Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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rufipes Fabricius, 1801, Dineutus [Gyrinus] 26

rufipes Dejean, 1825, Dyschirius [Clivina] 229

rufipes DeGeer, 1774, Harpalus [Carabus] 385

rufipes Motschulsky, 1844, Harpalus 381

rufipes Dejean, 1825, Lebia 428

rufipes Chaudoir, 1843, Leistus 84

rufipes Ganglbauer, 1891, Leistus [HN] 84

rufipes Chaudoir, 1843, Molops 474



rufipes Schilsky, 1889, Molops [HN] 473

rufipes Goeze, 1777, Nebria [Carabus] 92

rufipes Motschulsky, 1864, Neocollyris [Collyris] 117

rufipes Chaudoir, 1844, Notiophilus 97

rufipes Curtis, 1829, Notiophilus 97



rufipes Dejean, 1828, Platynus [Anchomenus] 464

rufipes Dejean, 1828, Pterostichus [Feronia] 513



rufipes Motschulsky, 1859, Pterostichus [Lyperopherus] 497

rufipes Fabricius, 1792, Siagona [Cucujus] 218

rufiscapis Bates, 1873, Apristus 431

rufiscapus Gebler, 1833, Harpalus 381



rufitarsis Fischer von Waldheim, 1844, Acinopus 368

rufitarsis Duftschmid, 1812, Harpalus [Carabus] 381

rufitarsis Rambur, 1838, Harpalus 375

rufitarsis Curtis, 1839, Laemostenus [Pristonychus] 536

rufitarsis Westwood, 1833, Paussus 210

rufitarsis Fischer von Waldheim, 1823, Pseudotaphoxenus [Sphodrus] 542

rufitarsis Dejean, 1828, Pterostichus [Feronia] 494

rufitarsoides Schauberger, 1934, Harpalus 381

rufithorax Pecchioli, 1837, Apotomus 237

rufithorax C. R. Sahlberg, 1827, Dicheirotrichus [Harpalus] 403

rufithorax Jedlika, 1936, Dolichus 530

rufithorax Kirschenhofer, 1996, Lachnoderma 438

rufithorax Jedlika, 1960, Stenolophus 405

rufiventris Régimbart, 1895, Cybister 50

rufiventris Walker, 1858, Miscelus [Cymindis] 437

rufoabdominalis Kataev, 1997, Stenolophus 405



rufoaenea Dejean, 1828, Amara 565

rufoaeneus Chaudoir, 1843, Dyschirius 228

rufoaeneus Dejean, 1828, Pogonistes [Pogonus] 286

rufobrunneum Heer, 1838, Bembidion 269

rufocastaneus Wollaston, 1862, Calathus 526



rufocincta C. R. Sahlberg, 1827, Amara [Harpalus] 557

rufocuprescens Deuve, 1993, Carabus 178



rufocupreus Breuning, 1943, Carabus 134

rufofemoralis Maan, 1941, Leistus 18, 83

rufofemoratus J. R. omnicki, 1892, Carabus [Trachycarabus] 199

rufofemoratus Motschulsky, 1866, Carabus 171

rufofuscus Jedlika, 1940, Synuchus [Parcalathus] 546

rufoguttatum Netolitzky, 1943, Bembidion 259

rufolimbata Maindron, 1905, Glycia 409

rufolimbatus Jedlika, 1935, Stenolophus [Egadroma] 405

rufomaculatum Motschulsky, 1845, Bembidion [Notaphus] 266

rufomarginata Fischer von Waldheim, 1828, Nebria 92

rufomarginatus Duftschmid, 1812, Leistus [Carabus] 84



rufomarginatus Curtis, 1827, Pterostichus [Argutor] 486

rufomarginatus Ménétriés, 1832, Zabrus 572

rufoniger Clark, 1863, Hydrovatus [Hyphydrus] 70

rufonitens Fairmaire, 1866, Pterostichus [Feronia] 492

rufopedestris Mandl, 1971, Neocollyris 116

rufopiceus Heyden, 1890, Pterostichus 486

rufosuturalis Mandl, 1954, Cosmodela [Cicindela] 108

rufotarsatulus Mandl, 1992, Chlaenius 348

rufotestacea Jedlika, 1934, Parena 434

rufotestaceum Jedlika, 1960, Agonum 455

rufotestaceum Pic, 1904, Cymbionotum [Graniger] 235

rufotibiellum Fairmaire, 1888, Bembidion 250



rufotibis Jedlika, 1934, Dicranoncus [Colpodes] 458

rufulus Fairmaire, 1859, Agabus 40

rufulus Bedel, 1898, Brachynopterus 419

rufulus Dejean, 1829, Bradycellus [Acupalpus] 401

rufulus Géhin, 1885, Carabus 198

rufulus Aubé, 1838, Hydaticus 53

rufulus Régimbart, 1888, Laccophilus 77

rufulus Aubé, 1838, Stictonectes [Hydroporus] 69

rufulus Habu, 1978, Synuchus 546

rufulus Rey, 1882, Tachys 275

rufulus Dejean, 1831, Trechus 340

rufum Mikhailov, 1996, Bembidion 258

rufus P. Rossi, 1790, Apotomus [Scarites] 237



rufus Breuning, 1933, Carabus 178

rufus Ragusa, 1892, Demetrias 418

rufus L. Dufour, 1820, Dromius 419

rufus Brüggemann., 1873, Harpalus 376

rufus Schilsky, 1889, Harpalus 373

rufus Clark, 1863, Herophydrus [Hyphydrus] 71

rufus Matits, 1911, Molops 474

rufus Chaudoir, 1878, Platidiolus 285

rufus Duftschmid, 1812, Platyderus [Carabus] 523

rufus A. Zimmermann, 1922, Rhantus 46



rugatinus Géhin, 1880, Carabus 171

rugatus Breuning, 1934, Carabus [Sphodristocarabus] 194

rugibasis Straneo, 1982, Pterostichus 510

rugiceps Chaudoir, 1846, Bembidion 20, 269

rugiceps Kraatz, 1884, Calosoma [Callisthenes] 119

rugiceps Klug, 1832, Clivina 220

rugiceps Straneo, 1984, Pterostichus [Amolopsa] 494

rugicolle Chaudoir, 1846, Agonum 451



rugicolle Motschulsky, 1844, Bembidion [Peryphus] 244

rugicolle Saulcy & Reiche, 1855, Bembidion [HN!] (as Reiche & Saulcy) 252

rugicollis J. Schmidt, 1998, Broscus 236

rugicollis Putzeys, 1873, Calathus 527



rugicollis LaFerté‑Sénectère, 1851, Chlaenius [Epomis] 353

rugicollis K. Daniel & J. Daniel, 1898, Cychrus 205



rugicollis Chaudoir, 1861, Desera 358

rugicollis Fairmaire & Laboulbène, 1854, Dyschirius 228

rugicollis Motschulsky, 1860, Harpalus [Platus] 385

rugicollis R. F. Sahlberg, 1844, Pterostichus [Feronia] 491

rugicollis Stephens, 1828, Pterostichus [Omaseus] 500

rugifer Kraatz, 1877, Carabus [Procrustes] 187



rugifrons Reitter, 1889, Carenochyrus [Ditomus] 363

rugimarginata Marsham, 1802, Nebria 92

rugipennis Motschulsky, 1861, Carabus [Damaster] 148

rugipennis Faldermann, 1836, Pseudotaphoxenus [Sphodrus] 542

rugipennis Chaudoir, 1855, Scarites 234

rugipleuris Reitter, 1919, Brachinus 212

rugosicollis Joseph, 1870, Laemostenus [Sphodrus] 535

rugosicollis Brullé, 1832, Myas 475

rugosifrons Nietner, 1856, Pseudoclivina [Clivina] 221

rugosior W. Horn, 1896, Neocollyris [Collyris] 116



rugosipennis Guignot, 1932, Agabus 40

rugosipennis Géhin, 1882, Carabus 139

rugosipennis Jedlika, 1932, Pterostichus 497

rugosissimus Breuning, 1943, Carabus 182

rugosofemoralis Balkenohl, 1999, Clivina 220



rugosothoracis Mandl, 1955, Carabus 137

rugosus Dejean, 1826, Carabus 187

rugosus Fabricius, 1792, Carabus 161

rugosus Lucas, 1846, Carabus 152

rugosus W. S. MacLeay, 1825, Cybister [Dytiscus] 50

rugosus Landin, 1955, Haplogaster 232

rugosus Poppius, 1905, Hydaticus 54

rugosus Nietner, 1857, Platymetopus [Harpalus] 393

rugosus Gebler, 1823, Pterostichus [Poecilus] 499

rugosus Ménétriés, 1832, Zabrus [Pelor] 572



rugulosa Gistel, 1857, Amara [Celia] 556

rugulosior Deuve, 1996, Carabus 190

rugulosipennis Deuve & Kaláb, 1992, Carabus 175

rugulosum Motschulsky, 1850, Calosoma [Callisoma] 120

rugulosus Birthler, 1885, Carabus 173

rugulosus L. Redtenbacher, 1844, Cybister [Trochalus] 50



rugulosus Heer, 1837, Harpalus 378

rugulosus Heer, 1837, Pterostichus 508

rugulosus Kraatz, 1884, Zabrus 571

ruizi Jeanne, 1996, Platyderus 523

ruiztapiadori Zaballos, 1996, Geocharis 239

rumelica Apfelbeck, 1904, Cicindela 106



rumelicum Apfelbeck, 1902, Bembidion 254

rumelicus Chaudoir, 1867, Carabus 198

rumelicus Apfelbeck, 1904, Harpalus 381

rumelicus J. Müller, 1934, Speluncarius [Pterostichus] 516



rungsi Antoine, 1942, Zuphium 574

rupa Jedlika, 1935, Amara 558

rupestre Dawson, 1854, Bembidion 254

rupestris Rotrou, 1946, Laemostenus 539

rupestroides Netolitzky, 1943, Bembidion 261

rupicola C. Zimmermann, 1832, Amara 557

rupicola Sturm, 1818, Ophonus [Harpalus] 391

rupicoloides Sharp, 1912, Ophonus 390

ruri Ashida, 1998, Trechiama 323



russicum Fischer von Waldheim, 1828, Calosoma 121

russicus Jedlika, 1957, Taphoxenus 544

russipes Mulsant & Wachanru, 1852, Cymindis 411

rusticum Casey, 1918, Bembidion 266



rusticus Geoffroy, 1785, Ophonus [Buprestis] 390

rusticus Andrewes, 1920, Pseudognathaphanus [Gnathaphanus] 363

rutenus Jeannel, 1942, Pterostichus [Haptoderus] 489

ruteri Colas, 1935, Trechus 332

ruteri Morvan, 1972, Xestagonum [Colpodes] 468

ruthena Tschitschérine, 1894, Amara 562

ruthenum Tschitschérine, 1895, Bembidion 250

ruthenus Reitter, 1878, Duvalius [Trechus] 307


ruthenus Motschulsky, 1849, Dyschirius 224

ruthenus Motschulsky, 1866, Pterostichus [Logarus] 486

ruthi Jeannel, 1929, Trechus 340

rutilans Wollaston, 1862, Broscus 236

rutilans Dejean, 1826, Carabus 141

rutilans Bates, 1889, Loxoncus [Anoplogenius] 404

rutilans Dejean, 1828, Pterostichus [Feronia] 513

rutilicollis Reitter, 1915, Lebia 428

rutilipes Csiki, 1930, Pterostichus 504

rutilus Schaum, 1863, Aephnidius [Masoreus] 357

rutilus Schaum, 1860, Perileptus 289

rutishauseri G. Hauser, 1912, Carabus [Coptolabrus] 141

rybinskii Reitter, 1896, Carabus 200

rybinskii Knirsch, 1924, Duvaliopsis [Trechus] 298

rybinskii Kinel, 1936, Nebrioporus [Deronectes] 65

ryosukei Uéno, 1960, Yamautidius [Rakantrechus] 345

ryugashiensis Uéno, 1988, Kurasawatrechus 313

ryukyuensis Habu, 1964, Amphimenes 435

ryukyuensis Habu, 1975, Clivina 220

ryukyuensis Satô, 1961, Copelatus 47

ryukyuensis Habu, 1978, Dicaelindus 443

ryukyuensis Satô, 1971, Gyrinus 28

ryukyuensis Uéno, 1974, Perileptus 289



ryukyuensis Habu, 1969, Trichotichnus 397

saadii Morvan, 1976, Trechus 329

sabaudus Géhin, 1880, Carabus 171

sabaudus Fauvel, 1882, Cychrus 203

sabaudus Fauvel, 1865, Hydroporus 63

sabdensis Deuve, 1994, Cychrus 205



sabrosensis Reitter, 1896, Carabus 167

sabuleti Fischer von Waldheim, 1828, Scarites 234

sabulicola Waltl, 1835, Cylindera [Cicindela] 110

sabulicola Lacordaire, 1835, Dyschirius [Clivina] 227

sabulicola Panzer, 1796, Ophonus [Carabus] 392

sabulosa Audinet‑Serville, 1821, Amara [Harpalus] 557

sabulosa Motschulsky, 1850, Cymindis [Tarus] 415



sabulosa Fabricius, 1787, Nebria [Carabus] 94

sabulosus Fabricius, 1792, Acinopus [Scarites] 368



sabulosus Olivier, 1795, Scarites 232

sacarum Kabak, 1998, Carabus 146



sacha Zaitzev, 1910, Hydroporus 63

sachalinensis Lapouge, 1906, Carabus 136

sachalinensis Matsumura, 1911, Carabus 136

sachalinensis A. Morawitz, 1862, Cicindela 107

sachalinensis Kamiya, 1936, Gyrinus 28


sachalinensis Matsumura, 1911, Harpalus 378

sachalinensis Kamiya, 1938, Ilybius [Agabus] 43

sachalinensis Kirschenhofer, 1985, Pterostichus 506

sachalinensis Lafer, 1989, Trechus 340



sachalinensis Habu, 1954, Trichotichnus 395

sachalinica Hieke, 1999, Amara 567

sachalinica Nakane, 1952, Clivina 219

sachalinicus Breuning, 1932, Carabus 133

sacheri C. G. Thomson, 1875, Carabus 151

sacheri I. Frivaldszky von Frivald, 1865, Pterostichus 513

sachiaensis Lassalle, 1999, Acalathus 529

sachianus Mandl, 1981, Carabus 166

sachtlebeni Jedlika, 1962, Pterostichus 509

sackeni Semenov, 1898, Carabus 189

sackenioides Deuve, 1991, Carabus 189

sacra Putzeys, 1875, Clivina 220

sadoense Habu, 1975, Agonum [Platynus] 453

sadoensis Schauberger, 1932, Anisodactylus 361

sadoensis Schauberger, 1929, Harpalus 386

sadona Bates, 1883, Nebria 94

saetosus Knirsch, 1913, Chaetoduvalius [Anophthalmus] 297

saeviens Bates, 1883, Nebria 94

saezi Vuillefroy, 1868, Platyderus 523

safei Abdul‑Karim & Ali, 1986, Agabus 40

safonovae Budarin, 1995, Pterostichus 492

saga Cavazzuti, 1997, Carabus 126



saganicola J. Müller, 1942, Apotomus 237

sagax Motschulsky, 1866, Pterostichus [Badistrinus] 487

sagax Jeannel, 1960, Trechus 340

saghaliensis Takizawa, 1933, Hydroporus 64

saginata Ménétriés, 1849, Amara [Celia] 557

saginatus Schaum, 1857, Hygrotus [Hydroporus] 72

sagitta Coye, 1870, Demetrias 419

sagitta Reitter, 1896, Syntomus [Metabletus] 432

sagittalis Jeannel, 1937, Pterostichus [Haptoderus] 492

saglensis Shilenkov, 1998, Trechus 340

sagowskii Lutshnik, 1909, Harpalus [Ophonus] 385

sagrensis Mateu, 1952, Trymosternus 410



saharae Bedel, 1900, Paussus 211

saharensis Mateu, 1984, Masoreus 357

sahendensis Morvan, 1981, Taphoxenus [Pseudotaphoxenus] 544

sahlbergi Sharp, 1882, Agabus 38

sahlbergi Poppius, 1906, Amara [HN] 559

sahlbergi Zetterstedt, 1838, Amara [Harpalus] 556

sahlbergi Dejean, 1826, Carabus 170


sahlbergi A. Zimmermann, 1932, Deronectes 59

sahlbergi J. Müller, 1924, Dyschirius 226

sahlbergi Falkenström, 1939, Haliplus 31

sahlbergi Zaitzev, 1905, Hydaticus 54

sahlbergi Nilsson, 2001, Hydroporus [Hydrocolus] 64



sahlbergi Sharp, 1882, Hygrotus [Coelambus] 72

sahlbergi Holdhaus, 1912, Microlestes 423



sahlbergi Tschitschérine, 1894, Pterostichus [Feronia] 490

sahlbergianus Géhin, 1885, Carabus 134

sahlbergianus Lutshnik, 1922, Ophonus [Harpalus] 390

sahlbergii Chaudoir, 1850, Agonum [Anchomenus] 449

sahlbergii Dejean, 1831, Bembidion 271

sahlbergii Fischer von Waldheim, 1824, Cicindela 107



sahlbergii Seidlitz, 1887, Graphoderus 49

sahlbergioides Munster, 1932, Bembidion 272

saidaensis Deuve, 1992, Carabus 158

saigonica Kult, 1951, Clivina 220

sainteclarei Schatmayr, 1929, Pterostichus 504

sainvali Branes, 1987, Carabus 140

saishutoicus Csiki, 1927, Carabus 156

saitoi Morita, 1997, Bradycellus 402

sajanensis Reitter, 1896, Carabus 195

sajanensis P. Moravec, 1993, Trechus 340

sajanica Bänninger, 1932, Nebria 89

sajanicus Csiki, 1901, Carabus 170

sajanum Shilenkov, 1995, Bembidion 244

sajanus Breuning, 1927, Carabus 164

sakagutii Nakane, 1957, Nebria 93

sakagutii Nakane & Ishida, 1958, Pterostichus 507

sakatae Jedlika, 1956, Trichotichnus 396

sakataensis Habu, 1954, Parabroscus [Synuchus] 544

sakishimanus Nakane, 1990, Hydaticus 53

sakuraii Nakane, 1978, Omoglymmius 78

salassus Jeannel, 1927, Trechus 340



salax Motschulsky, 1866, Carabus 198

saldaitisi Obydov, 1997, Carabus 170

saldoensis Bruneau de Miré, 1955, Reicheia 222

salebrosus Zamotajlov & Sciaky, 1999, Chinapenetretus 281

salechardensis Obydov, 1999, Carabus 170

salibai Antoine, 1953, Antoinella 293



salicola Motschulsky, 1864, Brachinus 214

salignicola Schweiger, 1969, Carabus 186

salignus Schweiger, 1969, Carabus 158

salina Fairmaire & Laboulbène, 1854, Nebria 93

salinarius Wallis, 1933, Haliplus 32


salinator Pantel, 1888, Harpalus 379

salinulus Reitter, 1900, Harpalus 381



salinus Baudi di Selve, 1864, Acupalpus [Stenolophus] 398

salinus Schaum, 1843, Dyschirius 227

salinus Dejean, 1829, Harpalus 381

salinus Joly, 1840, Nebrioporus [Hydroporus] 67

salinus Motschulsky, 1844, Pogonus 286

salinus Dejean, 1825, Scarites 234

salisburgensis Kraatz, 1879, Carabus [Megadontus] 166

salmantinus Bolívar y Pieltain, 1922, Carabus 140

salmantinus Jeanne, 1996, Platyderus 523

salpansis Deuve, 1984, Carabus 162

salpensis Deuve, 1985, Deltocolpodes [Agonum] 457

salsipotens Wollaston, 1864, Pogonus 287

saltatoria Gistel, 1837, Cicindela 104

salyensis Jeannel, 1942, Molops 474

salzmanni Germar, 1824, Abacetus [Harpalus] 346

samai Cavazzuti & Rapuzzi, 1998, Carabus 182

samai Schizzerotto, 1988, Stictonectes 69

samai Straneo, 1986, Tapinopterus 520



sambucensis Schaefer, 1973, Carabus 195

sambylensis Shilenkov, 1996, Carabus 171

sambylensis Belousov & Kabak, 1994, Trechus 340

samharicus Putzeys, 1873, Dyschirius 225

samnis Jeannel, 1921, Trechus 332

samnitica A. Fiori, 1899, Amara 565

samniticus A. Fiori, 1896, Pterostichus 503

samojedorum J. R. Sahlberg, 1880, Haliplus 31

samojedorum J. R. Sahlberg, 1880, Poecilus [Feronia] 481

samokovi Fery & Nilsson, 1993, Ilybius [Agabus] 43



samosir Csiki, 1938, Laccophilus 77

sampoi Casale, 1980, Doderotrechus 298

samson Reiche & Saulcy, 1855, Odotoncarus [Ditomus] 366

samurai Lutshnik, 1916, Poecilus [Platysma] 483



sanadai Habu, 1960, Celaenephes [Fukuchina] 410

sanatum Bates, 1883, Bembidion 271

sanatus Jedlika, 1934, Peliocypas [Risophilus] 434

sanchari Andrewes, 1921, Carabus 176

sanctaecrucis Wollaston, 1864, Nesacinopus [Cratognathus] 388

sanctaeluciae J. Müller, 1931, Anophthalmus 292

sanctus Motschulsky, 1864, Brachinus 216

sanctus Sharp, 1882, Hyphydrus 75

sandaligera Bates, 1873, Lebia 430

sandankyonis Habu, 1974, Hikosanoagonum [Platynus] 458

sanfilippoi Fery & Brancucci, 1997, Deronectes 58


sanfilippoi Ghidini, 1958, Hydroporus 64

sanfilippoi Fery, 1992, Hygrotus [Coelambus] 72

sanggarparensis Deuve, 1991, Carabus 191

sanggarparicus Deuve, 1996, Carabus 191



sangilenus Obydov, 1997, Carabus 164

sangjaei Kwon & Lee, 1984, Carabus [Damaster] 144

sanguinea Dejean, 1825, Clivina 219

sanguinipes Kryzhanovskij & Mikhailov, 1975, Laemostenus [Pristonychus] 538

sankhuana Hieke, 1990, Amara 563

sanmarkii C. R. Sahlberg, 1826, Oreodytes [Hyphydrus] 68

sansii Aubé, 1838, Nebrioporus [Hydroporus] 66

santschii Normand, 1916, Typhlocharis 241



saouensis David & Marchal, 1968, Carabus 166

sapancaensis Blumenthal & Breuning, 1967, Carabus 154



sapaudiae Jeannel, 1937, Nebria 93

saphyrea Dejean, 1828, Amara 551



saphyreum Gautier des Cottes, 1869, Bembidion 258

saphyrina Gené, 1836, Cicindela 104



saphyrina Barthe, 1922, Cylindera [Cicindela] 109

saphyrina Chaudoir, 1850, Neocollyris [Collyris] 117


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