Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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sculptus Schaum, 1860, Trechus 326

scurra Tschitschérine, 1901, Pterostichus [Platysma] 502



scurroides Jedlika, 1958, Pterostichus 502

scutellaris Chaudoir, 1842, Brachinus 214

scutellaris Sharp, 1882, Nebrioporus [Deronectes] 66

scutellaris Stephens, 1828, Tachys 276



scuticeps Bates, 1892, Clivina 220

scuticollis Fairmaire, 1888, Pterostichus [Steropus] 514



scydmaenoida Nietner, 1858, Tachyura [Bembidium] 277

scytha Tschitschérine, 1899, Harpalus 373

scythicus Schaum, 1856, Carabus 197

scythropus Andrewes, 1923, Pheropsophus 218

scythus Motschulsky, 1847, Carabus 182



scythus Schaum, 1864, Nebrioporus [Hydroporus] 67

scyticum K. Daniel, 1902, Bembidion 262

sebaouensis Bruneau de Miré, 1955, Reicheia 222

sebenelloi Daffner, 1983, Orotrechus 317

secalis Paykull, 1790, Trechus [Carabus] 325

sechellensis Basilewsky, 1973, Lophocoryza 221



seclusus Casey, 1914, Harpalus 379

secretus Fall, 1926, Dyschirius 226

secticollis Bates, 1873, Apristus 431

secundus Wrase, 1998, Bradycellus 401

securiger Uéno, 1982, Yamautidius 345



sedakovi Fischer vonWaldheim, 1844, Carabus 183

sedakowi Poppius, 1908, Pterostichus [Cryobius] 493

sedilloti Régimbart, 1895, Bidessus 55

sedilloti Régimbart, 1878, Graptodytes [Hydroporus] 60



sediquii Ledoux, 1988, Calosoma [Callisthenes] 119

seductum Casey, 1918, Bembidion 271

sedula Lutshnik, 1930, Amara 565

sedulus Knirsch, 1926, Anophthalmus [Haplotrechus] 291



sedulus Dejean, 1828, Pterostichus [Argutor] 486

segnis Dejean, 1829, Harpalus 376

segnis Schaum, 1864, Zabrus 571

segonzaci Bedel, 1903, Platydela 114

segregatus A. Morawitz, 1886, Carabus 146



seguranus Lauffer, 1905, Carabus [Chrysocarabus] 161

seguranus J. Vives & E. Vives, 1982, Laemostenus [Ceuthosthenes] 535



seidlitzi Kraatz, 1890, Cephalota [Cicindela] 102

seidlitzi Gerhardt, 1899, Hydroporus 61

seidlitzi Jakobson, 1908, Orectochilus 29

seidlitzi Schaum, 1864, Zabrus 569



seidlitzii Ragusa, 1887, Cybister 50

seijasi Español, 1969, Geotrechus 310



seileri Heer, 1837, Carabus 135

seimczanensis Obydov, 1993, Carabus 134

seishinensis Lapouge, 1932, Carabus [Leptocarabus] 159



seishini Jedlika, 1957, Amara 561

seistanus Andrewes, 1929, Dyschirius 227

seizaburoi Imura, Dejima & Mizusawa, 1993, Carabus 177



sejunctus Casey, 1914, Harpalus 377

sejunctus Bates, 1883, Pterostichus 494

sejunctus Leleup, 1945, Trechus 337


sekerai Reitter, 1906, Anillus 238

seladon Holdhaus, 1912, Microlestes 424



seladon Schauberger, 1926, Ophonus [Harpalus] 391

selene Schmidt‑Göbel, 1846, Scarites 234

selengensis Mandl, 1936, Cicindela 104

sellae Kraatz, 1878, Carabus 180

sellae Stierlin, 1881, Carabus 140

sellae Stierlin, 1881, Pterostichus 512



sellata Schmidt‑Göbel, 1846, Lebia 429

sellata Heller, 1921, Parena [Phloeodromus] 434



sellatoides Jedlika, 1940, Parena [Crossoglossa] 434

sellatus Peyerimhoff, 1931, Hydroglyphus [Bidessus] 56

sellatus Motschulsky, 1855, Mesolestes [Dromius] 421

sellatus Gebler, 1843, Mnuphorus [Cymindis] 358

selmanni Duftschmid, 1812, Pterostichus [Carabus] 488



semashchoensis Imura & Zamotajlov, 1993, Carabus 133

semecensis Winkler, 1926, Duvalius 300



semedanensis Jeannel, 1928, Duvalius 303

semelai Deuve, 1995, Carabus 126

semelai Deuve, 1997, Cychrus 205

semelederi Chaudoir, 1861, Cymbionotum [Graniger] 235

semenovi Lindroth, 1965, Bembidion 247

semenovi Belousov & Kataev, 1990, Broscosoma 235

semenovi Fedorenko, 1993, Dyschirius 228

semenovi Jeannel, 1962, Epaphiopsis [Epaphiama] 309

semenovi Perrault, 1986, Leistus 82

semenovi Glasunov, 1913, Mnuphorus 358



semenovi Tschitschérine, 1903, Notiophilus 96

semenovi Belousov & Kabak, 1992, Trechus 340



semenoviana Lutshnik, 1935, Amara 551

semenoviana Shilenkov, 1976, Nebria 89

semenovianus Breuning, 1932, Carabus 150

semenoviellus Breuning, 1932, Carabus 123

semenovii Motschulsky, 1859, Calosoma [Callisthenes] 118

semenowi Meyer‑Darcis, 1902, Carabus [Coptolabrus] 143

semenowi Dokhtouroff, 1888, Chaetodera [Cicindela] 103

semenowi Tschitschérine, 1895, Chlaenius 350

semenowi Tschitschérine, 1899, Chydaeus [Acrogeniodon] 362

semenowi A. I. Jakovlev, 1896, Colymbetes [Cymatopterus] 45

semenowi V. E. Jakovlev, 1890, Cymindis 415

semenowi Tschitschérine, 1899, Dicheirotrichus [Trichocellus] 403

semenowi Reitter, 1893, Eocarterus [Odontocarus] 366

semenowi Tschitschérine, 1901, Harpalus 382

semenowi A. I. Jakovlev, 1897, Hydroporus 64

semenowi A. I. Jakovlev, 1899, Hygrotus [Coelambus] 72



semenowi A. I. Jakovlev, 1897, Platambus [Anagabus] 44

semenowi Lutshnik, 1915, Platynus [Agonum] 463



semenowi Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 542

semenowi Tschitschérine, 1888, Pterostichus [Pseudohaptoderus] 510



semetricus Kraatz, 1878, Carabus 173

semiadelaidae Stark, 1890, Carabus 133

semiaeneum Fairmaire, 1886, Xestagonum [Colpodes] 468



semiapicalis Beuthin, 1896, Cicindela 104

semiapicalis Beuthin, 1905, Cylindera [Cicindela] 109

semiaratus Peyerimhoff, 1941, Dyschirius 229

semibraccatum Netolitzky, 1911, Bembidion 258

semicarinata Putzeys, 1877, Clivina 220

semicastaneus Reitter, 1900, Harpalus 377

semicircularis W. S. MacLeay, 1825, Scarites 232



semicirculatus O. Schneider, 1898, Rhantus 46

semicircumcincta Mandl, 1959, Myriochila [Cicindela] 114



semiclusus Walker, 1871, Nebrioporus [Hydroporus] 66

semicoeruleum J. Schmidt, 1995, Xestagonum 468

semicoriaceus Kraatz, 1881, Carabus 199

semicostatus Reineck, 1921, Dytiscus 52

semicupreonitens Schulz, 1900, Carabus 139

semicupreum Fairmaire, 1887, Agonum 451



semicupreus Fügner, 1891, Poecilus 483

semicyaneum Meier, 1899, Bembidion 247

semicyaneus Solsky, 1874, Chlaenius 356

semicylindricus Piochard de la Brûlerie, 1872, Dixus [Aristus] 365

semiferrugineum Kirschenhofer, 1984, Bembidion 244

semifuscus Cavazzuti & Ratti, 1999, Carabus 128

semigranosus Palliardi, 1825, Cychrus 205



semilineatus A. Zimmermann, 1915, Hygrotus 73

semilineatus A. Fleischer, 1898, Rhantus 47

semilotum Netolitzky, 1911, Bembidion 255

semilucidum Motschulsky, 1862, Asaphidion [Tachypus] 243

semiluitum Bates, 1883, Bembidion 243

semilunium Netolitzky, 1914, Bembidion 244

seminarius Motschulsky, 1859, Hydrovatus 70



seminiger Reitter, 1913, Brachinus 214

seminiger LeConte, 1862, Colymbetes 45

seminiger DeGeer, 1774, Hydaticus [Dytiscus] 54



seminigra Fairmaire, 1886, Merizomena 409

seminitens Kirschenhofer, 1991, Pterostichus 497

seminitidus Chaudoir, 1856, Chlaenius 354

seminskiense Shilenkov, 1990, Bembidion 269

semiobscurus Pic, 1905, Oedesis [Eriotomus] 366

semiopacus Reitter, 1895, Carabus 159



semiopacus Lassalle, 1993, Pseudotaphoxenus 542

semiplagiatus Reitter, 1887, Dromius 420



semipunctata Eschscholtz, 1833, Loricera 98

semipunctatum Donovan, 1806, Bembidion [Carabus] 254



semipunctatum Graëlls, 1858, Bembidion 252

semipunctatus Géhin, 1885, Carabus 187

semipunctatus Dejean, 1829, Harpalus 372

semipunctatus Fabricius, 1792, Noterus [Dytiscus] 34

semipunctatus Fabricius, 1775, Notiophilus [Elaphrus] 96

semipunctatus Reitter, 1904, Penetretus 285

semipunctatus Fairmaire, 1859, Zabrus 571

semipurpureus Motschulsky, 1865, Chlaenius 353

semirubra Andrewes, 1926, Coryza 221

semirubricum Reitter, 1914, Cymbionotum [Graniger] 235

semirufescens Semenov, 1889, Acalathus 529



semirufipes Semenov, 1898, Carabus 176

semirufus A. Zimmermann, 1921, Colymbetes 45

semirufus Germar, 1844, Deronectes [Hydroporus] 59



semirugosus Schaufuss, 1882, Carabus 158

semirugosus Chaudoir, 1855, Scarites 234



semirutilipes Semenov, 1898, Carabus 190

semisericeus Fairmaire, 1879, Calathus 528

semistriatus Dejean, 1828, Broscus [Cephalotes] 236

semistriatus Fischer von Waldheim, 1823, Carabus 197

semistriatus Kraatz, 1885, Carabus 147

semistriatus Dejean, 1825, Dyschirius [Clivina] 229



semistriatus Linné, 1758, Dytiscus 52

semistriatus Chaudoir, 1879, Platynus [Colpodes] 465

semisulcatus O. F. Müller, 1776, Dytiscus 52

semisuturalis Dupuis, 1912, Pentagonica 448



semiviolaceus Dejean, 1829, Harpalus 375

semiviolaceus Chaudoir, 1868, Lesticus [Triplogenius] 472

semiviridis Louwerens, 1953, Platynus [Colpodes] 465



semiviridis Pic, 1892, Pterostichus 504

semivittata Chaudoir, 1850, Cymindis 417

semivittata Fabricius, 1801, Lophyra [Cicindela] 113

senegalensis Gestro, 1901, Carabidomemnus [Arthropterus] 209

senegalensis Aubé, 1838, Cybister 50



senegalensis Dejean, 1831, Diplocheila [Rembus] 439

senegalensis Laporte, 1835, Heterhydrus [Hyphydrus] 74

senegalensis Dejean, 1831, Siagona 218

senenti Español, 1967, Subilsia 320

senilis Schaum, 1859, Molops [Pterostichus] 475

senilis Nietner, 1857, Platymetopus [Harpalus] 393

seoanei Fairmaire, 1878, Chlaenius 352


seonhyeongae Kwon & Lee, 1984, Carabus [Damaster] 123

seoulensis Deuve, 1998, Carabus 142

separandus Kraatz, 1881, Carabus [Sphodristus] 192

separandus Harald Lindberg, 1950, Dicrodontus 573

separata Fleutiaux, 1894, Cosmodela [Cicindela] 108

separatus Lapouge, 1907, Carabus 192

separatus Dejean, 1826, Licinus 442

sepositus Obydov, 1998, Carabus 183

seppenhoferi Bognolo, 1997, Anophthalmus 293

septemcarinatus Motschulsky, 1840, Carabus 164

septemlineatus Reitter, 1888, Carabus [Megadontus] 164

septemlineatus Jedlika, 1934, Platynus [Colpodes] 465



septemseriatus J. R. Sahlberg, 1875, Agabus [Gaurodytes] 40

septemtrionalis Gyllenhal, 1827, Dytiscus 52

septemtrionalis Habu, 1947, Trichotichnus [Oxycentrus] 397



septensis Lagar Mascaró, 1968, Deronectes 58

septentrionalis Curtis, 1840, Amara 549

septentrionalis Schiødte, 1837, Amara 556

septentrionalis Dupré, 1991, Aphoenops 294

septentrionalis Breuning, 1932, Carabus 172

septentrionalis Motschulsky, 1852, Dyschirius 227

septentrionalis Naviaux, 1999, Neocollyris 116

septentrionalis Gyllenhal, 1826, Oreodytes [Hyphydrus] 68

septentrionis Dejean, 1828, Patrobus [NP] 22, 285

septentrionis Chaudoir, 1868, Pterostichus [Feronia] 506

septentrionum Munster, 1923, Dyschirius 226

sequanus Lapouge, 1916, Carabus 171

sequensi Reitter, 1899, Omphreus 445

sequensi Reitter, 1899, Scarites 233

sequistriata Panzer, 1789, Cymindis [Carabus] 413

serbicum Apfelbeck, 1902, Bembidion 270



serbicus Hopffgarten, 1878, Carabus 174

serbicus Ganglbauer, 1900, Dicropterus [Scotodipnus] 239

serbicus Matits, 1910, Harpalus 386

serbicus Winkler, 1933, Omphreus 446

serbicus Csiki, 1904, Platynus 463

serbicus Apfelbeck, 1899, Pterostichus 495

serbicus Apfelbeck, 1902, Trechus 338

serdicana Apfelbeck, 1904, Amara 565



serdicanum Apfelbeck, 1904, Bembidion 254

serdicanus Apfelbeck, 1904, Harpalus 386



serenus Gredler, 1863, Patrobus 285

sergeii Zamotajlov, 1988, Deltomerus 283

sergeii Farka, 1999, Leistus 81

serianensis Breit, 1913, Boldoriella [Trechus] 296



seriatissimus Reitter, 1896, Carabus 174

seriatofoveatus Reitter, 1896, Carabus 181

seriatoporus Fischer von Waldheim, 1828, Carabus 200

seriatoporus Schaum, 1864, Zabrus 572

seriatum Motschulsky, 1844, Bembidion [Leja] 253



seriatus Motschulsky, 1850, Carabus 164

seriatus Roeschke, 1907, Cychrus 205



seriatus Chaudoir, 1846, Harpalus 379

seriatus Sharp, 1882, Nebrioporus [Deronectes] 66

seriatus Chaudoir, 1850, Pterostichus [Feronia] 506

seriatus Ganglbauer, 1915, Zabrus 571

sericans Bates, 1873, Tachys 276

sericata Lutshnik, 1935, Amara 550

sericatus Cavazzuti, 1999, Carabus 126

sericatus Tschitschérine, 1906, Harpalus 385

sericea Jedlika, 1953, Amara 551

sericea Andrewes, 1922, Dioryche 370



sericeum Fabricius, 1792, Calosoma [Carabus] 120

sericeus C. Zimmermann, 1832, Aephnidius [Masoreus] 357

sericeus Csiki, 1946, Carabus 179

sericeus Duftschmid, 1812, Harpalus [Carabus] 373

sericeus Geoffroy, 1785, Harpalus [Buprestis] 385

sericeus Motschulsky, 1844, Harpalus 382

sericeus Eschscholtz, 1818, Hydroporus [Dytiscus] 61

sericeus Fischer von Waldheim, 1824, Laemostenus [Sphodrus] 539



sericeus A. Costa, 1847, Nebrioporus [Hydroporus] 66

sericeus Klug, 1834, Orectogyrus 29

sericeus Fischer von Waldheim, 1824, Poecilus 483

sericeus J. T. Fleischer, 1829, Trechoblemus [Trechus] 324

sericimicans Chaudoir, 1876, Chlaenius 348



sericum Lapouge, 1930, Calosoma [Campalita] 120

sericus Lapouge, 1925, Carabus 178

seriepunctata W. Horn, 1892, Cylindera [Cicindela] 112



seriepunctata L. Redtenbacher, 1843, Cymindis 412

seriepunctatus Schauberger, 1932, Acinopus 369

seriepunctatus Peyron, 1858, Badister 440



seriepunctatus Sturm, 1818, Harpalus 373

seriepunctatus Chaudoir, 1846, Pterostichus [Feronia] [HN] 495

seriepunctatus Mannerheim, 1843, Pterostichus 487

sermeti Mateu & Colas, 1954, Mesolestes [Microlestes] 421

sermeti Jeannel, 1948, Trichaphaenops 344

serorum Netolitzky, 1934, Bembidion 244



serotinum Hochhuth, 1871, Bembidion 247

serpentina Germar, 1844, Calomera [Cicindela] 100

serrae E. Vives, 1976, Apoduvalius 295



serratensis Coiffait, 1958, Microtyphlus 240

serrator Forsskål, 1775, Graphipterus [Carabus] 359

serratulus Balkenohl, 1999, Trilophus 222

serratus Balkenohl, 1999, Trilophus 222

serricornis Paykull, 1799, Agabus [Dytiscus] 37

serripes Latreille, 1804, Harpalus 375

serripes Quensel, 1806, Harpalus [Carabus] [HN] 382



serrorum Csiki, 1944, Carabus 167

serrulatus Winkler, 1926, Cimmerites [Nannotrechus] 297

serrulatus Jeannel, 1946, Geotrechus 310

serrulatus Jeannel, 1946, Geotrechus 310

sertarensis Deuve, 1995, Carabus 135

sertarensis Deuve, 1995, Cychrus 204

servillei Solier, 1835, Carabus 161

servillei Solier, 1835, Cymindis 413

servus Duftschmid, 1812, Harpalus [Carabus] 382



servus Stephens, 1828, Harpalus 378

serxuensis Deuve, 1995, Carabus 135

seserligensis Shilenkov, 1998, Trechus 340

sesostris Chaudoir, 1876, Siagona 218

sesquisulcatus Wasmann, 1899, Paussus 210



sesquivittatus Fairmaire, 1880, Hydaticus 54

sessitanus Monguzzi, 1985, Trechus 340

seticeps Straneo, 1984, Pterostichus [Amolopsa] 494

seticeps J. Müller, 1934, Speluncarius [Pterostichus] 517



seticollis Ganglbauer, 1900, Pterostichus 512

seticornis Lindroth, 1980, Cillenus [Bembidion] 272



seticornis O. F. Müller, 1776, Loricera [Carabus] 98

setifeensis Lucas, 1842, Cymindis 414



setifensis Marseul, 1871, Mastax 217

setifer Kataev & N. Ito, 1999, Trichotichnus 394

setiferus Shilenkov, 1984, Archastes 79

setiger Uéno & Namkung, 1968, Kurasawatrechus 313



setiger Sloane, 1903, Lymnastis 274

setipennis Apfelbeck, 1899, Speluncarius 517



setipes Motschulsky, 1848, Microderes [Bioderus] 387

setipes Tschitschérine, 1898, Pterostichus [Feronia] 508



setiporus Reitter, 1894, Dicheirotrichus [Trichocellus] 402

setiporus Reitter, 1894, Harpalus [Loxophonus] 382

setiporus Reitter, 1893, Platynus 464

setniki Reitter, 1904, Neotrechus [Anophthalmus] 315

seto Ishikawa & Kubota, 1994, Carabus 178

setosa Ledoux & Roux, 1995, Nebria 95

setosa Holdhaus, 1924, Reicheia 222

setosa Walker, 1859, Selina [Pselaphanax] 407


setosomalaris W. Horn, 1913, Cosmodela [Cicindela] 108

setosulus Deuve, 1990, Cychrus 205

setosus Lapouge, 1910, Carabus 130

setosus J. R. Sahlberg, 1880, Licinus [Derostichus] 443

setosus Belousov & Dolzhanski, 1994, Taniatrechus 321

setosus Uéno, 1990, Trechiama 323



setteli Langenhan, 1913, Carabus 195

setulata Ledoux & Roux, 1995, Nebria 94

setulosus J. R. Sahlberg, 1895, Agabus [Gaurodytes] 36

seunglaki Kwon & Lee, 1984, Carabus [Leptocarabus] 159

seunglaki Park & Kwon, 1996, Pterostichus 502

seungmoi Park & Kwon, 1996, Pterostichus 495

seurati Alluaud, 1935, Omophron 208

sevanense Belousov, 1990, Bembidion 257

sevanensis Iablokoff‑Khnzorian, 1962, Dyschirius 228

sevanensis Shilenkov, 1983, Nebria 18, 85

sevastopolitanus Schauberger, 1926, Ophonus [Harpalus] 389

severi Ganglbauer, 1897, Anophthalmus [Trechus] 293

severini Régimbart, 1892, Hyphoporus 73

severini Régimbart, 1892, Orectochilus 29



severior Gozis, 1912, Agabus 40

severovi Deuve, 1992, Carabus 145

severum Chaudoir, 1850, Calosoma 120

sewertzowi Ballion, 1871, Calosoma [Callisthenes] 118

sewertzowi Semenov, 1898, Carabus 135

sewertzowi Tschitschérine, 1896, Cymindis 24, 417

sexdentatus Schiødte, 1841, Ilybius 43

sexguttatus Aubé, 1838, Rhithrodytes [Hydroporus] 68

sexmaculata Fabricius, 1787, Anthia [Carabus] 347

sexmaculata Baudrimont, 1923, Eurynebria 79

sexmaculata Patkiewicz, 1922, Lebia 428

sexmaculatus Dejean, 1825, Brachinus 216

sexmaculatus Andrewes, 1925, Tachys 276

sexpunctata Souverbie, 1855, Eurynebria [Nebria] 79

sexpunctatum Linné, 1758, Agonum [Carabus] 451



sexpunctatum Heer, 1838, Bembidion 269

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