Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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apiceplana Uéno, 1989, Epaphiopsis 309



apicerugosus Harald Lindberg, 1953, Calathus 528

apicesculptus Burgeon, 1935, Dyschirius 225

apicicornis Reitter, 1899, Laccophilus 76

apiculatiphallus Nemoto, 1988, Pterostichus 497



apiculatus Bates, 1892, Orthogonius 446

aplutenkoi O. Berlov & E. Berlov, 1996, Pterostichus 487

apoduvalipenis Salgado Costas & Ortuño, 1998, Trechus 326

apollo Zolotarev, 1913, Carabus [Plectes] 131

apollo Dvoák, 1993, Tschitscherinellus [Carterus] 367

aporus Puel, 1931, Anisodactylus 361

appendiculatus Wollaston, 1862, Calathus 526



appendiculatus Gautier des Cottes, 1869, Pterostichus [Feronia] 512

applanatus Uéno, 1980, Trechiama 321

approximans Fairmaire, 1888, Pogonus 286

approximatum Reiche & Saulcy, 1855, Agonum [Anchomenus] 451

appulsus Balkenohl, 1999, Trilophus 222



aprica Stephens, 1827, Cicindela 105

apricalis Cavazzuti, 1997, Carabus 126



apricans Motschulsky, 1850, Pogonus 287

apricaria Paykull, 1790, Amara [Carabus] 553



apricaria Gistel, 1838, Cylindera [Cicindela] 109

apristoides Rottenberg, 1870, Tachyura [Tachys] 278

aprutiana Bänninger, 1922, Nebria 92



aprutianus Gridelli, 1926, Pterostichus 495

apschuanus Rost, 1893, Carabus [Plectes] 200

aptera Kasahara, 1989, Cintaroa 446

apterum Netolitzky, 1930, Bembidion 255

apterus J. Müller, 1926, Bradycellus 401

apterus Holdhaus, 1904, Microlestes 422



apterus Bedel, 1899, Ophonus 388

aptinoides Wollaston, 1865, Dicrodontus [Polystichus] 573

aptinoides Tarnier, 1860, Pseudanchomenus [Anchomenus] 466


apuana Busi & Rivalta, 1980, Nebria 92

apuanus Dodero, 1917, Duvalius [Trechus] 301



apunctatus Puel, 1935, Harpalus 375

apusenicus P. Moravec, 1986, Trechus 326



aquatica Modeer, 1776, Orectochilus [Gyrinus] 29

aquaticum Panzer, 1796, Bembidion [Carabus] 270

aquaticus Linné, 1758, Notiophilus [Cicindela] 96



aquatile Illiger, 1798, Bembidion [Elaphrus] 270

aquatilis Bates, 1883, Carabus 160

aquila Coquerel, 1859, Orthomus [Feronia] 476

aquilarum Cavazzuti, 1997, Carabus 175

aquilo Andrewes, 1934, Pristosia [Calathus] 530

aquilo Andrewes, 1930, Trichotichnus 395

aquilonius Tschitschérine, 1904, Pterostichus [Platysma] 490

aquilum Andrewes, 1924, Bembidion 245

aquilus Jeannel, 1962, Trechus 326

aquitaniensis Mollard, 1996, Carabus 153

aquitanum Aubry, 1970, Bembidion [Trepanes] 248

arabiana Brouerius van Nidek, 1984, Cephalota [Cicindela] 102

arabica Hieke, 1983, Amara 555



arabica Dejean, 1831, Calomera [Cicindela] 99

arabica Fairmaire, 1896, Discoptera 356

arabica Mateu, 1986, Matabele 430

arabica Mateu, 1986, Merizomena [Glycia] 409



arabica Britton, 1948, Tachyura 280

arabicum Motschulsky, 1866, Calosoma [Caminara] 121

arabicus Mateu, 1986, Apristus 431

arabicus Sharp, 1882, Canthydrus 33

arabicus Régimbart, 1907, Dineutus [Dineutes] 26



arabicus Zaitzev, 1953, Haliplus 33

arabicus Guignot, 1951, Hydaticus 53

arabicus Mateu, 1979, Metadromius 421

arabicus Mateu, 1956, Microlestes 422

arabicus Sharp, 1882, Nebrioporus [Deronectes] 66

arabicus Raffray, 1885, Paussus 210

arabicus Arrow, 1901, Pheropsophus 217

arabicus Gestro, 1889, Triaenogenius 407



arabikensis Gottwald, 1985, Carabus 132

arachnopus G. Hauser, 1914, Carabus [Coptolabrus] 144

aradensis Kirschenhofer, 1984, Cymindis 411



aragonensis Lapouge, 1913, Carabus 141

aragonensis Colas, 1935, Pterostichus 498



aragonicus Ganglbauer, 1886, Carabus 167

aragonicus Jeannel, 1937, Laemostenus [Ceuthosphodrus] 533

aragonicus Jeanne, 1985, Platyderus 522



aragonicus Jeannel, 1942, Pterostichus [Haptoderus] 492

aragonicus Jeannel, 1930, Trechus 330

aralarensis Español & Mateu, 1945, Pterostichus [Haptoderus] 489

aralensis Glasunov, 1909, Poecilus [Platysma] 480



araneipes Lapouge, 1903, Carabus 184

aranensis Jeannel, 1921, Trechus 334



araratensis Kraatz, 1897, Carabus [Mimocarabus] 169

araratensis Casale, 1988, Laemostenus 538



araraticum Chaudoir, 1846, Calosoma 118

araraticus Mlyná, 1979, Harpalus 373

araraticus Kirschenhofer, 1987, Pterostichus 504

araschinica Reitter, 1892, Nebria 18, 85



arata Andrewes, 1925, Tachyura [Tachys] 278

arator Faldermann, 1836, Pterostichus [Abax] 495

aratus Andrewes, 1932, Apristus 431

araxellus Iablokoff‑Khnzorian, 1963, Elaphrus 207



araxicola Reitter, 1889, Calomera [Cicindela] 99

araxidis Iablokoff‑Khnzorian, 1976, Bembidion 246

araxidis Reitter, 1889, Zabrus 572

araxidis Iablokoff‑Khnzorian, 1972, Zuphium 574

arbensis Gridelli, 1944, Cylindera [Cicindela] 111

arbensis J. Müller, 1911, Dyschirius 224



arborensis Krausse, 1908, Carabus 161

arboreus Lewis, 1882, Carabus 159



arbustorum Gistel, 1857, Nebria 91

arcadicus J. Müller, 1935, Caecoparvus [Scotodipnus] 239

arcadicus Gistel, 1850, Carabus 138

arcadicus J. Müller, 1946, Tapinopterus 519

arcana Emetz, 1972, Cymindis 416

arcanoides Lorenz, 1998, Pterostichus [Aphaonus] 485

arcanus Semenov, 1898, Carabus 148

arcanus Kurnakov, 1962, Pterostichus 485

arcensis Herbst, 1784, Carabus 135

arcensis Ledoux & Roux, 1990, Nebria 85

archaeicus Uéno, 1996, Phreatodytes 34



archangelicum J. R. Sahlberg, 1875, Agonum [Anchomenus] 449

archangelicus Poppius, 1907, Pterostichus 506

archastoides Ledoux & Roux, 1997, Nebria 89

arcifer Znojko, 1928, Dyschirius 224

arctica Kryzhanovskij & Emetz, 1979, Cymindis 416

arctica Gyllenhal, 1810, Diacheila [Harpalus] 206

arctica Paykull, 1798, Miscodera [Scarites] 237

arctica Dejean, 1826, Nebria 88

arctica Dejean, 1826, Pelophila 98

arcticola Poppius, 1906, Amara 567



arcticollis Jeannel, 1928, Duvalius 307

arcticum Lindroth, 1963, Bembidion 265

arcticus Paykull, 1798, Agabus [Dytiscus] 35

arcticus Sparre‑Schneider, 1888, Carabus 166

arcticus Olivier, 1795, Dromius [Carabus] 420

arcticus Dejean, 1826, Elaphrus 207

arcticus C. G. Thomson, 1856, Hydroporus 63

arcticus J. R. Sahlberg, 1880, Pterostichus [Feronia] [HN] 490

arcticus Reiche, 1857, Pterostichus [Platysma] 487

arcuata Putzeys, 1865, Amara [Leiocnemis] 554



arcuata Kollar, 1836, Calomera [Cicindela] 99

arcuaticarinatus Kasahara, 1986, Pterostichus 511



arcuaticollis Motschulsky, 1860, Chlaenius 349

arcuaticollis Motschulsky, 1866, Pterostichus [Argutor] 486

arcuaticollis N. Ito, 1997, Stenolophus 405

arcuaticollis Motschulsky, 1860, Synuchus [Pristodactyla] 545

arcuatus Gautier des Cottes, 1870, Calathus 524



arcuatus Panzer, 1795, Graptodytes [Dytiscus] 59

arcuatus Tschitschérine, 1898, Harpalus 373

arcuatus Reiche, 1855, Phloeozetus [Lebia] 438

arcuatus Tschistchérine, 1897, Pterostichus [Feronia] 496

arcuatus Dejean, 1829, Tetragonoderus 357

arcuatus Jeannel, 1927, Trechus 334

arcuatus Jeannel, 1928, Typhlotrechus 344

ardoisi Puel, 1938, Agonum 449

ardosiacus Lutshnik, 1922, Ophonus [Harpalus] 391

ardua Stark, 1894, Nebria 85

arelatensis Lapouge, 1903, Carabus 160

arenaria Putzeys, 1865, Amara [Leiocnemis] 555



arenaria Fabricius, 1792, Cassolaia [Cicindela] 101

arenaria Fabricius, 1792, Clivina [Scarites] 219

arenaria Füessly, 1775, Cylindera [Cicindela] 110



arenaria Fabricius, 1775, Eurynebria [Carabus] 79

arenarius Bonelli, 1813, Scarites 232

arenarius V. Petagna, 1787, Scarites 232

arenicola Wesmael, 1872, Harpalus 373

arenicola Wollaston, 1863, Masoreus 357



arenicola Wollaston, 1854, Syntomus [Dromius] 432

arenicus Kalashyan, 1979, Laemostenus [Pristonychus] 538

arenosus Stephens, 1827, Dyschirius 227

arenstorffianus Absolon, 1913, Scotoplanetes 320

areolatus Creutzer, 1799, Perileptus [Carabus] 23, 288

areolatus Duftschmid, 1805, Scarodytes [Dytiscus] 68

areolatus Gravenhorst, 1807, Scarodytes [Dytiscus] 68

arerae Schauberger, 1927, Abax 469



arettensis Jeannel, 1941, Geotrechus 310

argaeicola Ganglbauer, 1905, Bembidion 256

argaensis Bilton & Fery, 1996, Rhithrodytes 68

argenteolum Ahrens, 1812, Bembidion 247



argodi Carret, 1898, Poecilus 482

argodi Ganglbauer, 1900, Scotodipnus 240



argodi Abeille de Perrin, 1886, Trichaphaenops [Anophthalmus] 344

argonautarum Semenov, 1898, Carabus 168



arguta Casey, 1918, Amara [Curtonotus] 558

argutoriformis Poppius, 1906, Pterostichus [Cryobius] 489

argutoroides Bates, 1873, Oxycentrus [Harpalus] 392

ariasi Bolívar y Pieltain, 1914, Trymosternus 410

arisana Jedlika, 1951, Lebia 428

arisanum Jedlika, 1940, Agonum [Colpodes] 452

aristidis Leprieur, 1879, Hydrovatus 70

aristochroides Deuve, 1992, Carabus 134



arizonensis Puel, 1936, Pterostichus [Haptoderus] 490

arkhyzicus Belousov, 1998, Alanorites 290

arlequinus Deuve, 1994, Carabus 127

arlettae Morvan, 1998, Andrewesius 456

armandi Fairmaire, 1886, Cylindera [Cicindela] 109

armata Ledoux & Roux, 1999, Nebria 89

armata Coiffait, 1969, Typhlocharis 241

armatus Wollaston, 1867, Dineutus 26

armatus Wollaston, 1864, Dyschirius 227

armellinii Ganglbauer, 1900, Binaghites [Scotodipnus] 239

armena Motschulsky, 1844, Amara [Bradytus] 553

armeniaca Motschulsky, 1839, Amara [Leirus] 547



armeniaca Brullé, 1834, Megacephala 118

armeniacum Chaudoir, 1846, Bembidion 20, 269



armeniacus Sharp, 1882, Agabus 40

armeniacus Mannerheim, 1830, Carabus 194



armeniacus Motschulsky, 1865, Chlaenius [Epomis] 353

armeniacus Chaudoir, 1879, Cychrus 202

armeniacus Casale, 1979, Duvalius 301

armeniacus Chaudoir, 1846, Harpalus 382

armeniacus Zaitzev, 1927, Hygrotus [Coelambus] 71

armeniacus Casale, 1988, Laemostenus 538

armeniacus Osculati, 1844, Sphodrus [Sphodrus] 543

armeniacus Ganglbauer, 1915, Zabrus 570



armeniae Lutshnik, 1937, Amara 552

armenica Laporte, 1834, Megacephala 118

armenius Zaitzev, 1930, Carabus 185

armenus Motschulsky, 1850, Calathus 528

armenus Jedlika, 1950, Chlaenius 352



armenus K. Daniel, 1904, Harpalus [Ophonus] 377

armenus Faldermann, 1836, Pterostichus [Omaseus] 495

armenus Iablokoff‑Khnzorian, 1963, Trechus 326

armiger Imura, 1997, Carabus 188

armiger Morita, 1997, Trichotichnus 395

armoricana Chaudoir, 1873, Cymindis 411

armoricanus Razet, 1951, Brachinus 214

armoricanus Oberthür, 1935, Carabus 138

armoricus Porta, 1923, Calathus 528

arnoldiana Kryzhanovskij, 1974, Amara [Curtonotus] 559



arnoldii Lutshnik, 1934, Amara 556

arnoldii Kryzhanovskij, 1962, Chilotomus 364

arnoldii Kabak, 1999, Cymindis 414

arnoldii Jeannel, 1962, Duvalius 301

arnoldii Gryuntal, 1984, Dyschirius 228

arnoldii Kataev, 1988, Harpalus 373

arnoldii Kabak & Komarov, 1995, Paradromius [Paradromius] 424

arnoldii Erjiomin, 1996, Pterostichus 489

arnoldii Belousov, 1987, Trechus 326

arnosti Nakane, 1978, Bembidion 267

arnostiana Kabak, nom. nov., Cymindis 24, 416

aroaniae Casale, Giachino, Vailati & Vigna Taglianti, 1996, Duvalius 301



arotus Alluaud, 1936, Dyschirius 225

arpaalaniensis P. Machard, 1985, Carabus 148

arpadis Csiki, 1907, Amara 551

arragonense Wagner, 1926, Bembidion 244

arragonensis Heyden, 1883, Zabrus 569

arribasi Ortuño, 1991, Pterostichus 497

arribasi Jeanne, 1988, Trechus 326

arrisi Jedlika, 1937, Bembidion 261

arrisi Jeannel, 1962, Trechus 333

arrogans Schaum, 1859, Carabus 151



arrogans Péringuey, 1908, Polyderis [Tachys] 274

arrogans Duftschmid, 1812, Pterostichus [Carabus] 514

arrogantior Deuve, 1991, Carabus 123

arrowi Baliani, 1934, Amara 563

arrowi G. Hauser, 1913, Carabus [Apotomopterus] 123

arrowi Lutshnik, 1931, Diodercarus 286

arrowi Guignot, 1936, Haliplus 31

arrowi Régimbart, 1907, Metagyrinus [Aulonogyrus] 28

arrowi Jedlika, 1936, Pterostichus 486

arrowi Grouvelle, 1908, Yamatosa [Rhysodes] 78

arrowi R. T. Bell & J. R. Bell, 1978, Yamatosa [Yamatoa] [HN] 78

arrowianum Breuning, 1934, Calosoma 120

arrowianus Breuning, 1934, Carabus [Apotomopterus] 177


arrowianus Jedlika, 1938, Pterostichus 486

arsenjevi Lutshnik, 1935, Amara 550

arsenjevi Lafer, 1979, Pterostichus 495

arsenjevi Jeannel, 1962, Trechus [Epaphius] 325

arsiyandagensis P. Machard, 1985, Carabus 194

artemisiae Putzeys, 1872, Trechus 326

arthuri Morvan, 1982, Laemostenus [Ceuthosphodrus] 537

arthuri P. Moravec & Lompe, nom. nov., Trechus 22, 326

articervix Cavazzuti, 1999, Carabus 182

articulatoides Jedlika, 1932, Bembidion 248

articulatum Panzer, 1796, Bembidion [Carabus] 270

artipunctatus Dvoák, 1993, Pachycarus [Mystropterus] 367

artus Andrewes, 1937, Omophagus 424

arunachalensis Saha & Biswas, 1985, Clivina 219

arunensis Heinertz, 1980, Carabus 161

arunensis Naviaux, 1994, Neocollyris 116

aruspex Clark, 1864, Hydaticus 54

arvarum Morvan, 1996, Xestagonum 467

arvena Jeannel, 1942, Cymindis 412

arvernus Jeannel, 1937, Pterostichus [Haptoderus] [HN!] 489



arvernus Sirguey, 1924, Pterostichus [Omaseus] 499

asahiensis Habu & Baba, 1962, Cymindis 418

asahinae Nakane, 1957, Amphimenes 435

asahinai Uéno, 1974, Perileptus 289

asahinus Habu & Baba, 1960, Pterostichus 497

asakawaensis Habu, 1973, Stenolophus 404

asakoae Morita, 1994, Negreum [Colpodes] 460

asaphus Antoine, 1941, Harpalus 374

ascendens K. Daniel, 1902, Bembidion 245

ascendens Wollaston, 1862, Calathus 526

ascendens Semenov, 1898, Carabus 124

ascetes Kataev & Wrase, 1997, Harpalus 373

aschnae Makhan, 1999, Peltodytes 33

asetosus Uéno, 1997, Trechus 326

ashutorensis Belousov & Kabak, 1992, Trechus 331

asiabadensis Jedlika, 1961, Cymindis 416

asiaeminoris Netolitzky, 1935, Bembidion 256

asiaeminoris Casale, 1988, Laemostenus 537

asiatica Jedlika, 1957, Amara 559

asiatica Audouin & Brullé, 1839, Cicindela 103

asiatica Ledoux & Roux, 1998, Nebria [add to incertae sedis] 95

asiaticum Jedlika, 1965, Bembidion 265

asiaticus Ballion, 1871, Broscus 236

asiaticus Sharp, 1882, Cybister 50

asiaticus Laporte, 1834, Licinus 442


asiaticus Mateu, 1971, Microlestes 422

asiaticus Chaudoir, 1852, Odotoncarus [Ditomus] 366

asiaticus Poppius, 1906, Pterostichus [Cryobius] 489

asiaticus Jeannel, 1927, Trechus 326

asiaticus Laporte, 1834, Zabrus 570

asiorum Müller‑Motzfeld, 1990, Bembidion 257

askalensis Wewalka, 1992, Hydroporus 60

askellek Morvan, 1998, Xestagonum 467

askhicus Belousov, 1992, Carabus 131

askhicus Belousov, 1991, Pterostichus 496

asmarensis Jedlika, 1956, Abacetus 347

asonis Uéno, 1974, Rakantrechus 319

asper Jedlika, 1936, Platynus 462

asper Morita, 1997, Trichotichnus 395

asperatus Dejean, 1826, Carabus [Calosoma] 137

asperatus Mulsant & Wachanru, 1853, Carabus [Procrustes] 188

aspericolle Germar, 1829, Bembidion [Periphus] 268

aspericollis Bates, 1873, Chlaenius 354

asperipennis Lapouge, 1921, Carabus 166

asperipennis Habu, 1978, Myas 475



asperulus Kraatz, 1878, Carabus 166

asperum Bates, 1883, Lachnoderma 438



asperus Beuthin, 1892, Carabus 129

aspettatii Magrini, 1982, Duvalius 304



asphaltinus Klug, 1832, Scarites 233

aspromontis Hille, 1914, Harpalus 371

assalemi Jedlika, 1968, Bembidion 252

assamensis Parry, 1844, Calochroa [Cicindela] 99

assamensis Mandl, 1965, Carabus 162

assamensis Vazirani, 1969, Hyphydrus 75

assamensis Schauberger, 1935, Lampetes [Lamprophonus] 387

assamensis Kirschenhofer, 1992, Oxycentrus 392

assamensis Rivalier, 1964, Prothyma 115

assamensis Putzeys, 1846, Pseudoclivina [Clivina] 221

assamensis Deuve, 1986, Synuchus 545

assequens G. Ochs, 1936, Orectochilus 29

assimile Motschulsky, 1865, Agonum [Agonothorax] 451

assimile Gyllenhal, 1810, Bembidion 248



assimile Stephens, 1828, Bembidion [Lopha] 270

assimilis Sturm, 1834, Agabus [Colymbetes] 37

assimilis Chaudoir, 1844, Amara 550

assimilis Hope, 1831, Calomera [Cicindela] 100



assimilis Cristoforis & Jan, 1837, Carabus 186

assimilis Duftschmid, 1812, Carabus 196

assimilis Kwon & Lee, 1984, Carabus [Tomocarabus] 149

assimilis Chaudoir, 1843, Cicindela 107

assimilis Dejean, 1829, Harpalus 371

assimilis Paykull, 1798, Nebrioporus [Dytiscus] 66

assimilis Chaudoir, 1844, Patrobus 285

assimilis Chaudoir, 1876, Pheropsophus 217

assimilis Paykull, 1790, Platynus [Carabus] 464

assimilis Hochhuth, 1871, Pterostichus [Argutor] 507

assimilis Heer, 1837, Trechus 331

assingi Wrase & Jaeger, 1996, Bradycellus 400

assingi Lompe, 1999, Trechus 326

assmanni Balkenohl, 1999, Reicheiodes 230



assonensis Jeannel, 1941, Geotrechus 309

assuanensis Mjöberg, 1905, Tetragonoderus 357

asteriscus White, 1844, Macrocheilus [Acanthogenius] 407

asthenes Uéno, 1987, Rakantrechus 319

astictus Bates, 1883, Morimotoides [Colpodes] 460

astrabadense Mannerheim, 1844, Bembidion [Peryphus] 245

astrabadensis Lutshnik, 1935, Amara 562

astrabadensis Semenov, 1903, Carabus 193

astrabadensis Reitter, 1902, Licinus 442

astrakhanus Vondel, 1991, Haliplus 31

astur Sharp, 1873, Anchomenidius [Anchomenus] 529



astur Lapouge, 1898, Carabus 195

astur Sharp, 1882, Hygrotus [Hydroporus] 72

asturiae Schauberger, 1928, Harpalus 375

asturicus Heinz, 1965, Anchomenus [Platynus] 455

asturicus Jeanne, 1966, Pterostichus [Lianoe] 497


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