Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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persica Chaudoir, 1842, Amara 547

persica Faldermann, 1836, Cicindela 106

persica Jedlika, 1968, Cymindis 24, 417



persica Morvan, 1971, Tachyta 277

persicola W. Horn, 1934, Lophyra [Cicindela] 113

persicum Ménétriés, 1832, Bembidion 262

persicum Géhin, 1885, Calosoma [Callisthenes] 118



persicus Jedlika, 1968, Chlaenius 354

persicus L. Redtenbacher, 1850, Chlaenius 354



persicus LaFerté‑Sénectère, 1851, Chlaenius 352

persicus Morvan, 1975, Chlaenius 354

persicus Peschet, 1914, Deronectes 58

persicus Fedorenko, 1994, Dyschirius 226

persicus Wehncke, 1876, Dytiscus 52

persicus Brancucci, 1983, Laccophilus 76

persicus Holdhaus, 1912, Microlestes 423

persicus Breit, 1914, Notiophilus 97

persicus Mandl, 1963, Pheropsophus 217


persicus Jedlika, 1961, Phloeozetus 438

persicus Apfelbeck, 1904, Pogonus 288

persicus Chaudoir, 1842, Scarites 232

persicus Mannerheim, 1844, Stenolophus 405

persicus Jedlika, 1952, Taphoxenus 544

persicus Chaudoir, 1878, Tapinopterus [Hoplodactylus] 518

persimile A. Morawitz, 1862, Bembidion 257

persimilis Uéno, 1969, Awatrechus 296

persimilis Breuning, 1950, Carabus 176

persimilis Harald Lindberg, 1950, Orthomus [Pterostichus] 477

persuasum Netolitzky, 1938, Bembidion 265

pertinax Tschitschérine, 1895, Pterostichus [Feronia] 495

perturbatocostatus M. Grotz & R. Grotz, 1980, Carabus 166

pertusus Schaum, 1858, Poecilus [Pterostichus] 480



pertyi Gistel, 1837, Myriochila [Cicindela] 113

pertyi Heer, 1837, Trechus 338

pervarius Kleinfeld, 1997, Carabus 126

pervasus Gozis, 1914, Graptodytes [Hydroporus] 59

pervenustus Wollaston, 1864, Metadromius [Dromius] 422



perversus Roubal, 1917, Harpalus 378

perviridis Reitter, 1896, Carabus 139

pesarinii Sciaky, 1982, Boldoriella 296

pesarinii Heinz & Pavesi, 1994, Calosoma 118

pesarinii Bucciarelli, 1979, Speluncarius 516



pescheti Guignot, 1930, Hydroporus 61

pescheti Peyerimhoff, 1929, Yola 57

pesianus Focarile, 1949, Trechus 335

petegariensis Uéno, 1971, Trechiama 322

petercatei Kirschenhofer, 1992, Trichotichnus 397

peterfrankei Kleinfeld, 1998, Carabus 127



petisii Duftschmid, 1812, Harpalus [Carabus] 382

petraeus Knirsch, 1927, Duvalius [Neoduvalius] 307



petreus Motschulsky, 1844, Harpalus [Microderus] 372

petri Hummel, 1825, Amara 553

petri Semenov, 1902, Calosoma 120

petri Semenov & Znojko, 1932, Carabus 138

petri Semenov & Znojko, 1929, Colpostoma 441

petri Tschitschérine, 1902, Harpalus 380



petrii Knirsch, 1914, Duvalius [Trechus] 299

petrimagni Netolitzky, 1920, Bembidion 258

petrimagni Kryzhanovskij & Mikhailov, 1975, Laemostenus [Pristonychus] 538

petrimontii Hieke, 1995, Amara 551



petrobia Gistel, 1838, Cicindela 107

petrochilosi Coiffait, 1965, Duvalius 306

petroffi Antoine, 1933, Platytarus 410


petrogorbatschevi Zamotajlov, 1996, Diplous 283

petrosum Gebler, 1833, Bembidion 262



petrovitzi Mandl, 1973, Brachinus [Dromius] 214

petrovitzi Mandl, 1973, Cymindis 414

petrovitzi Mandl, 1973, Trichis 439

petulans Casey, 1918, Bembidion 262

petulans Jedlika, 1938, Pterostichus 495



petzi Sokolá, 1907, Carabus 139

petzianus Born, 1911, Carabus 181

pewtzowi Tschitschérine, 1897, Harpalus 374

pexus Ménétriés, 1848, Harpalus 374

peyerimhoffi Antoine, 1959, Acinopus 368



peyerimhoffi Bruneau de Miré & Legros, 1963, Agabus 39

peyerimhoffi Sainte‑Claire Deville, 1935, Bembidion 255

peyerimhoffi Régimbart, 1906, Deronectes [Hydroporus] 58

peyerimhoffi Jakobson, 1906, Duvalius [Trechus] 303

peyerimhoffi Puel, 1929, Dyschirius 227

peyerimhoffi Alluaud, 1923, Nebria 93

peyerimhoffi Paulian & Villiers, 1939, Ocys [Oreocys] 273

peyerimhoffi Antoine, 1933, Tachyura [Tachys] 277

peyerimhoffi Jeannel, 1907, Trechus 22, 326

peyroni Gemminger & Harold, 1868, Chlaenius 352

peyroni Putzeys, 1866, Dyschirius 224

peyroni Carret, 1905, Gynandromorphus 362

peyroni Piochard de la Brûlerie, 1873, Penthophonus [Penthus] 393

pfefferi Maan, 1933, Bembidion 263

pfefferi Kult, 1949, Dyschirius 230

pfefferi Wewalka, 1974, Hydroporus 64



pfefferi Jedlika, 1964, Pseudotaphoxenus [Taphoxenus] 542

pfeiferi Apfelbeck, 1908, Aphaenopsis [Trechus] 293



pfeiferi Apfelbeck, 1912, Aphaenopsis [Anophthalmus] 293

pfeiffii C. R. Sahlberg, 1827, Bembidion 246

pfizenmayeri Netolitzky, 1943, Bembidion 266

pfizenmayeri Poppius, 1906, Pterostichus 499

phaedimus Schaufuss, 1882, Carabus 158

phaedrum Andrewes, 1923, Bembidion 271

phaeopus Sainte‑Claire Deville, 1902, Pterostichus [Platysma] 513

phaeus Lutshnik, 1922, Pterostichus [Platysma] 496

phami Deuve, 1988, Carabus 127

phami Deuve, 1981, Trechus 338

phamianus Deuve, 1990, Carabus 133

phanagoriacus Belousov, 1990, Trechus 338

pharao Lutshnik, 1916, Poecilus [Platysma] 480

pharao Schatzmayr & Koch, 1934, Tachys [DA] 276

pharaonis Motschulsky, 1865, Chlaenius [Epomis] 353


pharaonum Bedel, 1914, Anthia 347

pharina Andrewes, 1929, Nebria 93

pharisaeus Reiche & Saulcy, 1855, Harpalus 379

phenax Antoine, 1941, Microlestes 423

philippensis Jeannel, 1937, Duvalius 308

philippinensis Jedlika, 1935, Dicranoncus 458



philippinensis Jedlika, 1935, Sericoda [Agonum] 466

philippinus Jedlika, 1939, Craspedophorus 447

phillipinensis Kult, 1951, Clivina 220

philoscius Cavazzuti, 1995, Carabus 127



phlagochensis Imura & Zamotajlov, 1993, Carabus 133

phobon Netolitzky, 1930, Bembidion 254

phoebus Andrewes, 1932, Apristus 431

phoebus Kurnakov, 1962, Carabus 131

phoeniceum Huber & Marggi, 1997, Bembidion 265

phoenicius Vigna Taglianti, 1973, Duvalius 305

phoenix Tschitschérine, 1894, Amara 566

phoenix Wewalka, 1989, Bidessus 55

phoenix Lapouge, 1925, Carabus 130

phoenix Tschitschérine, 1902, Ophonus [Harpalus] 389

phreatica Uéno, 1957, Morimotoa 75

phryganobium Belousov & Sokolov, 1996, Bembidion 259



phryganobius Iablokoff‑Khnzorian, 1963, Trechus 335

phrygius Guignot, 1942, Graptodytes 60

phrygius J. Müller, 1932, Tapinopterus 518

phygas Antoine, 1956, Bembidion [Metallina] 251

physignathus Andrewes, 1937, Sphodropsis 543

piazzolii Focarile, 1950, Trechus 338

picanyoli Español & Escolá, 1983, Geotrechus 310



picciolii Bedel, 1876, Duvalius [Trechus] 302

picea Jedlika, 1956, Cymindis 24, 416

picea Dejean, 1826, Oreonebria [Nebria] 96



piceatus Sharp, 1882, Dytiscus 52

picena Binaghi, 1936, Typhloreicheia 223

picenus A. Villa & G. B. Villa, 1838, Carabus 166

piceocyaneum Solsky, 1874, Bembidion 246



piceola Fairmaire, 1887, Lachnocrepis [Oodes] 445

piceola Chaudoir, 1877, Parena [Crossoglossa] 434

piceolus Zaitzev, 1908, Agabus 41

piceolus Bates, 1873, Amphimenes 435

piceolus Chaudoir, 1850, Pterostichus [Feronia] 511

piceonigrum Jedlika, 1936, Agonum [Colpodes] 454

piceophilus Deuve, 1994, Carabus 175



picescens Ganglbauer, 1904, Duvalius [Trechus] 301

picescens Wollaston, 1864, Licinopsis [Pristonychus] 540



picescens Jedlika, 1936, Platynus [Colpodes] 464

picescens Fairmaire, 1887, Pristosia [Pristonychus] 531

piceum Linné, 1758, Agonum [Carabus] 453

piceus Rottenberg, 1870, Acupalpus [Stenolophus] 399

piceus A. I. Jakovlev, 1897, Agabus 39

piceus Zaitzev, 1905, Agabus 41

piceus Andrewes, 1931, Amblops 435

piceus Klug, 1834, Colymbetes 45

piceus Stephens, 1828, Hydroporus 64

piceus Sturm, 1835, Hydroporus 61

piceus Motschulsky, 1850, Laemostenus [Pristonychus] 538

piceus Frölich, 1799, Leistus 18, 82

piceus Panzer, 1793, Molops [Scarites] 474

piceus Nietner, 1856, Nanodiodes [Oodes] 445

piceus Dejean, 1828, Platynus 463

piceus Dejean, 1831, Platynus [Anchomenus] [HN] 464

piceus Fischer von Waldheim, 1829, Pterostichus [Argutor] 505

piceus Ballion, 1871, Stenolophus [Badister] 406

piceus Edmonds, 1934, Tachys 275

piceus Putzeys, 1847, Trechus 331

pici Schatzmayr, 1909, Badister 440

pici Régimbart, 1895, Haliplus 33

picicollis Chaudoir, 1876, Pheropsophus 218

picicolor Lindroth, 1956, Synuchus 546

picicornis Stephens, 1828, Agabus [Colymbetes] 38

picicornis Mandl, 1972, Chlaenius [Stenochlaenius] 355

picicornis Dejean, 1831, Distichus [Scarites] 231

picicornis J. R. Sahlberg, 1875, Hydroporus 64

picicornis Dejean, 1831, Laemostenus [Sphodrus] 539

picicornis Fabricius, 1801, Nebria [Carabus] 91

picicornis Faldermann, 1836, Parophonus [Ophonus] 393

picicornis Zetterstedt, 1828, Patrobus [Harpalus] 285

picilabris Stephens, 1835, Harpalus 373

picimanus Duftschmid, 1812, Pterostichus [Carabus] 485

picina Solsky, 1874, Amara [Celia] 555

picinus Marsham, 1802, Agabus [Dytiscus] 40

picinus Baudi di Selve, 1864, Amblystomus 360



picipennis J. R. Sahlberg, 1903, Agabus 39

picipennis Duftschmid, 1812, Harpalus [Carabus] 380



picipennis Motschulsky, 1844, Pterostichus 509

picipes Olivier, 1795, Acinopus [Scarites] 23, 368



picipes Fabricius, 1787, Agonum [Carabus] 453

picipes Motschulsky, 1844, Amara [Leirus] 561

picipes Duftschmid, 1812, Asaphidion [Elaphrus] 242

picipes Kirby, 1837, Bembidion [Peryphus] 271

picipes Sturm, 1825, Bembidion 249

picipes W. S. MacLeay, 1825, Caelostomus [Anaulacus] 471



picipes Bonelli, 1813, Clivina 219

picipes Herbst, 1806, Cychrus 202

picipes Fabricius, 1787, Hygrotus [Dytiscus] 72

picipes Kirby, 1837, Ilybius [Colymbetes] 43



picipes Chaudoir, 1868, Pterostichus [Feronia] [HN] 512

picipes Dejean, 1831, Pterostichus [HN] 512

picipes Fischer von Waldheim, 1829, Pterostichus [Argutor] [ND] 516



piciventris C. G. Thomson, 1868, Graphoderus 49

piciventris Graëlls, 1858, Trechus 339

picoensis Machado, 1988, Trechus 338

picotae Foster & Bilton, 1997, Agabus 40

picta Motschulsky, 1864, Cylindera [Parmecus] 109

picta Pallas, 1771, Cymindis [Carabus] 414

picta Semenov, 1891, Nebria 91

picteti Westwood, 1874, Platyrhopalopsis [Platyrhopalus] 211

picticornis Chaudoir, 1878, Abacetus 347

picticornis Lindroth, 1956, Synuchus 545

picticornis A. Fleischer, 1898, Trechus 335

pictipennis Sharp, 1882, Laccophilus 77

pictipennis Sharp, 1873, Platambus [Agabus] 44

pictipennis Putzeys, 1875, Tachys 276

pictiventris Fauvel, 1888, Nebria 93



pictiventris Régimbart, 1905, Rhantus 46

pictulum Bates, 1874, Cymbionotum [Coscinia] 235



pictulus Bates, 1873, Anomotarus [Cymindis] 408

pictum Kolenati, 1845, Asaphidion [Tachypus] 21, 242



pictum Duftschmid, 1812, Bembidion [Elaphrus] 270

pictum Faldermann, 1835, Bembidion [Peryphus] 262

picturatum Fairmaire, 1898, Bembidion 258

pictus Meier, 1899, Agabus 37

pictus Bates, 1873, Badister 440

pictus Hope, 1833, Brachinus [Aploa] 212

pictus Chaudoir, 1856, Chlaenius 354

pictus Fischer von Waldheim, 1823, Daptus 369

pictus Fabricius, 1787, Graptodytes [Dytiscus] 59



pictus Mannerheim, 1844, Haliplus 30

pictus Klug, 1834, Hyphydrus 75



pictus Küster, 1851, Laccophilus 76

pictus Andrewes, 1923, Platymetopus 393

pictus Sharp, 1882, Prodaticus 54

pieli Guignot, 1936, Hyphydrus 74

pieltaini Jeannel, 1920, Trechus 338

pieperi Erber, 1990, Thalassophilus 346


pietrii Normand, 1933, Graptodytes 59

pietrorattii Deuve, 1991, Carabus 174

piffli Mandl, 1961, Carabus 155

piffli Jedlika, 1964, Cymindis 414



piger Duftschmid, 1812, Harpalus [Carabus] 373

piger Dejean, 1828, Zabrus 572

piger Geoffroy, 1785, Zabrus [Buprestis] 573

pigmentatus Cavazzuti, 1998, Carabus 128

pignataroi Cavazzuti, 1996, Carabus 146

pihyonensis Deuve & Li, 1998, Carabus 122



pilicasius Nakajima, 1993, Carabus [Acoptolabrus] 122

pilicornis Fabricius, 1775, Loricera [Carabus] 98

pilicornis Donovan, 1804, Paussus 210

pilifer Ganglbauer, 1891, Duvalius [Trechus] 300

pilifer Bedel, 1900, Paradromius [Dromius] 425

pilifer Bates, 1878, Pterostichus [Molops] 494

piligera Chaudoir, 1883, Trichocoptodera [Coptodera] 437

pilipes Piochard de la Brûlerie, 1868, Acinopus 368



pilipes J. R. Sahlberg, 1921, Hydroporus 61

pilipes Germar, 1817, Pogonus [Carabus] 287

pilisensis Csiki, 1918, Trechus 338

pilleri Csiki, 1916, Abax 470

pillichi Bernau, 1914, Carabus 179

pilosa Gebler, 1825, Cymindis 415

pilosella L. Miller, 1868, Duvaliopsis [Anophthalmus] 298



pilosella Motschulsky, 1862, Lebia [Lebida] 427

pilosicoerulans Mandl, 1989, Chlaenius 350

pilosicornis Deuve & Quéinnec, 1993, Trechus 338

pilosipennis Machado, 1992, Calathus 528

pilosipennis Escalera, 1922, Cymindis 414

pilosissima Reitter, 1894, Cymindis 411

pilosistriata Uéno, 1980, Ushijimaella 345

pilosohumeralis Zamotajlov, 1993, Parapenetretus 284

pilosula Antoine, 1963, Lebia 428

pilosus Uéno, 1957, Awatrechus 296



pilosus Dejean, 1825, Carterus [Ditomus] 364

pilosus Huang, 1995, Harpalus 385

pilosus Guignot, 1949, Hydrotarsus 65

pilosus Fischer von Waldheim, 1817, Licinus [Scales] 442

pilosus Bates, 1892, Lymnastis 274

pilosus Host, 1789, Pterostichus [Carabus] 487

pimalicus Casey, 1914, Harpalus 378

pimelioides Lapouge, 1922, Carabus 152

pinchuanensis Imura, 1999, Carabus 126

pincum De Monte, 1957, Bembidion 247


pindarica Andrewes, 1925, Nebria 94

pindica Apfelbeck, 1904, Amara 551

pindica Jeanne, 1974, Nebria 92

pindicola Apfelbeck, 1901, Cymindis 414

pindicum Apfelbeck, 1901, Bembidion 257



pindicus Mandl, 1989, Cychrus 205

pindicus J. Müller, 1930, Molops 474

pindicus Maan, 1939, Tapinopterus 519

pinetorum Lapouge, 1910, Carabus 140

pinetorum Graëlls, 1858, Pterostichus [Corax] 489

pinganensis G. Hauser, 1920, Carabus [Coptolabrus] 144

pingshiangi Jedlika, 1957, Amara 554

pinguedineus Eschscholtz, 1823, Pterostichus [Poecilus] 492

pinguis Andrewes, 1930, Amara 558

pinguis Lapouge, 1914, Carabus 188

pinguis Régimbart, 1892, Hydrovatus 70



pinguis Dejean, 1828, Pterostichus [Feronia] 503

pinguis Andrewes, 1929, Scarites 234

pinguis Kiesenwetter, 1850, Trechus 334

pinguis Dejean, 1831, Zabrus 568

pinguiusculus Habu, 1978, Synuchus 546

pingwuensis Deuve, 1997, Carabus 125



pingyangensis G. Hauser, 1929, Carabus [Coptolabrus] 143

pinicola Graëlls, 1851, Laemostenus [Pristonychus] 536

pinipes Kabak, 1997, Leistus 84

piniphilus Gistel, 1857, Pterostichus [ND] 516

pinkeri Netolitzky, 1935, Bembidion 253

pinkeri Ganglbauer, 1891, Trechus 338

piochardi Géhin, 1884, Carabus [Chaetomelas] 188

piochardi Saulcy, 1874, Paussus 210



piochardi Régimbart, 1878, Stictotarsus [Hydroporus] 69

piperi Casey, 1924, Amara [Celia] 565

pippiai G. Fiori, 1961, Laemostenus [Actenipus] 533

pirata Schatzmayr, 1909, Agonum [Platynus] 451

pirata Semenov & Znojko, 1932, Carabus 133

piraticus Fairmaire, 1880, Carabus 152

pirazzolii Putzeys, 1873, Calathus 525



pirazzolii Géhin, 1876, Carabus 130

pirica Uéno, 1992, Trechus 325

piriformis Deuve, 1997, Carabus 124

pirinensis Mandl, 1985, Carabus 165

pirinensis Horvatovich, 1973, Nebria 86



pirinicus V. B. Guéorguiev, 1957, Ilybius 42

pirinicus Pawowski, 1972, Trechus 338

pirithous Reitter, 1896, Carabus 181


pisangana Hieke, 1995, Amara 564

pisanus Laporte, 1835, Dytiscus 52

pisidicus Jeannel, 1937, Anillidius [Nannotrechus] 290

pisidicus Peyron, 1855, Carabus [Procrustes] 182

pisidicus J. Müller, 1932, Laemostenus 538

pisidicus J. Müller, 1932, Tapinopterus 518

pithyusicus Lutshnik, 1933, Pterostichus [Platysma] 492

pivai Vigna Taglianti & Sciaky, 1988, Lessinodytes 314



pjasinensis Gottwald, 1993, Carabus 174

pjatigorianus Zamotajlov, 1996, Deltomerus 283



placida Putzeys, 1866, Clivina 220

placidus Gyllenhal, 1827, Dicheirotrichus [Harpalus] 403

placidus Rosenhauer, 1847, Tapinopterus [Pterostichus] 518

placidus Jeannel, 1962, Trechus 338



plagella Reitter, 1908, Rhopalostyla [Cymindis] 430

plagiata Klug, 1834, Drypta 358

plagiata Bänninger, 1923, Nebria 91

plagiata Putzeys, 1875, Perigona 448

plagiatus Reiche, 1868, Brachinus 213

plagiatus Duftschmid, 1812, Microlestes [Lebia] 423

plagiatus Reiche & Saulcy, 1855, Phloeozetus [Coptodera] 438

plagipennis Wollaston, 1865, Dromius 420

plana Bonelli, 1813, Siagona 218

planarius Obydov, 1994, Carabus 199

planaticollis Kirschenhofer, 1987, Pterostichus 505

planatus Chaudoir, 1843, Carabus 161



planatus Motschulsky, 1850, Carabus 199

planatus Motschulsky, 1844, Harpalus 383

planatus Régimbart, 1899, Hydaticus 53

planatus Schaufuss, 1885, Miscelus 436

planatus Peyron, 1858, Pedius [Feronia] 478

planatus Ménétriés, 1849, Poecilus 479

planatus Duftschmid, 1812, Trechoblemus [Carabus] 324

plancyi Tschitschérine, 1897, Harpalus 380



planicola Semenov, 1897, Cylindera [Cicindela] 111

planicola Tschitschérine, 1899, Pterostichus [Feronia] 509



planicolle Motschulsky, 1860, Bembidion [Metallina] 271

planicollis Schaum, 1857, Acupalpus [Stenolophus] 399

planicollis Küster, 1846, Carabus 164


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