Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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ovalis Reiche, 1861, Harpalus 370

ovalis Thunberg, 1784, Hygrotus [Dytiscus] 72

ovalis Illiger, 1798, Hyphydrus [Dytiscus] 74

ovalis Gschwendtner, 1934, Ilybius 43

ovalis Gschwendtner, 1936, Lacconectus 48

ovalis Breit, 1914, Notiophilus 97



ovalis Marsham, 1802, Porhydrus [Dytiscus] 68

ovalis Motschulsky, 1844, Pseudotaphoxenus [Sphodrus] 541

ovalis Gschwendtner, 1936, Rhantus 46

ovalis Fairmaire, 1859, Zabrus 571

ovata Fabricius, 1792, Amara [Carabus] 550

ovata Motschulsky, 1844, Amara 549

ovata Motschulsky, 1851, Tachyura [Lopha] 278



ovatoides Baliani, 1943, Amara 548

ovatus Belousov, 1998, Cimmerites 297



ovatus Sturm, 1835, Deronectes [Hydroporus] 58

ovatus W. J. MacLeay, 1871, Elaphropus [Bembidium] 273

ovatus Chaudoir, 1879, Haplogaster 232



ovatus Chaudoir, 1844, Harpalus [NO] 372

ovatus Linné, 1761, Hyphydrus [Dytiscus] 74



ovatus Hochhuth, 1871, Ilybius 43

ovatus Motschulsky, 1866, Percus 478

ovatus Putzeys, 1846, Trechus 337

ovicollis Putzeys, 1873, Dyschirius 224

ovicollis Solsky, 1874, Dyschirius 224

ovicollis Reitter, 1884, Tapinopterus [Steropus] 519

ovicollis Uéno, 1999, Toshiaphaenops 321

oviformis Breuning, 1943, Carabus [Eucarabus] 170

ovipenne Chaudoir, 1846, Bembidion 263

ovipennis Motschulsky, 1859, Amara [Leirus] 560

ovipennis Fairmaire, 1888, Andrewesius [Dyscolus] 456

ovipennis Motschulsky, 1864, Brachinus 212

ovipennis Chaudoir, 1846, Clivina 219

ovipennis Bates, 1883, Cylindera [Cicindela] 108

ovipennis Motschulsky, 1844, Cymindis 414

ovipennis Putzeys, 1866, Dyschirius 224

ovipennis Jeannel, 1941, Geotrechus 310

ovipennis Chaudoir, 1867, Leistus 83

ovipennis Semenov, 1889, Loricera 98



ovipennis Chaudoir, 1847, Molops 473

ovipennis Kraatz, 1875, Molops 473

ovipennis Motschulsky, 1844, Ophonus 390

ovipennis Chaudoir, 1850, Patrobus 285

ovipennis Casale & Vereschagina, 1986, Pseudotaphoxenus 541

ovipennis Sciaky, 1994, Straneostichus 518

ovipennis Lewis, 1896, Trechiama [Trechus] 322

ovipennis Chaudoir, 1844, Zabrus 571

ovitensis Perrault, 1974, Leistus 82

ovoidea Morvan, 1997, Lepcha 459

ovoideum Marggi & Huber, 1999, Bembidion 251

ovoideus Crotch, 1873, Agabus [Gaurodytes] 35

ovoideus Régimbart, 1895, Bidessus 55



ovoideus P. Machard, 1994, Carabus 193

ovoideus Sturm, 1824, Pterostichus [Platysma] 508

ovoideus Uéno, 1970, Trechiama 323

ovtchinnikovi Gottwald, 1987, Carabus 145

ovtchinnikovi Kabak, 1999, Cymindis 416

ovtshinnikovi Kabak, 1993, Amara [Cribramara] 558

ovtshinnikovi Belousov, 1992, Duvalius 305

ovtshinnikovi Kataev, 1990, Harpalus 380

ovtshinnikovi Kabak, 1994, Poecilus 485

ovtshinnikovi Belousov & Kabak, 1992, Trechus 337



ovula Poppius, 1908, Amara 550

ovuloides Jedlika, 1953, Amara 551

ovulum Netolitzky, 1910, Bembidion 267

oxiana Tschitschérine, 1898, Amara 561

oxiana Kabak, 1997, Cymindis 415

oxianus Glasunov, 1913, Mnuphorus 358

oxuroides Schaum, 1862, Carabus 148



oxycancellatus A. Fleischer, 1898, Carabus 196

oxydatum Netolitzky, 1939, Bembidion 265

oxyglymma Bates, 1883, Bembidion 271



oxygonum A. Morawitz, 1886, Calosoma [Callisthenes] 119

oxygonus Chaudoir, 1850, Tschitscherinellus [Ditomus] 367



oxygonus Chaudoir, 1846, Zabrus [Eutroctes] 569

oxyptera K. Daniel & J. Daniel, 1904, Nebria 90



ozaljensis J. Müller, 1913, Typhlotrechus [Trechus] 344

ozegaharanus Uéno, 1954, Trechus 325

pacatus Uéno, 1994, Trechiama 323

paccatus Motschulsky, 1850, Phobophorus 406

pacei Magrini & Vanni, 1993, Anillus 238

pacensis Zaballos & Jeanne, 1987, Typhlocharis 241

pachinus Bates, 1883, Pterostichus 513

pacholei Sokolá, 1911, Carabus 136



pachtoun Ledoux, 1975, Carabus [Procrustes] 155

pachycerus Chaudoir, 1880, Distichus 231

pachycerus Apfelbeck, 1918, Trechus 337

pachypterus Imura, 1999, Carabus 183

pachys Uéno, 1970, Stygiotrechus 320

pacificatorius Habu, 1954, Trichotichnus 397

pacificus Aubé, 1838, Hydaticus 54

pacificus Bates, 1873, Scarites 232

pacificus Uéno, 1978, Thalassoduvalius 321

pacistanus Guignot, 1959, Hyphoporus 73

paciuchensis Monguzzi, 1995, Anophthalmus 292

padana Schaum, 1856, Cylindera [Cicindela] 110

padewiethi Csiki, 1906, Carabus 151

paediscum Bates, 1883, Bembidion 244

paegunsanus Imura, 1997, Carabus 183

paektusanensis Deuve, 1994, Carabus 134



paeninsularis Deuve, 1992, Carabus 183

paganetti Magrini & Pavesi, 1998, Rhegmatobius 240

paganettianus Scholz, 1923, Hydroporus 63

paganettii Netolitzky, 1914, Bembidion 264



paganettii Schweiger, 1969, Bradycellus 24, 400

paganettii Born, 1905, Carabus [Tomocarabus] 197

paganettii Flach, 1907, Harpalus 372

paganettii Ganglbauer, 1897 (as 1896), Neotrechus [Trechus] 315

paganettii Breit, 1923, Tapinopterus [Pterostichus] 519

paganettii J. Müller, 1911, Winklerites [Scotodipnus] 240

paghmana Jedlika, 1968, Cymindis 412

pagmanicum Fassati, 1959, Bembidion 255

pahalgamensis Ledoux, 1984, Ledouxius [Penetretus] 284

paiafa White, 1845, Carabus 188



pailherensis Jeanne, 1972, Carabus 139

paivana Wollaston, 1860, Cymindis [Tarus] 414

pakistana Jedlika, 1963, Amara 566

pakistana Jedlika, 1963, Mastax 217

pakistanensis Heinz & Ledoux, 1987, Ledouxius [Penetretus] [DA] 284

pakistanensis Morvan, 1996, Platynus [Batenus] 463

pakistanica Jedlika, 1963, Tachyura [Tachys] 280

pakistanus Jedlika, 1964, Anomotarus [Cymindis] 408

pakistanus Jedlika, 1964, Harpalus 386

pakistanus Jedlika, 1962, Laemostenus [Pterostichus] 538



pakistanus Straneo, 1982, Pterostichus 510

pakistanus Casale, 1982, Sphodropsis 543

palaciosi Andrewes, 1932, Bembidion 254

palae C. Müller, 1900, Pterostichus 488

palaestina Putzeys, 1870, Amara 555


palaestinensis Fery & Hosseinie, 1999, Deronectes 58

palaestinus A. Zimmermann, 1934, Agabus [Gaurodytes] 39

palaestinus Reiche & Saulcy, 1855, Chlaenius 349

palaestinus Fery & Hosseinie, 1998, Deronectes 58

palaestinus Jedlika, 1958, Harpalus 374

palaestinus Piochard de la Brûlerie, 1873, Oedesis [Ditomus] 366



palaestinus Baudi di Selve, 1894, Stictotarsus [Deronectes] 69

palanda Jedlika, 1957, Amara 547

palei Fourès, 1962, Geotrechus 310

palekhai E. Berlov & O. Berlov, 1996, Pterostichus 491

paleki Kult, 1949, Dyschirius 230



palesius Sirguey, 1931, Carabus 140

palgongsanensis Kwon & Lee, 1984, Carabus 183

palgongsanus Nemoto, 1988, Pterostichus 497



palicari Laporte, 1834, Brachinus 214

pallasii Fischer von Waldheim, 1824, Cicindela 107

pallens Poppius, 1905, Agabus 37



pallens Sturm, 1825, Amara [NO] 557

pallens Lapouge, 1925, Carabus 140

pallens Fowler, 1887, Haliplus 30

pallens A. Zimmermann, 1915, Hydroporus 62

pallens Aubé, 1838, Hygrotus [Hydroporus] 72

pallens Fabricius, 1775, Plochionus [ 244 Carabus] 409

pallens Andrewes, 1935, Sirdenus 288

pallens Belousov & Kabak, 1994, Trechus 337

pallescens Jedlika, 1963, Cymindis 414

pallescens Seidlitz, 1887, Hydroporus 64

pallescens Uéno, 1960, Rakantrechus 319

pallescens Bates, 1873, Tachys [DA] 276

pallescens L. Redtenbacher, 1849, Trechus 337

palliardii Gradl, 1881, Carabus 166

palliata Fischer von Waldheim, 1823, Cymindis 412

pallida Ménétriés, 1849, Amara [Bradytus] 551

pallida Reiche, 1855, Cymindis 411

pallida Bedel, 1898, Pogonopsis 286



pallida Dupuis, 1913, Tachyura [Tachys] 277

pallida Klug, 1832, Trichis 439

pallidicolor Mandl, 1973, Psammodromius [Microdaccus] 426

pallidicorne J. Müller, 1921, Bembidion 259

pallidicornis Ballion, 1871, Chlaenius 352

pallidicornis Fairmaire, 1891, Dicranoncus 458

pallidicornis Jeanne, 1990, Tachyura 21, 279

pallidior Bates, 1892, Apotomus 237

pallidior Pic, 1901, Microlestes [Blechrus] 424

pallidior Uéno, 1981, Trechiama 323



pallidipenne Dejean, 1831, Bembidion 253

pallidipenne Illiger, 1802, Bembidion [Elaphrus] 243



pallidipennis Fassati, 1949, Acupalpus 399

pallidipennis A. I. Jakovlev, 1897, Agabus 38

pallidipennis Laporte, 1835, Agabus [Colymbetes] 40

pallidipennis A. Morawitz, 1862, Harpalus 380



pallidipennis Bänninger, 1928, Nebria 91

pallidipennis Dejean, 1828, Pogonus 287



pallidipennis Schaum, 1857, Trechus 341

pallidipes Reitter, 1919, Brachinus 216



pallidipes Breit, 1914, Nebria 94

pallidipes Bedel, 1913, Syntomus [Metabletus] 433

pallidiveste Carret, 1905, Bembidion 264

pallidula Motschulsky, 1844, Amara [Bradytus] 554

pallidula Chaudoir, 1846, Cymindis 415

pallidulus Aubé, 1850, Hygrotus [Hydroporus] 72

pallidulus A. Zimmermann, 1930, Laccophilus 76

pallidulus Chaudoir, 1843, Oxypselaphus 462

pallidulus Habu, 1978, Synuchus 546



pallidulus Antoine, 1944, Tachys 21, 276

pallidulus Ménétriés, 1832, Tachys 275

pallidulus Ganglbauer, 1891, Trechus 337

pallidum Jedlika, 1955, Callidagonum [Callidula] 457

pallidus Stephens, 1828, Bradycellus [Trechus] 401

pallidus Ménétriés, 1832, Cybister [Dytiscus] 51

pallidus Dejean, 1829, Dicheirotrichus [Harpalus] 402



pallidus J. R. Sahlberg, 1903, Haliplus 33

pallidus Semenov, 1904, Haliplus 32

pallidus Heer, 1837, Nebrioporus [Hydroporus] 66

pallidus Motschulsky, 1844, Stenolophus 406

pallidus Reitter, 1884, Tachys 276

pallidus Reitter, 1894, Trechoblemus 324

pallidus Sturm, 1825, Trechus 340

pallipes Bruneau de Miré, 1949, Apotomus 237

pallipes Duftschmid, 1812, Asaphidion [Elaphrus] 242



pallipes Dejean, 1829, Bradycellus [Acupalpus] 401

pallipes Stephens, 1828, Bradycellus [Trechus] 401

pallipes Andrewes, 1931, Calleida 408

pallipes Gebler, 1823, Chlaenius [Epomis] 352

pallipes Chaudoir, 1873, Cymindis 417

pallipes Chaudoir, 1850, Drypta 358

pallipes Motschulsky, 1844, Harpalus 384

pallipes Panzer, 1803, Leistus [Manticora] 84

pallipes Fabricius, 1801, Paranchus [Carabus] 462

pallipes Dejean, 1825, Syntomus [Dromius] 433



pallipes Andrewes, 1934, Synuchus [Calathus] 546

pallitibia Fairmaire, 1901, Clivina 220

pallorus Kopecký, nom. nov., Tachys 21, 276

pallulatus Cavazzuti, 1998, Carabus 127

palmata Motschulsky, 1844, Calomera [Cicindela] 100

palmeni J. R. Sahlberg, 1900, Bembidion 265

palmensis Wollaston, 1864, Olisthopus 461

palmensis Machado, 1992, Philorhizus 425

palmensis Mateu, 1956, Trechus 330

palmi Machado, 1992, Metadromius 421

palmyrus Görgner, 1999, Carabus 188

palpale Motschulsky, 1850, Bembidion [Metallina] 252

palpalis Dokhtouroff, 1888, Cicindela 107



palpalis Dejean, 1831, Trechus 338

palpalis Duftschmid, 1812, Trechus [Carabus] 340

paludicola Reitter, 1884, Acupalpus 399



paludicola J. Müller, 1903, Carabus 136

paludicola Gistel, 1857, Syntomus [Metabletus] [DA] 433

paludis Géhin, 1885, Carabus 154

paludosa L. Dufour, 1820, Cylindera [Cicindela] 110



paludosum Panzer, 1794, Bembidion [Elaphrus] 247

paludosus Fabricius, 1801, Agabus [Dytiscus] 40



paludosus Ahrens, 1830, Dyschirius [Clivina] 224

paludosus Olivier, 1790, Elaphrus 207

paludosus R. F. Sahlberg, 1844, Pterostichus [Feronia] 493

paludosus Gyllenhal, 1810, Trechus [Bembidium] 340

palumbinus Lutshnik, 1922, Harpalus 374

palumboi Ragusa, 1887, Bembidion 252



palustris Baudi di Selve, 1864, Amara 551

palustris Gistel, 1857, Amara 547

palustris Dejean, 1826, Carabus 136

palustris Dejean, 1829, Carabus 154

palustris Beuthin, 1896, Cicindela 103

palustris Motschulsky, 1840, Cicindela 104

palustris Linné, 1761, Hydroporus [Dytiscus] 63

palustris Wallis, 1926, Ilybius [Agabus] 43

palustris E. Jacquet, 1887, Lebia 427

palustris Duftschmid, 1812, Notiophilus [Elaphrus] 97

palustris Saulcy, 1870, Reicheia 222

palustris Reitter, 1894, Tachys [DA] 276

pamensis Deuve, 1992, Cychrus 205

pamirense Bates, 1878, Bembidion 261

pamirensis Bates, 1878, Amara [Curtonotus] 559

pamirensis Reinig, 1932, Cymindis 414

pamirensis Belousov & Kabak, 1996, Trechus 337


pamiricola Lutshnik, 1930, Amara 551

pamiricola Lutshnik, 1930, Bembidion 244



pamiricus Guignot, 1955, Agabus 41

pamphylicus Jeanne, 1996, Trechus 337

panda Deuve, 1988, Carabus 175

pandellei Géhin, 1882, Carabus 139

pandellei Linder, 1859, Hydraphaenops [Anophthalmus] 311

pandellei Saulcy, 1867, Hypotyphlus [Scotodipnus] 239

pandellei Kraatz, 1877, Trechus 340

pandellei Putzeys, 1870, Trechus 340

panderi Fischer von Waldheim, 1820, Calosoma [Callisthenes] 119

pandora Holdhaus, 1924, Typhloreicheia [Reicheia] 223



pandurus Bates, 1873, Carabus 148

pangi Deuve & Tian, 1999, Carabus 126



pangoides Reitter, 1913, Ophonus 388

pannonica Mandl, 1936, Cicindela 107



pannonicus Csiki, 1906, Carabus 173

panormitana Ragusa, 1906, Calomera [Cicindela] 101

panousei Antoine, 1956, Tachyura 278

panshiri Ledoux & Roux, 1997, Nebria 91

pantanellii A. Fiori, 1903, Poecilus [Metapedius] 481

panteli Ganglbauer, 1915, Zabrus 569

pantherinus Reclaire & van der Wiel, 1948, Nebrioporus [Deronectes] 67

pantomus Jedlika, 1958, Pterostichus 508



panurgus Jeannel, 1946, Tachys [Eotachys] 275

panzeri Dejean, 1829, Carabus 164

panzeri Panzer, 1803, Pterostichus [Carabus] 508

paolii Binaghi, 1940, Trechus 342

papasoffi Mandl, 1942, Duvalius [Paraduvalius] 308

paphius L. Redtenbacher, 1843, Carabus 169

paphlagonicus Maan, 1935, Calathus 527

paphlagonicus Maan, 1940, Trechus 337

paphlagoniensis Schweiger, 1966, Amara [Bradytus] 553

paphlagoniensis Breuning, 1954, Carabus 131

papingutica Horvatovich, 1974, Nebria 84

pappianus Mandl, 1980, Carabus 133

papukensis Koa, 1900, Carabus 152

paracenthesis Motschulsky, 1839, Metablus [Dromius] 431

paracomplanatum Nitzu, 1995, Bembidion 246

paradeuvei Ledoux & Roux, 1998, Archastes 79

paradiruptus Imura, 1999, Carabus 182

paradisiacus Kirschenhofer, 1990, Platynus [Agonum] 465



paradoxa Sciaky & Pavesi, 1994, Nebria 95

paradoxus Stark, 1890, Carabus 132

paradoxus Apfelbeck, 1908, Deltomerus 283



paradoxus Joseph, 1870, Laemostenus [Sphodrus] 535

paradoxus Putzeys, 1875, Miscelus 436

paradoxus Uéno, 1981, Paragonotrechus 318

paradoxus Putzeys, 1868, Syleter [Ardistomis] 221

paradoxus K. Daniel & J. Daniel, 1898, Trechus 329

paragraphus Kolenati, 1845, Stenolophus 405

paralaterale De Monte, 1953, Bembidion 252



parallela Chaudoir, 1842, Amara [Leirus] 553

parallela Lesne, 1896, Clivina 220

parallela Habu, 1975, Sugimotoa 439

parallela Motschulsky, 1851, Tachyta [Tachys] 277

parallela Holdhaus, 1924, Typhloreicheia [Reicheia] 223



parallelepipeda Motschulsky, 1866, Nebria 91

parallelepipedus Dejean, 1828, Abax [Feronia] 470

parallelepipedus Piller & Mitterpacher, 1783, Abax [Carabus] [NP] 470



parallelicollis Joseph, 1870, Laemostenus [Sphodrus] 535

parallelipenne Chaudoir, 1850, Bembidion 250



parallelipennis Desbrochers des Loges, 1871, Agabus 38

parallelipennis Chaudoir, 1868, Pterostichus [Feronia] 503

parallelipipedus Marsham, 1802, Pogonus [Carabus] 287

parallellogrammus Ahrens, 1812, Hygrotus [Dytiscus] 72

parallelocollis Reiche & Saulcy, 1855, Laemostenus [Pristonychus] 536

parallelogrammus Motschulsky, 1858, Oodes 445

parallelogrammus Motschulsky, 1844, Scarites 234

paralleloides Kirschenhofer, 1987, Pterostichus 505



parallelum Motschulsky, 1844, Calosoma 120

parallelus Duftschmid, 1812, Abax [Carabus] 470

parallelus Coiffait, 1955, Aphoenops 295

parallelus Belousov, 1998, Apocimmerites 295



parallelus Faldermann, 1836, Carabus 136

parallelus Fischer von Waldheim, 1844, Carabus 199

parallelus Kraatz, 1878, Carabus 151

parallelus Kraatz, 1878, Carabus 173

parallelus Sirguey, 1931, Carabus [HN] 165

parallelus Schmidt‑Göbel, 1846, Celaenephes 410

parallelus A. Zimmermann, 1920, Copelatus 48

parallelus Reitter, 1894, Dicheirotrichus [Trichocellus] 403

parallelus Motschulsky, 1844, Dyschirius 230



parallelus Chaudoir, 1869, Eurygnathus 441

parallelus Babington, 1836, Haliplus 32

parallelus Sharp, 1869, Hydroporus 62

parallelus Aubé, 1838, Hygrotus [Hydroporus] (as parallellus) 72

parallelus Mlyná, 1977, Molops 474



parallelus G. R Waterhouse, 1833, Notiophilus 97

parallelus Motschulsky, 1858, Oodes 445

parallelus Dejean, 1829, Ophonus [Harpalus] 390

parallelus Chaudoir, 1854, Oxycentrus 392



parallelus Chaudoir, 1871, Pogonus 287

parallelus Dejean, 1828, Pseudotaphoxenus [Sphodrus] 540

parallelus Ballion, 1871, Syntomus [Metabletus] 433



parallelus Motschulsky, 1850, Tachys 275

parallelus Reitter, 1894, Tachys 276

paraoxuroides Baba, 1939, Carabus 148

parapleurus Marsham, 1802, Hydaticus [Dytiscus] 54

pararboreus Ishikawa, 1992, Carabus [Leptocarabus] 159

parasimilis G. E. Ball, 1962, Pterostichus 491


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