Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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okulovi Lafer, 1988, Oreodytes 68

okumicus Belousov, 1992, Carabus 131

okumurai Ishikawa, 1966, Carabus [Apotomopterus] 176

okutamae Tanaka, 1963, Pterostichus 502

okutamaensis Ishikawa, 1966, Carabus 149

okutanii Habu, 1952, Callistomimus 348



okutanii Habu, 1958, Clivina 220

olcesii Fairmaire, 1871, Graniger [Ophonus] 366

olcesii Fairmaire, 1856, Paussus 210

olegi Belousov, 1992, Carabus 132

olegi Belousov, 1991, Pterostichus 496

olegi Kataev, 1999, Selenotichnus 393

olegiana Kabak, 1999, Cymindis 416

olegleonidovici Fassati, 1990, Bembidion 259

olemartini Kirschenhofer, 1984, Bembidion 262

olemartini Kirschenhofer, 1986, Peliocypas [Risophilus] 434

olenini Poppius, 1906, Harpalus 375

olens P. Rossi, 1790, Zuphium [Carabus] 574

olgae Arribas, 1993, Brachinus 212

olgae Semenov, 1898, Carabus 133

oliaroi Casale, 1979, Carabus 162

oligops Uéno, 1978, Epaphiopsis 309

oligops Bedel, 1896, Trechus 337

olisipennis Schatzmayr, 1943, Brachinus 213

olisipensis Schatzmayr, 1936, Geocharis [Anillus] 239



olivaceum Dejean, 1828, Agonum 450

olivaceum Gyllenhal, 1827, Bembidion 266

olivaceus G. Hauser, 1932, Carabus [Coptolabrus] 143

olivaceus Géhin, 1876, Carabus 180

olivaceus Boheman, 1858, Cybister 51

olivaceus Carret, 1903, Pogonus 287

oliveirae Reitter, 1884, Acupalpus 399

oliveirae Csiki, 1927, Carabus 179

oliverii Buquet, 1864, Eucolliuris [Casnonia] 443



olivia Bates, 1878, Cylindera [Cicindela] 110

olivieri Zaitzev, 1908, Agabus 38

olivieri Dejean, 1831, Calosoma 121



olivieri Dejean, 1826, Carabus 185

olivieri Laporte, 1834, Carabus 182

olivieri Crotch, 1871, Chlaenius 350

olivieri Dejean, 1826, Nebria 86

olivieria Brullé, 1832, Cicindela 104

olivius Bates, 1873, Metacolpodes [Colpodes] 460

olliveirianus Deuve, 1999, Carabus 183

ollivieri Morvan, 1997, Andrewesius 456

oltenicus Jeannel, 1928, Duvalius 299

olympiadicus Scheibel, 1937, Duvalius 305

olympiae Sella, 1855, Carabus 140

olympiana Jeanne, 1972, Nebria 84

olympica Maan, 1938, Nebria 18, 84



olympicola Breuning, 1969, Cychrus 205

olympicum De Monte, 1946, Bembidion 269

olympicum Meschnigg, 1947, Bembidion 246

olympicus Géhin, 1885, Carabus 186

olympicus Lapouge, 1902, Carabus 185

olympicus L. Redtenbacher, 1843, Morion 443

olympicus Kraatz, 1875, Tapinopterus [Pterostichus] 518

olympicus Piochard de la Brûlerie, 1876, Trechus 337

olympicus Casale, 1977, Winklerites 240

omanensis Schauberger, 1933, Crasodactylus 362

ombrophilus Deuve, 1990, Cychrus 205

omeishanicus Kryzhanovskij, 1994, Andrewesius 456

omensis Born, 1901, Carabus [Orinocarabus] 180

omiades Faldermann, 1836, Cymindis 412

omiensis G. Hauser, 1932, Carabus [Apotomopterus] 125

omisanensis Lassalle, 1999, Carabus 183

ommani Zaitzev, 1908, Agabus 40

omochlorus Andrewes, 1931, Chlaenius 349

omogoensis Nakane, 1972, Pterostichus 507

omostigma Netolitzky, 1914, Bembidion 263

omostigmoides Roubal, 1946, Bembidion 263

omphreodes Reitter, 1898, Carabus 123

ompoensis Deuve, 1991, Carabus 149

ompoensis Jedlika, 1932, Pterostichus 502

onaci Moldovan, 1993, Duvalius 307

ondai Morita, 1995, Amara 554

ondai Uéno, 1993, Trechiama 323

onerosus Belousov & Zamotajlov, 1993, Carabus 200

ongudaicus Belousov & Kabak, 1996, Trechus 337

oni Uéno, 1955, Trechiama 323

oniceps Uéno, 1989, Trechiama 323

onicus Belousov & Kabak, 1994, Trechus 337

onjinsanicus Rapuzzi, 1994, Carabus 174

onobrychidis Iablokoff‑Khnzorian, 1957, Metadromius [Glycia] 421

onocoro Uéno, 1983, Trechiama 323

onodai Imura, Dejima & Mizusawa, 1993, Carabus 177

onoha Bates, 1873, Calleida 408

onoi Takakura, 1987, Jujiroa 459

onthoporus Iablokoff‑Khnzorian, 1957, Laemostenus [Pristonychus] 538

onychinus Dejean, 1825, Trymosternus [Cymindis] 410

ooderus Chaudoir, 1850, Euleptus 458

oodiformis Casey, 1924, Amara 549

oodioides Dejean, 1829, Harpalus 380



ooligbukii Kirby, 1837, Dytiscus 51

ooptera Putzeys, 1865, Amara [Leiocnemis] 563



oopterum Chaudoir, 1850, Bembidion 263

oopterus Chaudoir, 1861, Leistus 82

oopterus Sciaky & Pavesi, 1997, Pseudotaphoxenus 541

oopterus Uéno, 1995, Trechiama 323

opaca Tschitschérine, 1894, Amara 566

opaca Štrba, 1910, Cicindela 103

opaca Chaudoir, 1868, Pseudozaena [Picrus] 208

opacicolor Reitter, 1897, Microdaccus [Dromius] 422

opacinus Ragusa, 1889, Aulonogyrus 26

opacipennis J. Müller, 1913, Anophthalmus [Trechus] 292

opacipennis Motschulsky, 1864, Brachinus 216

opacipennis Reitter, 1895, Carabus 174

opacipennis Jedlika, 1934, Pterostichus 502

opaculus Putzeys, 1875, Carabus 149

opaculus Chaudoir, 1876, Megalonychus [Anchomenus] 460

opacum Semenov, 1889, Holosoma 444



opacus Guignot, 1932, Bidessus 55

opacus Lucas, 1846, Calathus 525



opacus Haury, 1878, Carabus 139

opacus Kraatz, 1895, Carabus 155

opacus Chaudoir, 1852, Diplocheila [Rembus] 439

opacus Erichson, 1841, Dixus [Ditomus] 365

opacus Andrewes, 1925, Elaphropus [Tachys] 273

opacus C. R. Sahlberg, 1819, Gyrinus 27

opacus Wehncke, 1872, Hydroporus 61

opacus Falkenström, 1932, Hygrotus [Coelambus] 72

opacus Aubé, 1837, Ilybius [Agabus] 42

opacus Schaum, 1857, Microdaccus [Apristus] 422

opacus Dejean, 1829, Ophonus [Harpalus] 391

opacus Schmidt‑Göbel, 1846, Orthogonius 446

opacus A. Costa, 1886, Scarites 232

opacus Chaudoir, 1855, Scarites 232

opacus N. Ito, 1998, Trichotichnus 395

opatrinus Germar, 1824, Deronectes [Hydroporus] 58

opermanni Scheibel, 1933, Duvalius 307



operosus Chaudoir, 1859, Percus 478

ophonoides Jedlika, 1958, Ophonus [Penthophonus] 24, 390



opicus Casey, 1914, Harpalus 379

oportica Jedlika, 1958, Cymindis 411

optabilis Dejean, 1829, Harpalus 380



optata Fischer von Waldheim, 1828, Cicindela 104

optatus Andrewes, 1942, Abacetus 347

optatus Sharp, 1884, Platambus [Agabus] 44

optatus Vereschagina & Kabak, 1996, Pseudotaphoxenus 541

optatus Seidlitz, 1887, Stictonectes [Hydroporus] 69

optimus Bates, 1873, Philorhizus [Dromius] 426



opulentum Nietner, 1858, Bembidion 254

opulentus Oberthür, 1898, Carabus 141



opulentus Tschitschérine, 1889, Poecilus 481

oramarensis Shilenkov & Heinz, 1984, Nebria 86



oranensis Schulz, 1909, Calomera [Cicindela] 101

oranensis Géhin, 1885, Carabus 152

oranicus Csiki, 1927, Carabus 161

orbaraensis Dupré, 1988, Aphoenops 294

orbayi Hernando, 1999, Duvalius [to lindbergi] 23, 303

orbicolle Motschulsky, 1865, Agonum [Agonocyrtes] 454

orbicolle Motschulsky, 1850, Bembidion [Philochthus] 264

orbicollis Motschulsky, 1844, Carabus 199

orbicollis N. Ito, 1995, Harpalomimetes 363

orbicollis Motschulsky, 1865, Lindrothius [Calathus] 529

orbicollis A. Morawitz, 1862, Synuchus [Calathus] 546



orbicularis Régimbart, 1899, Hyphydrus 75

orbiculatum Motschulsky, 1839, Calosoma [Carabus] 118

orbiculatus Ledoux & Roux, 1998, Archastes 79

orbipennis Bedel, 1904, Atlantomasoreus [Masoreus] 357

orbitosus Obydov, 1998, Carabus 183

orcinulus Poppius, 1906, Pterostichus 507

orcinus Linder, 1859, Geotrechus [Anophthalmus] 310



ordinaria Semenov, 1904, Cephalota [Cicindela] 102

ordinarior Deuve & Quéinnec, 1993, Trechus 337



ordinata V. E. Jakovlev, 1884, Cephalota [Cicindela] 102

ordinatoides Kurnakov, 1958, Pterostichus 500

ordinatus Gautier des Cottes, 1870, Calathus 525

ordinatus Csiki, 1927, Carabus 164

ordinatus Bates, 1873, Dyschirius 229

ordinatus Fischer von Waldheim, 1823, Pterostichus [Myosodus] 501

ordossicus Semenov, 1889, Pogonus 287



ordubadense Iablokoff‑Khnzorian, 1960, Bembidion 262

orduensis Straneo, 1988, Pterostichus [Haptotapinus] 496

oreades Nègre, 1966, Calathus 525

oreas Bates, 1883, Trechiama [Trechus] 323

oreas Bates, 1891, Trichotichnus [Harpalus] 394

orehovscensis Daffner, 1996, Anophthalmus 293

orensis Breit, 1914, Nebria 90

oreoides Suenson, 1957, Trechiama 323

oreophila C. Zimmermann, 1832, Amara 557

oreophila Glasunov, 1902, Nebria 91

oreophilus K. Daniel & J. Daniel, 1890, Dicheirotrichus [Tachycellus] 402

oreophilus Knirsch, 1927, Neotrechus 315

oreophilus Tschitschérine, 1898, Pterostichus [Feronia] 501

orestes Andrewes, 1933, Dromius 421


orestes Jedlika, 1936, Pterostichus [Neohaptoderus] 501

orestias Andrewes, 1932, Nebria 90



oreteus Ragusa, 1881, Chlaenius 351

oribates A. Gaudin, 1935, Trechus 340



oricalcica O. F. Müller, 1776, Amara [Carabus] [NO] 555

orichalceum Panzer, 1796, Bembidion [Carabus] 247

orichalcicum Duftschmid, 1812, Bembidion [Elaphrus] 257

orichalcicum Illiger, 1798, Bembidion [Elaphrus] 247

orichalcicum Stephens, 1828, Bembidion [Tachypus] 251

orichalcina W. Horn, 1896, Neocollyris [Collyris] 117

oriens Dvoák, 1993, Ditomus [Carterus] 364

orientala Nietner, 1858, Tachyura [Bembidium] 278

orientale Apfelbeck, 1904, Bembidion 249

orientale Peyron, 1858, Bembidion 257

orientalis Kamiya, 1938, Agabus 35

orientalis Lutshnik, 1935, Amara 550

orientalis Gautier des Cottes, 1872, Anisodactylus 361

orientalis Jeannel, 1926, Anophthalmus 292

orientalis A. Gaudin, 1947, Aphoenops 295

orientalis Hrka, 1988, Aptinus 212

orientalis Sciaky & Wrase, 1997, Aristochroodes 471

orientalis Jeannel, 1928, Blemus [Lasiotrechus] 296



orientalis Apfelbeck, 1904, Brachinus 214

orientalis Chaudoir, 1876, Brachinus 216



orientalis Gautier des Cottes, 1866, Calathus 526

orientalis Wehncke, 1876, Canthydrus [Hydrocanthus] 34

orientalis Osculati, 1844, Carabus 147

orientalis Dejean, 1826, Chlaenius 349

orientalis Gschwendtner, 1931, Cybister 50

orientalis Schatzmayr, 1944, Cymindis 412

orientalis Modeer, 1776, Dineutus [Gyrinus] 26

orientalis J. R. Sahlberg, 1913, Dixus [Ditomus] 24, 365

orientalis Putzeys, 1866, Dyschirius 229



orientalis Semenov, 1904, Elaphrus 207

orientalis Schmidt‑Göbel, 1846, Galerita 359

orientalis Régimbart, 1883, Gyrinus 27

orientalis Clark, 1863, Hydroglyphus [Hydroporus] 56



orientalis Hlisnikovský, 1955, Hydroporus 64

orientalis Sharp, 1882, Hydrovatus 71

orientalis Clark, 1863, Hyphydrus 74



orientalis Aubé, 1838, Laccophilus 77

orientalis Jeannel, 1937, Laemostenus [Ceuthosphodrus] 533

orientalis K. Daniel, 1901, Lionychus 432

orientalis Dejean, 1828, Masoreus 357

orientalis Mateu, 1956, Microlestes 423



orientalis Dejean, 1825, Myriochila [Cicindela] 113

orientalis Bänninger, 1949, Nebria 88

orientalis Chaudoir, 1850, Notiophilus 97

orientalis Reitter, 1889, Notiophilus 97

orientalis Reiche & Saulcy, 1855, Olisthopus 461

orientalis Dejean, 1828, Pogonus 287



orientalis Dejean, 1829, Progonochaetus [Selenophorus] 363

orientalis Motschulsky, 1844, Pterostichus [Steropus] 493



orientalis Motschulsky, 1850, Pterostichus [Argutor] 509

orientalis Hope, 1845, Trichotichnus [Amara] 397

orientalis Apfelbeck, 1904, Zabrus 571

orientalis Aubry, 1956, Zabrus 569

orientalissimus Stichel, 1923, Dixus [Ditomus] 365

orienticola Lutshnik, 1935, Amara 550



orienticola Tschitschérine, 1903, Calomera [Cicindela] 99

orienticola Tschitschérine, 1902, Ophonus [Harpalus] 391

orientis Schauberger, 1926, Ophonus [Harpalus] 390

orientis Csiki, 1930, Pterostichus 509

orientosinicus Deuve, 1992, Trechus [Epaphius] 325



origin Nakajima, 1993, Carabus [Acoptolabrus] 122

originalis Schaufuss, 1865, Pseudotaphoxenus 541



orinomus Stephens, 1828, Pterostichus [Omaseus] 487

orinum Andrewes, 1922, Bembidion 246

orion Tschitschérine, 1901, Pterostichus [Platysma] 499

orionis Fagniez, 1913, Aphoenops 294

orionis Jedlika, 1962, Pterostichus 498

orissaensis Vazirani, 1969, Leiodytes [Clypeodytes] 57

orman Jedlika, 1963, Pterostichus 505

ormayanus Csiki, 1944, Carabus 167

ormayi Reitter, 1896, Carabus 173

ormayi Ganglbauer, 1891, Nebria 87

ormayi Ganglbauer, 1891, Trechus 337

ornata Fischer von Waldheim, 1823, Cymindis 414

ornata Klug, 1832, Glycia [Cymindis] 409

ornata Fleutiaux, 1898, Hypaetha [Cicindela] 112

ornata Apetz, 1854, Tachyura [Bembidium] 21, 278



ornatella Bates, 1883, Dolichoctis 436

ornatipennis Schilder, 1953, Hypaetha [Cicindela] 112



ornatipennis Régimbart, 1900, Pseuduvarus [Bidessus] 57

ornatum Andrewes, 1925, Asaphidion 242



ornatus W. J. MacLeay, 1888, Amblystomus [Acupalpus] 360

ornatus Sharp, 1882, Canthydrus 34

ornatus Heinz, 1976, Carabus 154

ornatus Schmidt‑Göbel, 1846, Pericalus 437

ornatus Herbst, 1784, Platambus [Dytiscus] 44


ornizensis Pierre Meyer, 1998, Carabus 155

orobicum De Monte, 1943, Bembidion 269

oromii Machado, 1987, Canarobius 345

orphaeus Pawowski, 1973, Trechus 337

orpheus LaFerté‑Sénectère, 1853, Amblystomus [Acupalpus] 360

orpheus Dieck, 1869, Geotrechus [Anophthalmus] 310

orpheus Dieck, 1870, Geotrechus [Anophthalmus] 310

orphnaeus Andrewes, 1935, Elaphropus [Tachys] 273

orphniopterus G. Hauser, 1920, Carabus [Coptolabrus] 156

orsinii A. Villa & G. B. Villa, 1838, Nebria 92

orsinii Dejean, 1831, Zabrus 571

ortali Vondel, 1991, Haliplus 32

ortali Wewalka, 1984, Hydroporus 63

orthapicalis Belousov & Kabak, 1998, Trechus 337

orthogenys Uéno & Saito, 1991, Jujiroa 459

orthogona Bremi‑Wolf, 1856, Cicindela 108

orthogonius Chaudoir, 1868, Molops 474



orthogrammus Sharp, 1882, Hygrotus [Coelambus] 72

orthomus Chaudoir, 1855, Scarites 234

ortizi Español, 1970, Trechus 337

ortunoi Arribas, 1992, Platyderus 523

oruzganensis Casale & Vereschagina, 1986, Pseudotaphoxenus 541

osakaense Jedlika, 1951, Bembidion 243

osakaensis Nakane, 1963, Brachinus 216

osakana Nakane, Ohkura & Uéno, 1955, Coptodera [Coptoderina] 436

osawai Imura, Zhou & Su, 1999, Carabus 192

oscari Apetz, 1854, Orectogyrus 29

oscensis Jeanne, 1985, Trechus 340

oschanini Semenov, 1897, Carabus 153

oschanini Tschitschérine, 1896, Cymindis 416

osculatii Osculati, 1844, Carabus 169

oscuratus Mateu, 1958, Brachinus 212

oscus A. Franzini & G. Franzini, 1984, Duvalius 305

osellai Breuning & Ruspoli, 1970, Carabus 160

osellai F. Battoni & Vigna Taglianti, 1994, Trechus 337

oseticus Belousov, 1998, Troglocimmerites 344



osiridis Peyron, 1856, Platytarus [Cymindis] 410

osismius Lapouge, 1922, Carabus 166

osmanilis Apfelbeck, 1904, Molops 473

osmanilis K. Daniel & J. Daniel, 1902, Trechus 337

osogovensis B. V. Guéorguiev, 1997, Molops 474

ossensis Maan, 1939, Zabrus 570

osseticus M. F. Adams, 1817, Carabus 200

osseticus Zamotajlov, 1992, Deltomerus 283

osseticus Fedorenko, 1993, Dyschirius 228



osseticus Reitter, 1909, Leistus 82

osseticus Kirschenhofer, 1987, Pterostichus [HN, repl. in vol. 2] 505

osseticus Poppius, 1908, Pterostichus 496

osseticus Jeannel, 1927, Trechus 336

ostariensis Born, 1912, Carabus 179

ostrinus Andrewes, 1924, Chlaenius 349



oszailensis Bedel, 1876, Typhlotrechus [Trechus] 344

otcharensis Kurnakov, 1972, Carabus 200

otingoensis Deuve, 1993, Carabus 194

otini Antoine, 1938, Amara 555

otini Antoine, 1963, Brachinus 212

otini Antoine, 1959, Bradycellus 401



otini Antoine, 1939, Carabus 152

otini Antoine, 1939, Cymindis 413

otini Antoine, 1953, Dyschirius 227



otini Antoine, 1941, Harpalus 373

otini Antoine, 1944, Paussus 210

otini Antoine, 1941, Platyderus 523

otini Guignot, 1941, Stictotarsus [Potamonectes] 69

otini Antoine, 1944, Tachys 276

otiosus Obenberger, 1917, Neotrechus [Anophthalmus] 316

otthi Gistel, 1838, Calomera [Cicindela] 101

ottofischeri B. Bodemeyer, 1928, Carabus [Phricocarabus] 179

ottomanus Reitter, 1896, Carabus 131

ottomanus Schweiger, 1970, Leistus 84

ottomanus Schweiger, 1968, Porotachys 275

ottomanus Apfelbeck, 1908, Pterostichus 508

ottonis Csiki, 1909, Carabus 166

ottonis Reitter, 1905, Neotrechus [Anophthalmus] 315

otuboi Habu, 1944, Morimotoides [Trephionus] 460

oudoti G. Hauser, 1913, Carabus [Coptolabrus] 144

ougsburgeri Heer, 1837, Amara 556

outereloi Novoa, 1978, Typhlocharis 241



ouzoumensis A. Gaudin, 1947, Hydraphaenops 311

ovale Motschulsky, 1844, Bembidion [Peryphus] 244



ovale A. Morawitz, 1886, Calosoma [Callisthenes] 119

ovaliceps Baehr, 1998, Gunvorita 573

ovaliceps Bates, 1878, Platynus [Colpodes] 465

ovalipenne Schuler, 1956, Bembidion [Peryphus] 259

ovalipenne Solsky, 1874, Bembidion 268



ovalipennis Jedlika, 1957, Amara 560

ovalipennis Ganglbauer, 1900, Binaghites [Scotodipnus] 239

ovalipennis Jaeger, 1996, Bradycellus 402

ovalipennis Schauberger, 1932, Parophonus [Tachyophonus] 393



ovalipennis Suenson, 1957, Trechiama 322

ovalis Duftschmid, 1812, Abax [Carabus] [NP] 470

ovalis J. R. Sahlberg, 1875, Agabus [Gaurodytes] 38

ovalis Mulsant & Rey, 1861, Amara 548

ovalis Dejean, 1831, Calathus 525

ovalis Sharp, 1884, Haliplus 32



ovalis Motschulsky, 1844, Harpalus 380

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