Ledouxius [Penetretus] 284

oblongus Dejean, 1826, Licinus 443


oblongus Schaum, 1858, Ophonus [Harpalus] 389

oblongus Fabricius, 1792, Oxypselaphus [Carabus] 462

oblongus Fabricius, 1792, Paranchus [Carabus] 462

oblongus Motschulsky, 1866, Percus 478

oblongus Chaudoir, 1843, Scarites 234

oblongus Tschitschérine, 1906, Trichotichnus [Amaroschesis] 394



oblongus Motschulsky, 1862, Typhlotrechus [Anophthalmus] 344

obockianus Fairmaire, 1892, Stenolophus [Hydroporomorpha] 405



obockianus Fairmaire, 1892, Triaenogenius 407

obotritica Schulz, 1909, Cicindela 107

obovalis Gebler, 1829, Carabus 164

obovatus J. R. Sahlberg, 1875, Agabus [Gaurodytes] 36

obovatus Fischer von Waldheim, 1828, Carabus 164



obovatus J. R. Sahlberg, 1880, Hydroporus 63

obscura Motschulsky, 1844, Amara 551

obscura Schilsky, 1888, Cicindela 106

obscura Fabricius, 1798, Cylindera [Cicindela] 109

obscura Laporte, 1834, Neocollyris [Collyris] 117

obscura Lesne, 1891, Neocollyris [Collyris] 117

obscura Mateu, 1977, Parena 434



obscurata Chaudoir, 1843, Cicindela 104

obscurata J. R. Sahlberg, 1903, Nebria 92

obscuratus Halbherr, 1908, Acupalpus 398

obscuratus G. Hauser, 1921, Carabus [Coptolabrus] 143

obscuratus Mannerheim, 1853, Colymbetes [Cymatopterus] 45

obscuratus Schaufuss, 1861, Laemostenus [Sphodrus] 535

obscuratus Poppius, 1906, Pterostichus [Cryobius] 491

obscurecoerulescens Ménétriés, 1848, Cylindera [Cicindela] 109

obscuredilatata W. Horn, 1914, Cylindera [Cicindela] 111

obscureguttata Dupuis, 1910, Calomera [Cicindela] 100

obscurellum Motschulsky, 1845, Bembidion [Peryphus] 262

obscurellus Dejean, 1831, Tilius [Dromius] 433

obscuriceps Régimbart, 1907, Orectochilus 28



obscuriceps Morvan, 1972, Trechus 335

obscuricollis Chaudoir, 1837, Calathus 527

obscuricollis Reitter, 1899, Dicheirotrichus [Trichocellus] 403



obscuricollis Motschulsky, 1844, Lebia [Omalomorpha] 428

obscuricollis Pic, 1905, Trichis 439

obscuricolor Breit, 1912, Pterostichus [Platysma] [HN] 485

obscuricolor Pic, 1903, Pterostichus 504

obscuricorne Fassati, 1949, Bembidion 262

obscuricornis Motschulsky, 1859, Amara 556

obscuricornis Brullé, 1834, Brachinus 213

obscuricornis Ménétriés, 1832, Brachinus 214

obscuricornis Beuthin, 1885, Carabus 194

obscuricornis Reitter, 1899, Dicheirotrichus [Trichocellus] 403

obscuricornis Motschulsky, 1844, Harpalus 382

obscuricornis Stephens, 1828, Harpalus 381

obscuricornis Sturm, 1818, Harpalus 376

obscuricornis J. R. Sahlberg, 1921, Hydroporus 62

obscuricornis Reitter, 1885, Leistus 84

obscuricornis Poppius, 1906, Pterostichus [Cryobius] 491

obscurior J. R. Sahlberg, 1875, Agabus [Acatodes] 36

obscurior G. Hauser, 1921, Carabus [Coptolabrus] [HN] 141

obscurior Semenov, 1888, Carabus 178

obscurior Schreber, 1759, Cylindera [Cicindela] 109

obscurior Kirby, 1837, Elaphrus 206

obscurior Pic, 1911, Oedesis [Eucarterus] 366

obscurior Desbrochers des Loges, 1871, Yola [Hydroporus] 57

obscuripennis J. R. Sahlberg, 1875, Agabus [Gaurodytes] 36

obscuripennis Motschulsky, 1864, Brachinus 216



obscuripennis Reitter, 1889, Calathus 525

obscuripennis Fairmaire, 1886, Paraglycia [Glycia] 409



obscuripennis Eichler, 1924, Thalassophilus 346

obscuripes Bates, 1873, Amara 550

obscuripes J. Müller, 1943, Bembidion 266

obscuripes Motschulsky, 1860, Hydroporus 63

obscuripes Everts, 1893, Molops 474

obscuripes Motschulsky, 1850, Ophonus 389

obscuripes Schaufuss, 1887, Platymetopus 393

obscuripes Poppius, 1906, Pterostichus [Argutor] 491

obscuritarsis Motschulsky, 1864, Brachinus 216



obscuritarsis Motschulsky, 1844, Dicheirotrichus [Ophonus] 403

obscuritarsis Chaudoir, 1879, Loxocrepis [Colpodes] 460



obscuriusculus Eidam, 1930, Carabus 195

obscurofemorata Mandl, 1971, Neocollyris 116

obscuroguttatus Duftschmid, 1812, Syntomus [Lebia] 433

obscuroides Paulino de Oliveira, 1876, Dixus [Aristus] 365

obscurum Chaudoir, 1842, Agonum 452

obscurum Paykull, 1790, Agonum [Carabus] 452

obscurum Andrewes, 1925, Asaphidion 242



obscurum L. Redtenbacher, 1849, Bembidion 250

obscurum Verhoeff, 1890, Bembidion 263

obscurus Heer, 1837, Brachinus 213

obscurus Géhin, 1885, Carabus [Procerus] 185

obscurus Klug, 1832, Chlaenius 350

obscurus Chaudoir, 1854, Chydaeus 362

obscurus Xie & Yu, 1991, Craspedophorus 447



obscurus Stephens, 1835, Demetrias 419

obscurus Dejean, 1825, Dixus [Ditomus] 365



obscurus Gyllenhal, 1827, Dyschirius [Clivina] 228

obscurus Gschwendtner, 1922, Dytiscus 52

obscurus P. Moravec & Wrase, 1995, Eotrechodes [Thalassophilus] 345

obscurus Chaudoir, 1856, Harpaglossus [Chlaenius] 356

obscurus Habu, 1979, Holcoderus 436

obscurus J. R. Sahlberg, 1903, Hydroglyphus [Bidessus] 56

obscurus Sturm, 1835, Hydroporus 63



obscurus Motschulsky, 1859, Hydrovatus 70

obscurus Régimbart, 1895, Hydrovatus 70

obscurus Marsham, 1802, Ilybius [Dytiscus] 43

obscurus Panzer, 1795, Laccophilus [Dytiscus] 77

obscurus Reitter, 1905, Leistus 82

obscurus Fabricius, 1792, Ophonus [Carabus] 392

obscurus Herbst, 1784, Oxypselaphus [Carabus] 462



obscurus Fairmaire & Laboulbène, 1854, Poecilus [Feronia] [HN] 481

obscurus Faldermann, 1836, Poecilus 482

obscurus Dejean, 1828, Pterostichus [Feronia] 501

obscurus Della Beffa, 1931, Stictotarsus [Hydroporus] 69

obscurus J. R. Sahlberg, 1913, Tachys 276

obscurus Uéno, 1955, Trephionus 547



obsidianus Kolenati, 1846, Sphaerius [Microsporus] 26

obsoleta Duftschmid, 1812, Amara [Carabus] 551

obsoleta Faldermann, 1836, Cymindis 417

obsoleta V. E. Jakovlev, 1885, Cymindis 415

obsoleta Semenov, 1889, Loricera 98



obsoletum Dejean, 1831, Bembidion 250

obsoletus Fischer von Waldheim, 1823, Carabus 199

obsoletus G. Hauser, 1921, Carabus [Damaster] [HN] 148

obsoletus Sturm, 1815, Carabus [HN] 151

obsoletus Dejean, 1829, Dicheirotrichus [Harpalus] 402

obsoletus Agazzi, 1970, Doderotrechus [Typhlotrechus] 298

obsoletus Putzeys, 1846, Dyschirius 228

obsoletus Westhoff, 1881, Haliplus 32

obsoletus Aubé, 1838, Hydroporus 63



obsoletus V. E. Jakovlev, 1885, Mnuphorus [Corsyra] 358

obsoletus Louwerens, 1953, Platynus [Colpodes] 465



obsoletus P. Rossi, 1790, Sphodrus [Carabus] 543

obsoletus Dejean, 1825, Syntomus [Dromius] 433

obtritus Chaudoir, 1857, Carabus [Procrustes] 187

obtritus Lapouge, 1898, Carabus 197

obtritus Lapouge, 1925, Carabus 130

obtusa LeConte, 1855, Amara 559

obtusa Stephens, 1828, Amara 548

obtusa Jeannel, 1942, Cymindis 412

obtusangalus Schaufuss, 1867, Laemostenus [Antisphodrus] 535



obtusangula Baliani, 1934, Amara 564

obtusangulus Ganglbauer, 1889, Molops 473



obtusangulus Faldermann, 1835, Neophygas [Harpalus] 387

obtusangulus J. Müller, 1923, Platyderus 523



obtusangulus Reitter, 1883, Pterostichus [Feronia] 501

obtusangulus Reitter, 1885, Zabrus 572

obtusicollis Schauberger, 1932, Chydaeus 362



obtusicollis Motschulsky, 1849, Cymindis 414

obtusicollis Putzeys, 1878, Harpalus 372

obtusicollis Bates, 1883, Leistus 83

obtusicollis Schauberger, 1936, Trichotichnus 394

obtusioides Jeannel, 1927, Trechus 337

obtusipennis Ishikawa & Kim, 1983, Carabus [Damaster] 142



obtusipennis J. R. Sahlberg, 1875, Hydroporus 62

obtusipennis Régimbart, 1892, Orectochilus 28



obtusiusculus Chaudoir, 1871, Patrobus 285

obtusiusculus Jeannel, 1941, Tachys [Eotachys] 275

obtusiusculus Ganglbauer, 1889, Trechus 337

obtusum Audinet‑Serville, 1821, Bembidion 265

obtusus Jeannel, 1926, Aphoenops 294

obtusus Ganglbauer, 1886, Carabus [Tribax] 133

obtusus Boheman, 1848, Copelatus 48

obtusus Motschulsky, 1844, Demetrias 419

obtusus Gebler, 1833, Harpalus [Anisodactylus] 372

obtusus Motschulsky, 1855, Hydrovatus 70

obtusus Sharp, 1882, Ilybius 42

obtusus Sharp, 1882, Laccophilus 77

obtusus Chaudoir, 1861, Laemostenus [Sphodrus] 533



obtusus Jeannel, 1942, Pterostichus [HN] 513

obtusus Jeannel, 1947, Pterostichus [Haptoderus] [HN!] 490



obtusus Sturm, 1824, Pterostichus 488

obtusus Erichson, 1837, Trechus 337



obversus Motschulsky, 1844, Carabus 170

obydovi E. Berlov & O. Berlov, 1996, Carabus 165

occasus Csiki, 1930, Pterostichus 507

occidentale J. Müller, 1918, Bembidion 260

occidentalis Mateu, 1947, Brachinus 214

occidentalis Born, 1907, Carabus 187

occidentalis Jeanne, 1969, Carabus 156

occidentalis Mandl, 1977, Cychrus 202



occidentalis Jakobson, 1908, Cymindis 413

occidentalis Knirsch, 1929, Duvalius [Duvaliotes] 300

occidentalis Lassalle, 1993, Laemostenus 533

occidentalis G. Ochs, 1955, Orectochilus 28

occidentalis Gautier des Cottes, 1870, Orthomus 476


occidentalis Antoine, 1933, Platytarus 410

occidentalis Coiffait, 1953, Pterostichus [Haptoderus] [HN!] 490



occidentalis Pécoud, 1927, Pterostichus 507

occidentalis Bedel, 1895, Scarites 233

occidentalis Fresneda & Fery, 1990, Stictonectes 69

occidentalis Uéno, 1987, Trechiama 323

occidentalis Colas & A. Gaudin, 1935, Trechus 327

occidentalis Freude, 1989, Zabrus 571

occiduum Marggi & Huber, 2001, Bembidion 248



occipitalis Bates, 1892, Clivina 220

occipitalis W. S. MacLeay, 1825, Pheropsophus [Aptinus] 217

occitanus Casale & Vigna Taglianti, 1993, Duvalius 304

occulta Holdhaus, 1924, Typhloreicheia [Reicheia] 223

occultus Sharp, 1882, Bidessus 55

occultus Sharp, 1882, Hydroporus 64

occultus Ballion, 1871, Pseudotaphoxenus [Sphodrus] 541



oceanicus Jeanne, 1971, Carabus 168

oceanicus Bedel, 1905, Graphipterus 359

oceanicus Régimbart, 1886, Haliplus 32

oceanicus Wollaston, 1865, Paradromius [Dromius] 424



oceanicus Carret, 1903, Pogonus 287

ocellata Herbst, 1806, Cicindela 106

ocellatus Báguena Corella, 1935, Hydroglyphus [Bidessus] 56

ochii Uéno, 1996, Ishikawatrechus 312

ochotica R. F. Sahlberg, 1844, Nebria 94

ochotica R. F. Sahlberg, 1844, Pelophila 98

ochoticus Poppius, 1908, Agabus 35



ochoticus Mannerheim, 1852, Carabus 170

ochoticus Motschulsky, 1860, Pterostichus [Steroderus] [HN] 506

ochoticus R. F. Sahlberg, 1844, Pterostichus [Feronia] 491

ochotonarum Bezina, 1996, Carabus 150

ochraceus Sturm, 1824, Pedius [Platysma] 478

ochraceus Gautier des Cottes, 1868, Pterostichus [Feronia] [ND] 515



ochreatus Reiche & Saulcy, 1855, Harpalus 380

ochreatus Dejean, 1831, Trechus 337

ochrias Andrewes, 1925, Polyderis [Tachys] 274

ochroides Andrewes, 1933, Peliocypas 434



ochropterus Duftschmid, 1812, Calathus [Carabus] 527

ochropus Andrewes, 1935, Bembidion 268



ochropus R. F. Sahlberg, 1844, Pterostichus [Feronia] 490

ochsi L. Gaudin, 1925, Aphoenops 294

ochsi Dodero, 1921, Duvalius [Trechus] 305

ochsi Puel, 1925, Laemostenus 532

ochsi Fagniez, 1921, Pterostichus [Troglorites] 514

ocreatus Bates, 1873, Chlaenius 348


octocelis Andrewes, 1924, Lebidia 426

octocola Mannerheim, 1853, Sericoda [Anchomenus] 466

octofoveolatus Mäklin, 1857, Platynus [Anchomenus] 463

octogramma Chaudoir, 1852, Calochroa [Cicindela] 99

octoguttata A. Morawitz, 1862, Lebidia 426

octoguttatus Régimbart, 1895, Pseuduvarus [Bidessus] 57

octomaculata Baudrimont, 1923, Eurynebria 79

octomaculatum Goeze, 1777, Bembidion [Carabus] 270

octonotata Wiedemann, 1819, Calochroa [Cicindela] 99

octopunctata Loew, 1843, Calomera [Cicindela] 100

octopunctata Souverbie, 1855, Eurynebria [Nebria] 79

octopunctatus Apfelbeck, 1904, Pterostichus [HN] 487

octopunctatus Stephens, 1828, Pterostichus 487

octostriata Netolitzky, 1929, Tachyura [Tachys] 277

octussis C. A. Dohrn, 1885, Cicindela 104

oculare J. Müller, 1934, Parazuphium [Zuphium] 573

ocularis Pascoe, 1872, Tenomerga [Cupes] 25

oculata Helliesen, 1912, Amara 556

oculatus Herbst, 1784, Rhantus [Dytiscus] 46

ocylum Jedlika, 1933, Bembidion 244

ocylus Jedlika, 1934, Platynus [Colpodes] 465

odaesaniculus Deuve, 1991, Carabus 149

odaesanus Ishikawa & Kim, 1983, Carabus [Lipaster] 174

odaiensis Kamiya, 1933, Orectochilus 28

odanakai Kasahara, 1995, Apatrobus 280

odelli Andrewes, 1930, Nirmala 476



odessanus Lutshnik, 1925, Pogonistes 286

odontocnemis J. Müller, 1932, Pterostichus 496

odoratus Motschulsky, 1844, Carabus 172

odvarkai Dvoák, 1994, Leistus 82

odvarkai Dvoák, 1995, Pterostichus 505

odysseus Breuning, 1932, Carabus 126



oedimera Jeannel, 1942, Cymindis 414

oenotrius Ravizza, 1975, Pterostichus 511

oertzeni Hieke, 1984, Amara 563

oertzeni Jeanne & F. Battoni, 1988, Calathus 525

oertzeni Ganglbauer, 1888, Carabus 182

oertzeni L. Miller, 1884, Duvalius [Anophthalmus] 305

oertzeni Reitter, 1889, Licinus 443

oertzeni Kraatz, 1886, Speluncarius [Elasmopterus] 516

oertzeni Reitter, 1885, Zabrus 571

oetensis Maan, 1940, Tapinopterus 520

ogaensis Morita, 1995, Pterostichus 502

ogasawarensis Kamiya, 1932, Copelatus 48

ogloblini Lutshnik, 1935, Amara 550


ogloblini Kabak, 1999, Cymindis 418

ogouzicus Deuve & Quéinnec, 1992, Trechus 337



ogumae Matsumura, 1911, Calosoma 120

ogurae Bates, 1883, Agonum [Anchomenus] 454

ogurai Ishikawa, 1969, Carabus 159

ohbayashii Satô, 1985, Gyrinus 27

ohbayashii Ishida, 1968, Pterostichus 502

ohdaiensis Nakane, 1957, Nebria 94

ohdaisanus Nakane, 1963, Apatrobus [Patrobus] 280

ohdaisanus Nakane, 1963, Pterostichus 502

ohkawai Nakane, 1968, Carabus [Apotomopterus] 177

ohkawai Uéno, 1988, Kurasawatrechus 313

ohkawai Uéno, 1993, Trechiama 323

ohkurai Morita, 1992, Bembidion 271

ohkurai Morita, 1996, Pterostichus 502

ohkurai Uéno, 1996, Trechiama 323

ohkurai N. Ito, 1996, Trichotichnus 397

ohminensis Harusawa, 1978, Carabus 159

ohruii Uéno, 1972, Trechiama 323



ohshimai Deuve, 1988, Carabus 124

ohshimai Uéno, 1952, Kurasawatrechus 313

ohshimai Uéno, 1951, Trechiama [Yosiitrechus] 323

ohshimaianus Deuve, 1988, Carabus 190

ohshimaorum Deuve, 1991, Carabus 129

ohtanii Uéno, 1969, Stygiotrechus 320

ohtsukai Morita, 1993, Apatrobus 280

ohtsukai Morita, 1996, Bembidion [Ocydromus] 266

oirat Kabak, 1994, Poecilus 483

oitalensis Obydov, 1998, Carabus 145

okadai Uéno, 1962, Epaphiopsis 309

okamotoi Imura, Su & Osawa, 1998, Cychrus 205

okawai Morita, 1997, Trichotichnus 397

oki Ishikawa & Kubota, 1994, Carabus 178

okianus Nakane, 1961, Carabus [Apotomopterus] 176

okiensis Nakane, 1989, Pterostichus 515

okikulumi Nakajima, 1993, Carabus [Damaster] 122

okinawana Nakane, 1957, Cicindela 108

okinawense Hori & Cassola, 1989, Callytron 101

okinoshimanus Imura, Dejima & Mizusawa, 1993, Carabus 177


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