Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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rechingeri Csiki, 1915, Celaenephes 410

recidivus Gschwendtner, 1923, Hydroporus 64

recondita W. Hansen, 1968, Desera 358

reconditus Ivanovs, 1993, Carabus 134

recta Kraatz, 1890, Cassolaia [Cicindela] 101

recta Motschulsky, 1844, Cylindera [Cicindela] 109

recta Walker, 1858, Pseudoclivina [Clivina] 221

rectangula Motschulsky, 1850, Cymindis [Tarus] 411

rectangula Chaudoir, 1876, Glycia 409


rectangularis Klug, 1832, Cylindera [Cicindela] 111

rectangularis Schaufuss, 1882, Typhlotrechus [Anophthalmus] 344

rectangulata Uéno & Saito, 1991 6, Jujiroa 459

rectangulum Jacquelin du Val, 1852, Bembidion 265

rectangulus Cuvier, 1833, Abax [Carabus] 470

rectangulus Reitter, 1883, Amblystomus 360

rectangulus Morvan, 1992, Deltocolpodes 457

rectangulus Sharp, 1882, Hydaticus 53

rectangulus C. G. Thomson, 1870, Ophonus 390

rectangulus Fairmaire, 1859, Orthomus [Feronia] 477

rectangulus N. Ito, 1994, Parophonus 392

rectangulus Lindroth, 1956, Synuchus 546

rectangulus Chaudoir, 1868, Tapinopterus [Feronia] 518

rectestriatus Bokor, 1921, Duvalius [Trechus] 300

recticaudis Kurnakov, 1961, Lindrothius 529

recticollis Schaufuss, 1865, Laemostenus [Pristonychus] 539

rectidens Wasmann, 1922, Paussus 210

rectifrons Bates, 1892, Distichus [Distichus] 231



rectilinea Roeschke, 1891, Cicindela 108

rectilobata Uéno, 1978, Epaphiopsis 308



rectiperlatus Maindron, 1906, Carabus [Coptolabrus] 143

rectitibius Leoni, 1907, Laemostenus [Pristonychus] 537

rectoimpressus Motschulsky, 1850, Carabus 186

rectokoreanus Ishikawa & Kim, 1985, Carabus [Leptocarabus] 159



rectus Babington, 1841, Agabus [Colymbetes] 41

rectus Wollaston, 1862, Calathus 526

rectus Sharp, 1882, Methles 75

rectus Uéno, 1958, Trechiama 324



recurvata Kraatz, 1890, Cephalota [Cicindela] 102

recurvicollis Andrewes, 1925, Porotachys [Tachys] 275

recurvus Putzeys, 1866, Dyschirius 230

recurvus Marsham, 1802, Hygrotus [Dytiscus] 73

redikortzewi Semenov & Breuning, 1931, Carabus 145



redikorzevi Lutshnik, 1934, Amara 566

redikorzevi Lutshnik, 1934, Harpalus 372

redoni Antoine, 1928, Deltomerus 283



redtenbacheri K. Daniel, 1902, Bembidion 245

redtenbacheri Géhin, 1876, Carabus 181

redtenbacheri I. Frivaldszky von Frivald & J. Frivaldszky, 1857, Duvalius [Anophthalmus] 300

redtenbacheri Schaufuss, 1865, Laemostenus [Antisphodrus] 534

redtenbacheri W. Horn, 1894, Neocollyris [Collyris] 117

reductecostatus Mandl, 1964, Carabus 173

reductelineata W. Horn, 1912, Cylindera [Cicindela] 110

reductemaculata Mandl, 1934, Calomera [Cicindela] 100

reductemaculata Nakane & Ohkura, 1956, Coptodera [Coptoderina] 435


reductescripta W. Horn, 1912, Abroscelis [Cicindela] 99

reductesignata Mandl & Wiesner, 1975, Prothyma 115

reductoculatus Uéno, 1992, Trechiama 323



reductus G. Hauser, 1921, Carabus [Acoptolabrus] 123

reductus J. Müller, 1930, Carabus 198

reductus Roeschke, 1897, Carabus 198



reductus Walker, 1858, Catascopus 435

reductus Jedlika, 1935, Chlaenius 349

reductus J. MüMotschulsky, 1866, Dyschirius 226

reductus Hlisnikovský, 1955, Hyphydrus 74

reflexa Putzeys, 1866, Amara [Curtonotus] 561

reflexa Bates, 1883, Nebria 94

reflexicollis Motschulsky, 1844, Amara 568

reflexicollis Uéno, 1977, Bhutanotrechus 296

reflexicollis Faldermann, 1839, Calathus 526

reflexicollis Gebler, 1832, Poecilus 483



reflexicollis Bates, 1883, Porotachys [Tachys] 275

refleximargo Machado, 1992, Calathus 528

refleximargo Reitter, 1894, Reflexisphodrus [Pseudotaphoxenus] 543

refleximargo Chaudoir, 1873, Trymosternus 410

reflexipennis Semenov, 1889, Reflexisphodrus [Taphoxenus] 543

reflexus Schaum, 1858, Calathus 526

reflexus Putzeys, 1878, Harpalus 381

reflexus Chaudoir, 1861, Laemostenus [Pristonychus] 536

reflexus Semenov, 1889, Leistus 81

reflexus Schaum, 1862, Zabrus 571

refulgens Reiche, 1876, Amara [Triaena] 568



refulgens Chaudoir, 1846, Carabus 175

regale Andrewes, 1922, Bembidion 250



regalis Petri, 1903, Agabus 39

regalis Fischer von Waldheim, 1820, Carabus 172

regalis Schmidt‑Göbel, 1846, Catascopus 435

regeli A. Morawitz, 1886, Carabus 178

regeli Ballion, 1878, Harpalus 386

regeli Tschitschérine, 1894, Pterostichus [Feronia] 502

regelianum A. Morawitz, 1886, Calosoma [Callisthenes] 119

regimbarti Zaitzev, 1906, Agabus 36



regimbarti Wilke, 1920, Cybister 50

regimbarti Peyerimhoff, 1931, Gyrinus 28

regimbarti Zaitzev, 1908, Haliplus 31

regimbarti Gschwendtner, 1936, Hydroglyphus [Bidessus] 56

regimbarti Sharp, 1884, Orectochilus 28

regimbarti A. I. Jakovlev, 1896, Rhantus 47

reginae Marcilhac, 1993, Aristochroodes [Pterostichus] 471

reginae Deuve, 1997, Carabus 137


regisborisi Buresch, 1926, Duvalius [Paraduvalius] 308

regisborisi Buresch, 1926, Duvalius [Paraduvalius] 308

regismontium Schweiger, 1966, Amara [Bradytus] 553

regismontium Netolitzky, 1943, Bembidion 254



regismontium Schweiger, 1964, Carabus 131

registanica Hieke, 1996, Amara 552

regiszogui Meschnigg, 1936, Duvalius 305

regularis Fischer von Waldheim, 1823, Carabus [Cechenus] 184

regularis Wissmann, 1848, Carabus 171

regularis Motschulsky, 1844, Harpalus 376

regularis Sharp, 1882, Hydroporus 64



regularis Fischer von Waldheim, 1823, Pterostichus [Myosodus] 501

regularis Motschulsky, 1844, Pterostichus [Lyperopherus] 499

regularis Putzeys, 1870, Trechus 339

regulus C. A. Dohrn, 1882, Carabus 192

reichardti Lutshnik, 1934, Amara 562

reichardti Lutshnik, 1930, Bembidion 262



reichardti Lutshnik, 1930, Cymindis 416

reichardti Lutshnik, 1930, Pseudotaphoxenus [Taphoxenus] 542



reichei Aubé, 1837, Agabus 37

reichei Coquerel, 1859, Amara 566



reichei Guérin‑Méneville, 1842, Calosoma [Callisthenes] 118

reichei Aubé, 1838, Cybister 50



reichei Guérin‑Méneville, 1859, Graphipterus 359

reichei Desbrochers des Loges, 1867, Harpalus 385

reichei Jakobson, 1907, Harpalus [HN!] 370

reichei Kraatz, 1858, Percus 478

reichei Chaudoir, 1875, Platytarus [Cymindoidea] 410

reicheianus Peyron, 1858, Poecilus [Feronia] 483

reichenbachii Schaufuss, 1861, Laemostenus [Pristonychus] 539

reichii Dejean, 1826, Nebria 87



reichii Waltl, 1835, Poecilus 484

reinigi Netolitzky, 1934, Bembidion 249



reinigi Schauberger, 1934, Harpalus 382

reischitzi Mandl, 1955, Carabus 169

reiseri Apfelbeck, 1902, Bembidion 257

reiseri Heinz, 1980, Carabus 193



reiseri Mandl, 1970, Cicindela 107

reiseri Apfelbeck, 1904, Cychrus 204

reiseri Ganglbauer, 1891, Duvalius [Trechus] 306

reiseri Apfelbeck, 1904, Molops 474

reiseri Ganglbauer, 1889, Pterostichus 516

reissi Mihók, 1911, Duvalius [Anophthalmus] 299

reissi Ganglbauer, 1911, Laemostenus 535

reissi J. Müller, 1932, Pterostichus 496


reitteri Ragusa, 1886, Acupalpus 399

reitteri Ragusa, 1885, Agonum 451

reitteri Tschitschérine, 1894, Amara 554



reitteri Wanka, 1915, Anthracus 400

reitteri Roeschke, 1897, Calosoma 121



reitteri Appel, 1916, Carabus 184

reitteri Retowski, 1885, Carabus [Plectes] 132

reitteri Jakobson, 1906, Chlaenius 352

reitteri W. Horn, 1897, Cicindela 106

reitteri Liebke, 1927, Cymindis 416

reitteri Miller, 1881, Duvalius [Anophthalmus] 307

reitteri Kult, 1949, Dyschirius 227

reitteri G. Ochs, 1942, Gyrinus 28



reitteri Zaitzev, 1908, Hygrotus [Coelambus] 71

reitteri A. Fiori, 1913, Leistus 84

reitteri Jakobson, 1906, Leistus 82

reitteri Holdhaus, 1912, Microlestes 423

reitteri Rybiski, 1902, Nebria 87

reitteri Spaeth, 1900, Notiophilus 97

reitteri Jedlika, 1937, Pristosia [Eucalathus] 531

reitteri Stark, 1889, Pterostichus 505

reitteri Koenig, 1904, Scarites 234

reitteri Csiki, 1928, Tachys 276

reitteri Jedlika, 1952, Taphoxenus 544

reitteri R. T. Bell, 1977, Yamatosa [Yamatoa] 78



reitteri Apfelbeck, 1904, Zabrus 570

reitterianus Breuning, 1932, Carabus 190

reitterii Seidlitz, 1887, Orectochilus 28

rektoriki Kult, 1954, Amara 558

relator Reitter, 1900, Harpalus 380

relicta Pace, 1976, Crowsoniella 25

relicta Breit, 1914, Oreonebria [Nebria] 96

relictum Apfelbeck, 1904, Bembidion 246

relictum Weissmandl, 1936, Broscosoma 235

relictum Apfelbeck, 1918, Calosoma 121



relictus Krausse, 1910, Carabus 161

relictus Semenov, 1898, Carabus 129



relictus Binaghi, 1959, Leistus 83

relictus Semenov, 1900, Leistus 83

relictus Jeanne, 1985, Mesolestes [Neomesolestes] 421

relictus Agazzi, 1959, Orotrechus 317

relictus Semenov & Bogachev, 1938, Panagaeus 447



relictus Neresheimer & Wagner, 1927, Patrobus 285

relictus Uéno, 1957, Phreatodytes 35

relictus Aubry, 1979, Pterostichus [Pyreneorites] 489


religiosus Uéno, 1957, Awatrechus 296

religiosus Normand, 1938, Tachys 276

relucens Mannerheim, 1853, Amara [Celia] 565

relucens Andrewes, 1923, Dicranoncus [Anchomenus] 458

relucens Bates, 1883, Trichotichnus 397

remboides Solsky, 1874, Harpalus 381

remondi Morvan, 1997, Andrewesius 456

remondi Morvan, 1998, Diacanthostylus 458

remondi Morvan, 1999, Xestagonum 468

remondianus Deuve, 1996, Carabus 189

remondorum Deuve, 1995, Carabus 138

remondorum Lassalle & Marcilhac, 1999, Reflexisphodrus 543



remota C. Zimmermann, 1832, Amara [Celia] 565

remotepunctatus Putzeys, 1866, Dyschirius 224

remotestriata Dejean, 1828, Amara 565

remotus J. R. Sahlberg, 1913, Agabus 39

remotus Reitter, 1888, Carabus [Trachycarabus] 147

remotus Marsham, 1802, Dyschirius [Carabus] 229

renardi Fleutiaux, 1890, Calomera [Cicindela] 99

renardi Chaudoir, 1846, Carabus 157

renardi Severin, 1890, Hyphydrus 75

renati Maindron, 1899, Hypaetha [Cicindela] 112

renati Jeannel, 1922, Trechus 337

renaudeti A. Gaudin, 1947, Geotrechus 310

rendschmidtii Germar, 1839, Abax [Feronia] 469

renei W. Horn, 1897, Hypaetha [Cicindela] 112

renei Belousov, 1990, Trechus 339

renitens Bates, 1886, Loxoncus [Anoplogenius] 404

renoicus Casey, 1914, Harpalus 377

repanda Zoubkoff, 1833, Cymindis 417

repandum J. R. Sahlberg, 1875, Bembidion 262

repandus Poppius, 1906, Pterostichus [Cryobius] 492

repercussus Drapiez, 1819, Carabus 173

reperiendus Deuve, 1992, Carabus 150

repletus Reitter, 1884, Carabus 194

resiczabanyensis Csiki, 1913, Carabus 196

resli Bulirsch, 1996, Dyschirius 225

resplendens Breuning, 1934, Carabus 193

resplendens Dokhtouroff, 1888, Cicindela 106

ressli Mandl, 1961, Carabus 187

ressli Wewalka, 1974, Nebrioporus [Potamonectes] 66



resslianus Mandl, 1963, Carabus 186

restricta Fischer von Waldheim, 1828, Cicindela 106



restrictus Sharp, 1882, Laccophilus 77

restrictus Fischer von Waldheim, 1829, Ophonus 392



retezari Gottwald, 1980, Carabus 200

reticulatum Iablokoff‑Khnzorian, 1976, Bembidion 259

reticulatum Fabricius, 1787, Calosoma [Carabus] 118



reticulatus Kirby, 1837, Agabus [Colymbetes] 35

reticulatus Schaum, 1857, Apristus 431



reticulatus Hampe, 1852, Carabus 198

reticulatus Zamotajlov, 1990, Chinapenetretus [Parapenetretus] 281

reticulatus Motschulsky, 1844, Chlaenius 350

reticulatus Brinck, 1940, Gyrinus 28

reticulatus Fabricius, 1792, Hygrotus [Dytiscus] 73

reticulatus Netolitzky, 1917, Ocys [Bembidion] 273

reticulatus Chaudoir, 1842, Platyderus [Lissotarsus] 523

reticulatus Jacquelin du Val, 1857, Poecilus [Feronia] 481

reticulatus Schaum, 1857, Pogonus 287

reticulatus Belousov, 1998, Porocimmerites 318

reticulatus Lindroth, 1956, Synuchus 546



reticulosus Falkenström, 1939, Scarodytes 68

retipenne J. Müller, 1918, Bembidion 253



retipennis Maan, 1933, Trichotichnus 396

retowskianus Mandl, 1955, Carabus 199

retowskianus Reitter, 1887, Harpalus 374

retowskii Reitter, 1888, Carabus [Neoplectes] 132

retowskii Heyden, 1883, Harpalus [Pseudoophonus] 382

retowskii Reitter, 1887, Harpalus 374

retowskii Reitter, 1897, Notiophilus 97

retractus Joseph, 1870, Laemostenus [Sphodrus] 535

retractus Walker, 1858, Mochtherus [Panagaeus] 437

retrofasciata Motschulsky, 1864, Lebia 430

retrospinosa Heyden, 1885, Nebria 87

revelierei Sharp, 1882, Hydroporus 62

revelieri Perris, 1866, Hypotyphlus [Scotodipnus] 239



revelieri Mulsant & Guillebeau, 1855, Leistus 82

rex Gottwald, 1980, Carabus 200

reyi Coiffait, 1953, Trechus 325

reymondi Antoine, 1957, Amara [Celia] 555

reymondi Colas & A. Gaudin, 1935, Aphoenops 294

reymondi Antoine, 1963, Brachinus 214

reymondi Antoine, 1953, Graphipterus 360

reymondi Colas, 1950, Licinus 442

reymondi Antoine, 1951, Nebria 93

reynieri Charet, 1992, Carabus 158

rhadamanthus Linder, 1860, Aphoenops [Anophthalmus] 295



rhaetica Heer, 1837, Amara 553

rhaetica K. Daniel & J. Daniel, 1890, Nebria 93

rhaeticum Heer, 1837, Bembidion 269


rhaeticus Heer, 1837, Leistus 83

rhaeticus Heer, 1837, Pterostichus 511

rhanis Tschitschérine, 1902, Pterostichus [Platysma] 502

rhantoides Sharp, 1882, Hydaticus 54



rhatica Antoine, 1936, Amara 555

rhatica Antoine, 1936, Cymindis 414

rhaticus Antoine, 1941, Calathus 528

rhiannon Morvan, 1996, Xestagonum 468

rhilensis Kraatz, 1876, Carabus 152

rhilensis Roeschke, 1907, Cychrus 205

rhilensis Apfelbeck, 1904, Molops 472

rhilensis J. Frivaldszky, 1879, Nebria 87

rhilensis Rottenberg, 1874, Pterostichus 485

rhilensis Kaufmann, 1884, Trechus 339

rhinopterus Hampe, 1852, Carabus 198

rhisobius Gistel, 1857, Dromius [ND] 421



rhodanensis Schaefer, 1952, Graptodytes 59

rhodanicus Lapouge, 1916, Carabus 171

rhodesianus Csiki, 1928, Polyderis [Tachys] 274

rhodius Straneo, 1935, Tapinopterus [Pterostichus] 520

rhododendron Deuve & Imura, 1991, Carabus 191

rhodopeius Jeannel, 1921, Trechus 339

rhodopense Apfelbeck, 1902, Bembidion 246

rhodopensis Apfelbeck, 1904, Carabus 179

rhodopensis Apfelbeck, 1904, Molops 474

rhodopensis Horvatovich, 1973, Nebria 86

rhodopensis Apfelbeck, 1904, Zabrus 570

rhodopus Schauberger, 1930, Harpalus 375

rhodoterena Tschitschérine, 1903, Cicindela 107

rhombophora Andrewes, 1925, Tachyura [Tachys] 277

rhytidera Chaudoir, 1876, Zolotarevskyella [Blechrus] 426

rhytidopteroides W. Horn, 1924, Setinteridenta [Cicindela] 114

rialensis Guillebeau, 1890, Hypotyphlus [Microtyphlus] 239

ribagorzanus Bolívar y Pieltain, 1919, Hypotyphlus [Scotodipnus] 239

ribbei Putzeys, 1877, Broscosoma 235



ribeiranus Franz, 1981, Trechus 329

riberai Fery & Hosseinie, 1998, Deronectes 59

richardi Knirsch, 1925, Duvalius [Duvalius] 307

richardsi Ali, 1967, Chlaenius 350

richteri Fassati, 1961, Bembidion 20, 249

richteri Jedlika, 1961, Brachinus [Cymindis] 214

richteri M. Quedenfeldt, 1887, Carabus 161

richteri Kryzhanovskij, 1951, Eobroscus 237

richteri Farka, 1995, Leistus 18, 81



richteri Shilenkov, 1999, Leistus 18, 83

richteri Morvan, 1996, Platynus [Batenus] 463

richterianum Müller‑Motzfeld, 1988, Bembidion 246

richterianus Kirschenhofer, 1987, Oxycentrus 392

ricinus Wewalka, 1979, Hydaticus 54

rickmersi Reitter, 1898, Bembidion 262

riedelii Ménétriés, 1832, Carabus 169



rifense Antoine, 1941, Bembidion 269

rifensis Cobos, 1961, Eocarterus 365

rifensis Alluaud, 1932, Siagona 218

rifensis Coiffait, 1969, Typhlocharis 241

rifensis Antoine, 1941, Zabrus 572

riffaudi Gory, 1833, Pheropsophus [Brachinus] 218

riffensis Fairmaire, 1872, Carabus 168

riffensis Antoine, 1933, Pterostichus 505

riffensis Mateu, 1954, Singilis 438

rigouti Basquin & Darge, 1986, Carabus 158

rigoutianus Deuve & Li, 1998, Carabus 122

rigrevanensis Coiffait, 1962, Laemostenus [Pristonychus] 538

rilvensis H. Kolbe, 1887, Carabus 166

rimanus Schauberger, 1936, Trichotichnus 397

rimmae Deuve, 1993, Laemostenus 537

rimoganensis Miwa, 1934, Rhyzodiastes [Clynidium] 78

rimosus Guignot, 1952, Copelatus 48



rimskyikorsakovi Lutshnik, 1933, Bembidion 258

rinaldi Toledano, 1999, Bembidion 245



rinaldoi Meggiolaro, 1959, Carabus 184

riojai Vidal y López, 1919, Cicindela 106

rionicum Müller‑Motzfeld, 1983, Bembidion 246

rionicus Belousov, 1992, Carabus 181

riozoi Uéno, 1999, Ishikawatrechus 312



riparia Dejean, 1822, Cicindela 108

riparia Stephens, 1827, Cicindela 105

riparioides Korell, 1965, Cicindela 107

riparius Mannerheim, 1844, Dyschirius 227

riparius Linné, 1758, Elaphrus [Cicindela] 207

riparius Dejean, 1828, Pogonus 287

ripensis Motschulsky, 1866, Pterostichus [Argutor] 508



riphaeus Motschulsky, 1844, Carabus 199

ripicola L. Dufour, 1820, Bembidion 250

rishirensis Nakane, 1957, Carabus 134

rishiriensis Habu, 1972, Nebria 89



rishwani Makhan, 1999, Haliplus 31

risnjakensis Depoli, 1926, Molops 473

ritchiei Vigors, 1825, Neolaphyra [Cicindela] 114



ritchii Lucas, 1846, Neolaphyra [Cicindela] 114

ritsemae Régimbart, 1880, Canthydrus [Hydrocanthus] 34

ritsemae Oberthür, 1883, Selina 407

rivalis Gyllenhal, 1827, Oreodytes [Hyphydrus] 68

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