Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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paraspinosus Korell & Kleinfeld, 1999, Cychrus 203



parcepunctatus Reitter, 1902, Ophonus 391

parcesparsa Antoine, 1939, Cymindis 413

parcestriatus Scheibel, 1938, Typhlotrechus 345

pardoi Mateu, 1957, Orthomus 477

pardoi Mateu, 1954, Singilis 438

parens Fairmaire, 1889, Diacanthostylus [Colpodes] 458

parens Tschitschérine, 1897, Pterostichus [Feronia] 493

parepum Jedlika, 1933, Bembidion 271

parexilis Nakane, 1961, Carabus 159

parilis Dvoák, 1993, Odotoncarus [Carterus] 366

paris Breuning, 1932, Carabus 128



parisiensis Gautier des Cottes, 1867, Calathus 527

parisii Gridelli, 1939, Graptodytes 59

parisii Schatzmayr, 1940, Oreonebria [Nebria] 95

parkapansis Morvan, 1982, Carabus 162

parkeri Baliani, 1934, Amara 566



parkeri Andrewes, 1935, Bembidion 261

parkeri G. Ochs, 1925, Orectochilus 29

parnassicola Kraatz, 1875, Molops 475

parnassicus Breit, 1923, Caecoparvus [Scotodipnus] 239

parnassicus Kraatz‑Koschlau, 1884, Carabus 185



parnassium L. Miller, 1884, Bembidion 257

parnassius Schaum, 1859, Pterostichus 504

paroecus J. Frivaldszky, 1865, Duvalius [Anophthalmus] 300

paroecus J. Frivaldszky, 1878, Duvalius [Anophthalmus] (as valid sp.) 300

paroecus Csiki, 1915, Endynomena [Thyreopterus] 437

paromoeus Guignot, 1954, Hyphydrus 74

paropamisicus Casale, 1982, Pseudotaphoxenus [Taphoxenus] 541



parowanum Casey, 1918, Bembidion 262

parowanus Casey, 1924, Harpalus 379

parreyssi Palliardi, 1825, Carabus 151

parreyssi Chaudoir, 1850, Mastax 217

parreyssi Kraatz, 1875, Molops 473



parreyssii Chaudoir, 1843, Nebria 90

parryi Putzeys, 1861, Clivina 219

parsorum Netolitzky, 1934, Bembidion 257



parthicus Lapouge, 1925, Carabus 193

parumpunctatum Fabricius, 1792, Agonum [Carabus] 451

parumpunctatus Semenov, 1898, Carabus 134

parumpunctatus Zamotajlov, 1990, Deltomerus 283



parumpunctatus W. Kolbe, 1924, Harpalus 372

parumpunctatus O. Schneider, 1903, Laccophilus 77

parumpunctatus Germar, 1824, Pterostichus 512

parumstriatus Fairmaire, 1872, Platyderus [Sphodrus] 521

parvicatenatus Mandl, 1955, Carabus 178



parviceps Lapouge, 1925, Carabus 123

parviceps Van Dyke, 1925, Elaphrus 207

parviceps Fairmaire, 1859, Laemostenus [Sphodrus] 532

parviceps Mlyná, 1977, Molops 474

parviceps Poppius, 1906, Pterostichus [Cryobius] 492

parvicolle J. R. Sahlberg, 1880, Bembidion 250

parvicollis Gebler, 1833, Amara 552



parvicollis Kraatz, 1878, Carabus 136

parvicollis Uéno, 1969, Coreoblemus 297

parvicollis Schaum, 1864, Deronectes [Hydroporus] 58

parvicollis Tschitschérine, 1900, Dicheirotrichus [Trichocellus] 402

parvicollis Uéno & Saito, 1991, Jujiroa 459

parvicollis Chaudoir, 1869, Leistus 18, 83



parvicollis Dodero, 1916, Microtyphlus [Scotodipnus] 240

parvicollis Motschulsky, 1859, Nebria 89

parvicollis Winkler, 1926, Neotrechus 315

parvicollis Uéno, 1987, Oroblemus 316

parvicollis Wollaston, 1865, Philorhizus [Dromius] 426

parvicollis Van de Poll, 1889, Physodera 438

parvicollis Sciaky & Wrase, 1997, Pterostichus 500

parvicollis Uéno, 1999, Shuaphaenops 320

parvicornis Deuve, 1990, Carabus 134

parvielongatus Deuve, 1994, Carabus 127

parvilimbus G. Ochs, 1925, Orectochilus 29

parvimaculata Fowler, 1912, Lophyra [Cicindela] 113

parvipes Cavazzuti, 1996, Carabus 124

parvithorax Cavazzuti & Ratti, 1999, Carabus 126

parvula Ledoux & Roux, 1995, Nebria 95

parvula Dejean, 1831, Tachyura [Bembidium] 279

parvulus Sturm, 1825, Acupalpus [Trechus] 399

parvulus Fresneda & Hernando, 1989, Agabus 40

parvulus Stark, 1894, Carabus 133

parvulus Ballion, 1878, Harpalus [Pangus] 380


parvulus Fabricius, 1792, Hygrotus [Dytiscus] 73

parvulus Aubé, 1838, Laccophilus 77



parvulus G. R. Waterhouse, 1833, Notiophilus 96

parvulus Herbst, 1784, Porhydrus [Dytiscus] 68

parvulus Semenov, 1889, Pseudotaphoxenus 542



parvulus Chaudoir, 1868, Pterostichus [Feronia] 489

parvulus Uéno, 1969, Stygiotrechus 320



parvus Géhin, 1885, Carabus 152

parvus Wiedemann, 1823, Distichus [Scarites] 231

parvus Uéno, 1980, Trechiama 323

pasanauricus Kirschenhofer, 1987, Pterostichus 505



paschkowensis G. Hauser, 1921, Carabus [Coptolabrus] 144

pascuorum Lapouge, 1908, Carabus 130

pascuorum Tschitschérine, 1890, Poecilus [Feronia] 484

pascuorum J. Daniel, 1903, Pterostichus 491

pasianax Fischer von Waldheim, 1828, Carabus 172

pasquinii Pomini, 1938, Oreonebria [Nebria] 95

pasquinii Cerruti, 1973, Speluncarius 516

pasquinii Vigna Taglianti, 1977, Troglocimmerites 344

passerinii Dejean, 1828, Percus [Feronia] 478

pastica Dejean, 1831, Amara 566

pasticus Germar, 1817, Acinopus [Carabus] 368

pastor Motschulsky, 1844, Harpalus 385



pastus Ménétriés, 1848, Harpalus 373

pasubiana K. Daniel & J. Daniel, 1898, Amara 563

pasubianum Weissmandl, 1936, Broscosoma 235

pasubianus Ganglbauer, 1901, Duvalius [Trechus] 301



pasubianus Jeannel, 1937, Laemostenus [Cryptotrichus] 536

pasubianus Jeannel, 1928, Orotrechus 317

pateri Puel, 1938, Brachinus 212

pateri Hauret, 1946, Gynandromorphus 362

pateri Puel, 1937, Poecilus 484

patinalis Bates, 1892, Loxoncus [Anoplogenius] 404

patrata Schiødte, 1841, Amara 553

patricia Duftschmid, 1812, Amara [Carabus] 566

patris Smetana, 1950, Pterostichus 491

patrizii Vigna Taglianti, 1999, Laemostenus 535

patroboides Lindroth, 1956, Synuchus 546

patroclus Breuning, 1933, Carabus 126

patruele Gemminger & Harold, 1868, Agonum [Platynus] 453

patruelis Dejean, 1829, Harpalus 379

patruelis Chaudoir, 1846, Nebria 86

patruelis L. Dufour, 1820, Percus [Broscus] 479



patruelis Chaudoir, 1846, Syntomus [Dromius] 432

patruelis Putzeys, 1847, Trechus 331

pauciseta C. G. Thomson, 1867, Poecilus [Feronia] 484

paulae Fagniez, 1924, Duvalius [Duvalites] 302



pauli Csiki, 1927, Carabus 139

paulinae Fagniez, 1922, Duvalius [Duvalites] 305

paulini Vuillefroy, 1868, Pterostichus [Feronia] 504

paulinoi Heyden, 1870, Bembidion 243

paulinoi Heyden, 1891, Stenolophus 406

paulmeyeri Fassati, 1958, Bembidion 261

paulo Wrase, 1995, Philorhizus 426

paulusi Kaláb, 1995, Carabus 175

paumashanica Hieke, 1997, Amara 554



pauper Schilsky, 1888, Agabus 38

pauper O. Schneider, 1903, Graptodytes [Hydroporus] 60

pauper Tschitschérine, 1897, Trichotichnus [Harpalus] 397

paupercula Knirsch, 1925, Duvaliopsis [Trechus] 298

paussoides Wasmann, 1904, Platyrhopalus 211

pauxillus Wollaston, 1864, Dyschirius 226

pavani C. Bari & R. Rossi, 1965, Allegrettia 290

pavani Straneo, 1988, Pterostichus [Haptotapinus] 496

paveli Csiki, 1899, Anophthalmus [Trechus] 293

pavesii Cavazzuti, 1992, Carabus 193

pavesii Sciaky, 1994, Leistus 81

pavesii Zamotajlov & Sciaky, 1996, Parapenetretus 284

pavidus Panzer, 1799, Paranchus [Carabus] 462

pavionis Jeannel, 1926, Duvalius 302

pavionis Meggiolaro, 1961, Orotrechus 317



pavlitscheki Born, 1907, Carabus 152

pavlovskii Kryzhanovskij, 1955, Calosoma [Callisthenes] 119

pavlovskii Jeannel, 1962, Trechus 337

pavruzum Morvan, 1996, Xestagonum 468

pawlowskianus Deuve, 1989, Carabus 183

pawlowskianus P. Moravec & Lompe, nom. nov., Trechus 22, 333

pawlowskii Farka & Fassati, 1999, Leistus 83

pawlowskii Shilenkov, 1983, Nebria 88

paykulli Erichson, 1837, Colymbetes 45

paykulli G. Ochs, 1927, Gyrinus 28

paykulli P. Rossi, 1792, Percus [Carabus] 478

paykullii Leach, 1817, Agabus 37

pazi Seidlitz, 1867, Nebria 95

pecignai Toribio, 1992, Trechus 337

pecirkai Jedlika, 1932, Chlaenius 353

pecirkai Jedlika, 1931, Colpoides 457

pecirkai Jedlika, 1946, Cymindis 416



pecirkai Jedlika, 1928, Harpalus 386

peckorum Deuve, 1992, Epaphiopsis [Epaphiama] 309

pecoudellus Deuve, 1998, Carabus 179

pecoudi Puel, 1925, Brachinus 213

pecoudi Colas, 1938, Calathus 528

pecoudi Jeannel, 1937, Duvalius 305



pecoudi Jeannel, 1928, Geotrechus 310

pecoudi A. Gaudin, 1938, Hydraphaenops [Aphoenops] 311

pecoudi Puel, 1925, Licinus 442

pecoudi Colas, 1957, Ocys 272

pecoudi Jeannel, 1943, Orthomus 476

pecoudi Mateu, 1954, Paraeutrichopus 521

pecoudi Antoine, 1922, Pterostichus [Steropus] 514

pecoudi Jeannel, 1947, Pterostichus [Haptoderus] [HN!] 489



pecoudi Puel, 1925, Pterostichus 512

pecoudi Colas & A. Gaudin, 1935, Trechus 337



pecoudi Jeannel, 1937, Trechus 336

pecoudi Antoine, 1934, Trymosternus 410

pecoudi Colas, 1942, Zabrus 572

pecoudianus Deuve, 1992, Carabus 147

pecoudianus A. Gaudin, 1947, Pterostichus [Lianoe] 498

pecoudianus Jeannel, 1952, Trechus 337

pectinatus Báguena Corella, 1935, Hydroglyphus [Bidessus] 56

pectoralis Dejean, 1825, Brachinus 214



pectoralis J. R. Sahlberg, 1880, Hydroporus 60

pectoralis Motschulsky, 1860, Hygrotus [Hydroporus] 72



pectoralis Gistel, 1857, Nebria 93

peculiaris Tschitschérine, 1894, Amara 561



peculiaris Tschitschérine, 1894, Poecilus [Feronia] 480

pedator Reitter, 1887, Carabus 179

pedemontana Vuillefroy, 1868, Oreonebria [Nebria] 96

pedemontanus Ganglbauer, 1891, Carabus 180



pedemontanus A. Fiori, 1904, Haliplus 30

pedemontanus Ganglbauer, 1891, Pterostichus 513

pederzanii Fery & Nilsson, 1993, Ilybius [Agabus] 43

pederzanii Toledo & Mazzoldi, 1996, Rhantus 46

pederzanii Angelini, 1973, Scarodytes 69

pedestre Motschulsky, 1844, Bembidion [Notaphus] 254



pedestris Putzeys, 1866, Amara [Curtonotus] 560

pegodi Jedlika, 1937, Chlaenius 353

peguensis Bates, 1892, Brachyodes [Oodes] 444



pehri Lapouge, 1913, Carabus 184

pehri Sokolá, 1907, Cychrus 203

peilingi Jedlika, 1937, Pterostichus 515

peipingensis Mandl, 1934, Calomera [Cicindela] 100

peiracavae Jeannel, 1926, Duvalius 305

peirolerii A. Villa & G. B. Villa, 1838, Carabus 184


peirolerii Bassi, 1834, Platynus 463

peirolerii Heer, 1837, Pterostichus 503

peisonis Ganglbauer, 1891, Pogonus 287



pekinensis Fairmaire, 1887, Carabus 136

pekinensis Kult, 1949, Dyschirius 229

pekinensis Vondel, 1992, Peltodytes 33

peksi Heinz & Pavesi, 1994, Calosoma 119

peksi Korell & Kleinfeld, 1993, Carabus 154

pektusanica Horvatovich, 1973, Nebria 89

pelagicus Wollaston, 1860, Nesacinopus [Harpalus] 388

pelates Guignot, 1953, Hyphydrus 74

pelelaensis J. Schmidt, 1997, Xestagonum 468



peletieri Laporte, 1835, Carabus 152

peletieri Laporte, 1840, Graphipterus 359

peletieri Lucas, 1848, Neolaphyra [Cicindela] 114

peleum Jedlika, 1933, Bembidion 244

peleus Schaufuss, 1861, Laemostenus [Sphodrus] 535

pelidnum Herbst, 1784, Agonum [Carabus] 453

pelionensis Breit, 1923, Tapinopterus [Pterostichus] 520

peliopterum Chaudoir, 1850, Bembidion 246

peliotes Habu, 1974, Negreum [Platynus] 460



pelissieri Darnaud, 1978, Carabus 195

pelissieri Pic, 1947, Lophyra [Cicindela] 113

pellegrinii Assmann, 1998, Pterostichus [Haptoderus] 490



pellucida Tschitschérine, 1894, Amara 562

pellucida Piochard de la Brûlerie, 1875, Cymindis 414

pellucidus Wollaston, 1854, Zargus 440

pelopis J. R. Sahlberg, 1908, Haliplus 32

peloponnesiacus Casale, 1982, Laemostenus 533

peltastes Andrewes, 1931, Brachinus 216

peltatus Panzer, 1796, Badister [Carabus] 441

peltatus Kolenati, 1845, Calathus 528

peltoides Dejean, 1826, Licinus 442

pelvicus Cavazzuti, 1973, Carabus 180

pelviger Kolenati, 1845, Pterostichus [Pseudomaseus] 511

penacollaradensis Dupré, 1991, Hydraphaenops 311

pendjabensis Guignot, 1954, Hydroglyphus [Guignotus] 56

peneckei Ganglbauer, 1897 (as 1896), Trechus 337

penelope Kleinfeld, 1997, Carabus 126

penglainus Huang, Hu & Sun, 1994, Harpalus 385



penialis Lapouge, 1913, Carabus 133

penibeticus Mateu & Colas, 1954, Orthomus 477

peninsularis Park & Kwon, 1996, Pterostichus 507

penitus Guignot, 1945, Hydroporus 65

pennatus Dejean, 1828, Pterostichus [Feronia] 500


pennboullek Morvan, 1999, Xestagonum 468

pennekum Morvan, 1995, Platynus [Colpodes] 465

pennina Binaghi, 1936, Alpiodytes [Reicheia] 221

pennina Gistel, 1838, Cicindela 107

pennina Jeannel, 1937, Nebria 93

penninum Netolitzky, 1918, Bembidion 246



penninus Ganglbauer, 1900, Binaghites [Scotodipnus] 239

pennisii Magrini & Vanni, 1984, Duvalius 305

pennisii Magrini, 1987, Nebria 93

pennisii Magrini & Vanni, 1989, Ocys 272

pennisii Magrini, 1984, Trechus 338

pentagrammus Schaum, 1864, Hydroglyphus [Hydroporus] 56

pentheri Apfelbeck, 1918, Calosoma 121

pentheri J. Müller, 1924, Laemostenus 534

pentheri Apfelbeck, 1904, Molops 473

pentheri Winkler, 1926, Neotrechus 316

pentheri Apfelbeck, 1918, Pterostichus 515

pentheri Ganglbauer, 1905, Zabrus 571



pentheroides Roubal, 1932, Calosoma 121

perabdita Antoine, 1941, Amara 566

peratrum De Monte, 1956, Bembidion 261

perauratus Reitter, 1896, Carabus 194

percontator Reitter, 1887, Pterostichus 505



percrenatus Reitter, 1896, Pterostichus 505

perditum Netolitzky, 1920, Bembidion 263



peregrina A. Morawitz, 1862, Amara 560

peregrina C. A. Dohrn, 1880, Galerita 359

peregrina Zaballos & Wrase, 1998, Typhlocharis 241



peregrinus Apfelbeck, 1904, Brachinus 213

peregrinus Cavazzuti & Ratti, 1998, Carabus 183

peregrinus Tschitschérine, 1898, Poecilus [Feronia] 480

perelegans Tschitschérine, 1898, Aristochroa 471



perelegans Jedlika, 1960, Platynus [Platynus] 464

perelloi Casale, 1979, Carabus 153



perezi Putzeys, 1867, Amara [Leiocnemis] 554

perezi Vuillefroy, 1868, Harpalus 379

perezii Martínez y Saez, 1873, Orthomus [Feronia] 477



perezii Crotch, 1869, Trechus 331

perfectus Schaufuss, 1882, Carabus 157

perfectus Andrewes, 1929, Dyschirius 225

perforata Motschulsky, 1850, Cymindis [Tarus] 415

perforata Bates, 1873, Parena [Bothynoptera] 434



perforatus Fischer von Waldheim, 1820, Carabus 182

perforatus Reiche & Saulcy, 1855, Dixus [Aristus] 365

perforatus Schaum, 1859, Haliplus 32

perforatus Sharp, 1882, Leiodytes [Bidessus] 57

pericallus L. Redtenbacher, 1867, Callistoides [Chlaenius] 348



pericaspius Guignot, 1956, Rhantus 46

periglabellus Huang, 1992, Harpalus 385

perignitus Reitter, 1896, Carabus 141

perigonoides Kasahara, 1994, Uenanthracus 406



peringueyi Kult, 1954, Dyschirius 225

perinii Tamanini, 1953, Orotrechus 317

perinii Holdhaus, 1950, Trechus 338

perisi Novoa, 1979, Pterostichus 508

perissus Uéno, 1983, Trechiama 323

perissus Andrewes, 1936, Trechus 338

peristerica Apfelbeck, 1901, Nebria 87

peristericus Roubal, 1930, Cychrus 205

peristericus J. Müller, 1914, Duvalius [Trechus] 300

peristericus Apfelbeck, 1901, Molops 473

peristericus Apfelbeck, 1901, Tapinopterus 520

peristericus Apfelbeck, 1904, Zabrus 571

perlaetus Reitter, 1900, Ophonus 389

perlamatus Straneo, 1989, Pterostichus 510



perlautus Gozis, 1912, Agabus 39

perlidaghensis Shilenkov, 1983, Nebria 85

perlidaghensis Reitter, 1902, Pterostichus 495

perlonga Heyden, 1885, Nebria 91

perlongus J. Müller, 1937, Dyschirius 225

perlucidus Schauberger, 1929, Harpalus 385

perlutus Bates, 1873, Elaphropus [Tachys] 273

perlutus Jedlika, 1938, Pterostichus 516



perminutus Reitter, 1898, Trechus 336

permoestum Puel, 1938, Agonum 451



pernigrum Reitter, 1897, Agonum 451

pernitidus Antoine, 1941, Platyderus 522

pernix Csiki, 1929, Amara 564

pernix Semenov, 1900, Bedeliolus 286

pernix Csiki, 1904, Carabus 152

perobscurus Deuve, 1980, Carabus 155

perobtecta Antoine, 1953, Amara [Percosia] 556

peronae Hopffgarten, 1885, Carabus 185

perplexa Dejean, 1828, Amara 549

perplexum Dejean, 1831, Bembidion 255

perplexus Schaum, 1857, Carabus 198

perplexus Lacordaire, 1835, Dytiscus 51

perplexus Sharp, 1882, Graphoderus 49



perplexus Gyllenhal, 1827, Harpalus 382

perplexus Schaum, 1847, Hydroporus [RN] [NO] [add, syn of lucasi Reiche] 62

perprasinum Reitter, 1897, Agonum 451

perpusillus Mateu & Deuve, 1979, Trechus 338

perpusillus Rottenberger, 1874, Winklerites [Microtyphlus] 240

perraulti Deuve, 1998, Amerizus 242

perraulti Sciaky, 1994, Leistus 81

perreaui Deuve, 1987, Cicindela 108

perreaui Deuve, 1985, Deltocolpodes [Agonum] 457

perreaui Deuve, 1988, Epaphiopsis [Epaphiama] 309

perreaui Perrault, 1986, Leistus 81

perreaui Deuve & Quéinnec, 1985, Trechus 338



perreti Born, 1907, Carabus 171

perrieri Fairmaire, 1901, Selina [Sphinctodera] 407

perrieri Jeannel, 1946, Tachys 275

perrinae Fery & Brancucci, 1997, Deronectes 58

perrinae Giordan, 1989, Duvalius 305

perrini Dejean, 1831, Carabus 199

perroti Guignot, 1950, Haliplus 32

perroti Jeannel, 1954, Trechiotes [Trechus] 324

persa Lapouge, 1925, Carabus 192

persana Dokhtouroff, 1885, Cicindela 105

persecretum Antoine, 1941, Agonum 453

persephone Andrewes, 1926, Bembidion 266

persianum Morvan, 1974, Calosoma [Callitropa] 119

persianus Roeschke, 1896, Carabus 164

persianus Stichel, 1923, Dixus [Ditomus] 365

persianus Jedlika, 1958, Harpalus 375

persianus Morvan, 1973, Sirdenus 288


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