Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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pseudosobaekensis Deuve & Li, 1998, Carabus 142

pseudosteuarti Lapouge, 1925, Carabus 179



pseudostrenua Kult, 1946, Amara 568

pseudostuxbergi Poppius, 1906, Pterostichus [Cryobius] 492



pseudosubsinuatus Hänel, 1940, Pterostichus 493

pseudosuturalis Bedel, 1906, Cymindis 414

pseudotanguticus Deuve & Kaláb, 1992, Carabus 176

pseudotaphum Morvan, 1973, Bembidion [Peryphus] 262

pseudotenellum Netolitzky, 1910, Bembidion 249

pseudotinctulus Schauberger, 1932, Harpalus 386

pseudotorosus Schweiger, 1969, Carabus 157

pseudotorquatus Deuve, 1995, Carabus 126

pseudotosanus Deuve & Mourzine, 1997, Carabus 129

pseudotroberti Lapouge, 1911, Carabus 140

pseudotypica Lutshnik, 1924, Cicindela 107

pseudouralense Fassati, 1945, Bembidion 263



pseudoustulatum J. Müller, 1934, Bembidion 263

pseudoviolaceus Kraatz, 1886, Carabus [Megadontus] 165

pseudowagae Deuve, 1995, Carabus 175

pseudoyunnanus Deuve, 1994, Carabus 129

pshuensis Gottwald, 1985, Carabus 132



psilax Andrewes, 1935, Bembidion 271

psilica Bates, 1866, Cylindera [Cicindela] 111

psilodorum Andrewes, 1933, Bembidion 262

psilopterus Kraatz, 1879, Carabus [Megadontus] 166

psittaceus Geoffroy, 1785, Harpalus [Buprestis] 376

pskemicus Deuve & Kaláb, 1993, Carabus 147

pslukhensis Fedorenko, 1996, Reicheiodes 230

psophia Audinet‑Serville, 1821, Brachinus 213



psota Jedlika, 1938, Pterostichus 486

psuchrum Andrewes, 1922, Bembidion 20, 271

psyllocephala K. Daniel & J. Daniel, 1898, Amara 563

pterostichus Reitter, 1900, Harpalus [Ophonus] 380



pubens Bedel, 1876, Anophthalmus [Trechus] 291

puberulus LeConte, 1850, Hydroporus 64



puberulus Mannerheim, 1853, Hydroporus 61

pubescens Joseph, 1871, Anophthalmus 291

pubescens Ménétriés, 1832, Chlaenius 348

pubescens Paykull, 1790, Dicheirotrichus [Carabus] 402

pubescens O. F. Müller, 1776, Harpalus [Carabus] 385

pubescens Gyllenhal, 1808, Hydroporus [Hyphydrus] 64

pubescens Chaudoir, 1850, Siagona 218


pubescens Mateu, 1986, Syntomus 433

pubescens Nietner, 1858, Zuphium 574

pubicollis Uéno, 1957, Yamautidius [Rakantrechus] 345

pubifer Reitter, 1897, Licinus 442

pubiger L. Dufour, 1820, Chlaenius 355

pubipennis L. Dufour, 1820, Lebia 428



publipennis Küster, 1849, Parophonus [Harpalus] 392

puchneri Lebenbauer, 1998, Orotrechus 317



pucholti Jedlika, 1963, Cymindis 417

pucholti Jedlika, 1963, Lebia 430

pucholti Jedlika, 1962, Poecilus [Pterostichus] 483



pucholti Jedlika, 1964, Pseudotaphoxenus [Taphoxenus] 541

pueli Antoine, 1941, Acinopus 368

pueli Schatzmayr, 1935, Acinopus 368

pueli Antoine, 1923, Amara 555

pueli Schauberger, 1933, Anisodactylus 361

pueli Chobaut, 1904, Geotrechus [Trechus] 310

pueli Antoine, 1920, Harpalus 383

pueli Barthe, 1916, Hydroporus 65

pueli Antoine, 1923, Mesolestes [Microlestes] 421

pueli Lutshnik, 1935, Pogonus 286

pueli Csiki, 1930, Pterostichus 512

puella Ledoux & Roux, 1998, Nebria 93

puellum Dejean, 1828, Agonum 453

puellus Lapouge, 1925, Carabus 145

puer A. Morawitz, 1886, Carabus 145

puer Kabak, 1997, Poecilus 485

puetzi Kleinfeld, Korell & Wrase, 1996, Cychrus 205

puetzi Kataev & Wrase, 1997, Harpalus 380

puetzi Sciaky & Wrase, 1997, Straneostichus 518

puetzi P. Moravec & Wrase, 1998, Trechus 338



pugetanum Fall, 1916, Bembidion 247

puigmalensis Lagar Mascaró, 1981, Geotrechus 310

puigmalensis Jeanne, 1976, Trechus 339

puisseguri Bourgin, 1953, Carabus 195

pukwonensis Deuve & Mourzine, 1993, Carabus 134

pulchella Hope, 1831, Heptodonta [Cicindela] 115

pulchella Dejean, 1831, Mastax [Brachinus] 217



pulchellum Lucas, 1846, Bembidion 252

pulchellum Marsham, 1802, Bembidion [Carabus] 251

pulchellum Panzer, 1796, Bembidion [Carabus] 244

pulchellus Brüggemann, 1873, Carabus 153

pulchellus Csiki, 1906, Carabus 170

pulchellus Klug, 1834, Copelatus [Agabus] 48

pulchellus Clark, 1863, Haliplus 32


pulchellus Clark, 1863, Hyphydrus 75

pulchellus Schaum, 1864, Microdaccus 422



pulchellus Heer, 1839, Platambus [Colymbetes] 44

pulchellus Faldermann, 1836, Pterostichus [Platysma] 505

pulchellus Putzeys, 1846, Trechus 338

pulchellus Carret, 1903, Pogonus 287

pulcher L. Redtenbacher, 1867, Callistoides [Chlaenius] 348

pulcher Kleinfeld, 1997, Carabus 141

pulcher Habu, 1978, Synuchus 546

pulcherrima Bates, 1873, Nebria 91

pulcherrimum Motschulsky, 1850, Bembidion [Peryphus] 268

pulcherrimus Magrini, Vanni & Zanon, 1997, Thaumastaphaenops 321

pulchra Baliani, 1943, Amara 564

pulchrinulus Reitter, 1900, Harpalus 380

pulchrior Maindron, 1906, Nebria 91

pulchripes Reitter, 1896, Carabus 133

pulchripes White, 1844, Tricondyla 118



pulchrum Gyllenhal, 1827, Bembidion 244

pulepilica Nakajima, 1993, Carabus [Acoptolabrus] 122

pulicaria Dejean, 1831, Tachyura [Bembidium] 279

pulicarius Sharp, 1882, Laccophilus 77



pulla Antoine, 1953, Amara [Celia] 555

pulla Jedlika, 1957, Amara [HN] 561

pulla Jacquelin du Val, 1852, Tachyura [Bembidium] 277

pullatus Zaitzev, 1908, Gyrinus 28



pullmani Casey, 1924, Amara 549

pullula Poppius, 1906, Amara 559

pullulus R. F. Sahlberg, 1844, Pterostichus [Feronia] 491

pullum Stephens, 1828, Agonum 453

pullus Semenov & Znojko, 1932, Carabus 147



pullus Delherm, 1891, Ophonus 391

pullus Gyllenhal, 1827, Pterostichus [Harpalus] 507

pulpani Kult, 1949, Amara 551

pulpani Fassati, 1955, Bembidion 260

pulpani Reška, 1965, Trechus 339

pulumuricus Cavazzuti, 1994, Carabus 148

pulverosus Stephens, 1828, Rhantus [Colymbetes] 47

pulverulentus P. Rossi, 1790, Harpalus [Carabus] 385

pulvillatus Breuning, 1934, Carabus [Coptolabrus] 144

pulvinatus Ménétriés, 1848, Harpalus 380

pulvinipenis Belousov & Kabak, 1999, Trechus 339

pumicatum Lapouge, 1907, Calosoma 119

pumicatus Panzer, 1796, Stomis [Carabus] 517

pumila Dejean, 1826, Cylindera [Cicindela] 111

pumila Andrewes, 1921, Siagona 218



pumilio Schaum, 1857, Acupalpus [Stenolophus] 398

pumilio Piochard de la Brûlerie, 1876, Amara [Zabrus] 566

pumilio Küster, 1846, Carabus 169

pumilio Duftschmid, 1812, Ocys [Elaphrus] 273

pumilio Dejean, 1829, Ophonus [Harpalus] 389

pumilio Dejean, 1828, Pterostichus [Feronia] 492

pumilio Jeannel, 1923, Trechus 339

pumilus Aubé, 1838, Bidessus [Hydroporus] 55

pumilus Kraatz, 1886, Carabus [Eutelocarabus] 183

pumilus Dejean, 1829, Harpalus 382

pumilus Sturm, 1818, Harpalus 380

pumilus Sharp, 1882, Hydrovatus 70

pumilus Jeannel, 1927, Trechus 339

pumoensis Deuve, 1998, Trechus 339

punctata Dokhtouroff, 1882, Cicindela 105

punctata Dejean, 1825, Cymindis 418

punctata Nietner, 1858, Eucolliuris [Odacantha] 443

punctata Gebler, 1843, Lebia 428

punctatella Motschulsky, 1844, Tachyura [Pericompsus] 280

punctatellum Motschulsky, 1844, Bembidion [Phyla] 252

punctatellus Reitter, 1894, Dicheirotrichus [Trichocellus] 403

punctatellus Duftschmid, 1812, Syntomus [Lebia] 432

punctaticollis Heller, 1923, Brachinus 216

punctaticollis N. Ito, 1999, Trichotichnus 394

punctatipennis A. Morawitz, 1862, Anisodactylus 361

punctatipennis Brancucci, 1984, Platambus 44

punctatipennis Tschitschérine, 1889, Pterostichus [Pseudadelosia] 513

punctatipennis Rambur, 1838, Typsiharpalus [Harpalus] 397

punctatissima Schaum, 1863, Abroscelis [Cicindela] 99

punctatissima Baliani, 1932, Amara 554

punctatissima Motschulsky, 1844, Cymindis 415

punctatissimus Fairmaire, 1859, Deltomerus [Patrobus] 283



punctatissimus Poppius, 1905, Hydroporus 64

punctatissimus Brancucci, 1983, Laccophilus 77

punctatissimus Breit, 1914, Leistus 83

punctatissimus Stephens, 1828, Ophonus 390

punctatissimus Baudi di Selve, 1894, Pseudaristus [Ditomus] 367

punctatoauratus Germar, 1824, Carabus 140

punctatocollis Gemminger & Harold, 1868, Poecilus 483

punctatocostatus Haury, 1885, Carabus 179

punctatocostatus Mandl, 1964, Carabus [HN] 173

punctatostriata Motschulsky, 1860, Amara [Amarocelia] 556

punctatostriata Putzeys, 1877, Epaphiopsis [Trechus] 309

punctatostriata Schaufuss, 1876, Nebria 87

punctatostriatum Motschulsky, 1844, Bembidion [Peryphus] 250


punctatostriatus Fischer von Waldheim, 1829, Broscus [Cephalotes] 236

punctatostriatus Bates, 1873, Carabus 177

punctatostriatus Heyden, 1886, Carabus 182

punctatostriatus Chaudoir, 1856, Chlaenius 351



punctatostriatus Motschulsky, 1864, Demetrias 419

punctatostriatus L. Redtenbacher, 1843, Distichus [Scarites] 231

punctatostriatus Dejean, 1829, Harpalus 380

punctatostriatus Fairmaire, 1859, Laemostenus [Pristonychus] 539

punctatostriatus Duftschmid, 1812, Paradromius [Lebia] 424

punctatostriatus Stephens, 1828, Poecilus 484

punctatostriatus Motschulsky, 1859, Pterostichus [Steroderus] 506

punctatostriatus Jedlika, 1953, Taphoxenus 544

punctatostriatus Heyden, 1883, Tapinopterus 520

punctatostriatus Brullé, 1835, Zabrus 571

punctatosulcatus Chaudoir, 1855, Oxylobus 231



punctatulus J. R. Sahlberg, 1889, Hydroporus 60

punctatulus Fabricius, 1792, Licinus [Carabus] 442



punctatulus Duftschmid, 1812, Ophonus [Carabus] 390

punctatus Scopoli, 1763, Acilius [Dytiscus] 49

punctatus Dejean, 1828, Broscus [Cephalotes] 236



punctatus Géhin, 1885, Carabus 156

punctatus Laporte, 1834, Carabus 157



punctatus Motschulsky, 1844, Chlaenius 350

punctatus Guérin‑Méneville, 1847, Crasodactylus 362

punctatus Heinz & Ledoux, 1987, Deltomerus 283

punctatus Dejean, 1825, Dyschirius [Clivina] 227



punctatus Olivier, 1795, Dytiscus 52

punctatus Motschulsky, 1844, Elaphrus 207



punctatus Zoubkoff, 1837, Eretes [Eunectes] 53

punctatus Marsham, 1802, Hygrotus [Dytiscus] 72

punctatus Dejean, 1828, Laemostenus [Pristonychus] 539



punctatus Reitter, 1885, Leistus 84

punctatus Dejean, 1828, Molops 474

punctatus Jedlika, 1934, Platynus [Colpodes] 465

punctatus L. Redtenbacher, 1843, Pterostichus [Feronia] [ND] 516

punctatus Geoffroy, 1785, Rhantus [Dyticus] 47

punctatus Hoppe, 1795, Rhantus [Dytiscus] 46

punctatus O. F. Müller, 1776, Suphrodytes [Dytiscus] 70

punctibasis Jedlika, 1957, Amara 568

punctibasis Reitter, 1894, Anchomenus [Agonum] 456

punctibasis Poppius, 1906, Harpalus 379

punctibasis Chaudoir, 1868, Poecilus [Feronia] 483



punctibasis Fairmaire, 1889, Pristosia [Calathus] 531

punctibasis Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 542



puncticeps Seidlitz, 1887, Hygrotus [Coelambus] 71

puncticeps Fairmaire & Laboulbène, 1854, Leistus 83

puncticeps Stephens, 1828, Ophonus 390

puncticeps Bedel, 1907, Paradromius [Dromius] 424

puncticeps Andrewes, 1919, Planetes 359



puncticeps C. G. Thomson, 1867, Poecilus [Feronia] 482

puncticeps Schaum, 1864, Zabrus 572



puncticolle L. Dufour, 1843, Bembidion 267

puncticollis Coquerel, 1859, Acupalpus [Bradycellus] 399

puncticollis Dejean, 1828, Amara 563

puncticollis Rylands, 1841, Amara [HN] 548

puncticollis Kraatz, 1878, Carabus [Orinocarabus] 180

puncticollis Dejean, 1826, Chlaenius 354

puncticollis Chaudoir, 1855, Distichus [Taeniolobus] 231

puncticollis Chaudoir, 1860, Neocollyris [Collyris] 117

puncticollis Küster, 1848, Notiophilus 98

puncticollis Mannerheim, 1844, Odacantha 444

puncticollis Lucas, 1842, Olisthopus 461

puncticollis Paykull, 1798, Ophonus [Carabus] 390

puncticollis Jedlika, 1934, Platynus [Colpodes] 465

puncticollis Dejean, 1828, Poecilus [Feronia] 480



puncticollis Motschulsky, 1844, Poecilus 483

puncticollis Brullé, 1832, Zabrus [HN] 571

puncticollis Dejean, 1831, Zabrus 572

punctidorsis Reitter, 1899, Dicheirotrichus [Trichocellus] 403



punctifrons Chaudoir, 1850, Poecilus [Feronia] 480

punctifrons Reitter, 1908, Pogonus 287

punctifrons Fairmaire, 1866, Zabrus 571

punctiger Semenov, 1897, Carabus 201



punctiger Stephens, 1828, Harpalus 381

punctiger Reiche & Saulcy, 1855, Platyderus [Feronia] 523

punctiger J. R. Sahlberg, 1880, Pterostichus [Feronia] 492

punctigera Kraatz, 1890, Cassolaia [Cicindela] 101

punctilabris W. S. MacLeay, 1825, Pseudognathaphanus [Harpalus] 363

punctipennis J. R. Sahlberg, 1880, Agabus [Gaurodytes] 35

punctipennis Reitter, 1889, Amara [Harpalodema] 566

punctipennis Jeannel, 1927, Anchotrechus 290

punctipennis Gebler, 1833, Anisodactylus 361

punctipennis Germar, 1824, Calathus 525

punctipennis Taschenberg, 1883, Cybister [Trogus] 50

punctipennis Reitter, 1901, Cychrus 204

punctipennis Casey, 1914, Dicheirotrichus [Trichocellus] 403

punctipennis Putzeys, 1846, Dyschirius 227

punctipennis Baehr, 1998, Gunvorita 573

punctipennis Mulsant, 1852, Harpalus 380

punctipennis C. G. Thomson, 1867, Hydaticus 54


punctipennis J. R. Sahlberg, 1880, Hydroporus 64

punctipennis Fauvel, 1882, Leistus 83

punctipennis Sharp, 1882, Methles 75

punctipennis Sharp, 1884, Orectochilus 28



punctipennis Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 542

punctipennis Chaudoir, 1846, Zabrus [Eutroctes] 569



punctiventre Reiche & Saulcy, 1855, Calosoma 119

punctiventris Schaum, 1864, Zabrus 571



punctulata Dejean, 1828, Amara 556

punctulata Chaudoir, 1876, Drypta 358

punctulatum Drapiez, 1820, Bembidion 266



punctulatus Reiche & Saulcy, 1855, Carabus 186

punctulatus Voet, 1778, Carabus [Buprestis] 187

punctulatus Chaudoir, 1844, Dixus [Aristus] 24, 365

punctulatus Fabricius, 1777, Dytiscus 52

punctulatus Penecke, 1901, Haliplus 32

punctulatus Chaudoir, 1857, Hololeius 356

punctulatus Wesmael, 1835, Notiophilus 97

punctulatus Dejean, 1828, Olisthopus 461

punctulatus W. S. MacLeay, 1825, Platymetopus [Harpalus] 393

punctulatus Schaller, 1783, Poecilus [Carabus] 483

punctulatus Dejean, 1828, Pogonus 286

punctulatus Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 542

punctulatus Lutshnik, 1922, Siopelus [Trichotichnus] 394



punctulicollis Bates, 1889, Platymetopus 393

punctulipenne Bates, 1878, Bembidion 258



punctum Gebler, 1829, Hygrotus [Hydroporus] 72

punctum Wiedemann, 1823, Scarites 232

punicum K. Daniel & J. Daniel, 1898, Parazuphium [Zuphium] 574

punicus Normand, 1933, Deltomerus 283

punicus Bedel, 1899, Dicheirotrichus 402

punjabensis Vazirani, 1970, Rhantus 46

pupieri Jeanne, 1988, Orthomus 477



pupillata Andrewes, 1924, Lebidia 426

pupulus A. Morawitz, 1889, Carabus 123

pupulus Busi & Rocca, 1983, Duvalius 301

puritanum Hayward, 1897, Bembidion 251

purkynei Jedlika, 1967, Agonum 455



purkynei Jedlika, 1967, Amara 548

purkynei Jedlika, 1946, Archastes 79



purkynei Jedlika, 1932, Bembidion 271

purkynei Maan, 1935, Calathus 527

purkynei Jedlika, 1946, Cymindis 414

purkynei Obenberger, 1917, Elaphrus 207

purkynei Jedlika, 1933, Lebia 430



purkynei J. Müller, 1933, Molops 473

purkynei Jedlika, 1934, Platynus [Colpodes] [HN if Platynus] 465

purkynei Obenberger, 1917, Platynus 463

purkynei Jedlika, 1931, Pristosia [Laemostenopsis] 531

purkynei Jedlika, 1928, Tapinopterus 520

purkynei Maan, 1940, Zabrus 570

purpurarius Machado, 1992, Paradromius 424

purpurascens Motschulsky, 1859, Amara [Celia] 565

purpurascens Fabricius, 1787, Carabus 166

purpurascens Dejean, 1828, Poecilus [Feronia] 480

purpurata Klug, 1834, Neocollyris [Collyris] 117

purpuratus Sirguey, 1931, Carabus 139

purpuratus Sturm, 1815, Carabus 173

purpuratus Schaufuss, 1865, Laemostenus [Cryptotrichus] 536

purpuratus Heer, 1838, Pterostichus 504

purpureodiscoides Kleinfeld, 1999, Carabus 142



purpureus Kraatz, 1876, Carabus 186

purpureus Oberthür, 1885, Carabus 139

purpuricollis Kraatz, 1886, Carabus 171

purpurifer Reitter, 1885, Carabus 151

purpuripenne Reitter, 1891, Calosoma 120

purpuripennis Kraatz, 1886, Carabus 171

purroyi Salgado Costas, 1987, Apoduvalius 295

puschkini M. F. Adams, 1817, Carabus 201

pusilla Uéno, 1955, Nippononebria [Nebria] 95



pusilla Péringuey, 1896, Perigona [Extromus] 449

pusillimus Péringuey, 1896, Polyderis [Tachys] 274

pusillum Schaum, 1858, Agonum [Anchomenus] 451

pusillum Stephens, 1828, Agonum 453

pusillum Gyllenhal, 1827, Bembidion 249

pusillum Stephens, 1828, Bembidion [Tachys] 265

pusillus Lapouge, 1915, Carabus 149

pusillus Sharp, 1882, Copelatus 48

pusillus Dejean, 1825, Dyschirius [Clivina] 227

pusillus Jeannel, 1928, Geotrechus 311

pusillus Motschulsky, 1850, Harpalus [Erpeinus] 380



pusillus Jeannel, 1947, Hydraphaenops [Aphoenops] 311

pusillus Fabricius, 1781, Hydroglyphus [Dytiscus] 56

pusillus G. R. Waterhouse 1833, Notiophilus 96

pusillus Schreber, 1759, Notiophilus [Cicindela] 96

pusillus Jeannel, 1923, Perileptus 289

pusillus Dejean, 1828, Pterostichus [Feronia] 492

pusio Semenov, 1898, Carabus 191




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