Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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asturicus Jeanne, 1969, Pterostichus [Haptoderus] [HN!] 489

asturicus Jeannel, 1921, Trechus 337



asturiens Born, 1925, Carabus 167

asturiensis Salgado Costas, 1991, Apoduvalius 295

asturiensis Vuillefroy, 1866, Calathus 527

asturiensis Jedlika, 1957, Harpalus 376

asturiensis Bruneau de Miré, 1964, Nebria 18, 87



asturiensis Wagner, 1928, Penetretus 285

asturiensis Jedlika, 1958, Platyderus 522

asturiensis Heyden, 1880, Zabrus 569

astutus Tschitschérine, 1903, Penthophonus 393

astutus Tschitschérine, 1903, Pterostichus [Platysma] 496

asumarokovi E. Berlov & O. Berlov, 1996, Pterostichus 505

asymmetrica Tanaka, 1957, Amara [Armatoleirides] 552

asymmetricus Bates, 1883, Pterostichus 501

asymmetricus Uéno, 1990, Trechiama 322

atalanta Andrewes, 1935, Bembidion 261

atamuradovi Kryzhanovskij & Mikhailov, 1987, Tachys 276



atava Jankowski, 1932, Cicindela 105

atavus W. Horn, 1921, Calochroa [Cicindela] 99



atavus Lapouge, 1908, Carabus 130

atavus Schauberger, 1932, Chilotomus 364

atavus J. Müller, 1934, Neotrechus 315



atavus Friederichs, 1903, Notiophilus 97

atbaschensis Obydov, 1998, Carabus 150

atchibachi Zamotajlov, 1992, Carabus 132

atelesta Chaudoir, 1854, Myriochila [Cicindela] 113

atentsensis Deuve, 1990, Carabus 188



ater Villers, 1789, Abax [Carabus] 470

ater Brinck, 1955, Aulonogyrus 26



ater A. Fleischer, 1898, Carabus 196

ater Motschulsky, 1859, Dromius 420



ater W. Redtenbacher, 1842, Haliplus 32

ater DeGeer, 1774, Ilybius [Dytiscus] 41



ater Panzer, 1796, Ilybius [Dytiscus] 41

ater Falkenström, 1936, Platambus [Colymbinectes] 43



ater C. R. Sahlberg, 1827, Pterostichus [Harpalus] 499

aterrima Grandi, 1906, Calomera [Cicindela] 100

aterrima A. Fiori, 1896, Nebria (as atterima) 90

aterrimum Motschulsky, 1844, Bembidion [Omala] 270

aterrimus Stephens, 1828, Ilybius [Colymbetes] 42

aterrimus Herbst, 1784, Pterostichus [Carabus] 499



aterrimus Küster, 1844, Pterostichus [HN] 488

aterrimus A. Morawitz, 1863, Scarites 232



aterrimus Heyden, 1880, Typsiharpalus [Ophonus] 397

ateuches Andrewes, 1936, Clivina 220

athalassicum De Monte, 1953, Bembidion 263



athesinus Depoli, 1939, Abax 470

athesinus Depoli, 1927, Carabus 198

athleta Semenov, 1897, Carabus 157

athleta Kryzhanovskij, 1976, Chlaenius 349

athonis Schatzmayr, 1909, Trechus 328

atienzai Zaballos & Ruiz‑Tapiador, 1997, Typhlocharis 241

ativsengueni Schweiger, 1962, Carabus 153

atlante Antoine, 1957, Agonum 449

atlantica Barthe, 1922, Cylindera 111

atlantica Oberthür, 1883, Nebria 85



atlantica Alluaud, 1932, Tachyura [Tachys] 279

atlanticns Mateu, 1974, Microlestes 423

atlanticum Wollaston, 1854, Bembidion 254



atlanticus Lapouge, 1925, Carabus 154

atlanticus Régimbart, 1892, Gyrinus 27

atlanticus Escalera, 1913, Laemostenus 539



atlanticus Schauberger, 1927, Nesarpalus 388

atlanticus Fairmaire, 1875, Orthomus [Feronia] 477

atlanticus Mateu, 1957, Philorhizus 425



atlantidis Pawowski, 1972, Trechus 330

atlantis Antoine, 1925, Amara 555

atlantis Antoine, 1944, Bembidion [Peryphus] 260

atlantis Alluaud, 1923, Carabus 152

atlantis Mandl, 1944, Cicindela 103

atlantis Théry, 1933, Graptodytes [Hydroporus] 59

atlasica Escalera, 1914, Cymindis 413

atlasicus Coiffait, 1969, Anillopsidius 237

atlasicus P. Moravec & Lompe, nom. nov., Trechus 22, 326

atomus P. Moravec & Wrase, 1998, Trechus 326

atorus Uéno, 1965, Ryugadous 319

atra Stephens, 1828, Amara 547

atramentarius Rosenhauer, 1856, Styracoderus [Pterostichus] 518

atrata Habu, 1975, Achaetocephala 449

atrata Pallas, 1776, Cephalota [Cicindela] 101

atrata Dejean, 1826, Oreonebria [Nebria] 95

atrata Dejean, 1825, Siagona 218



atratula Beuthin, 1898, Cicindela 107

atratulus Mandl, 1983, Chlaenius 349



atratum Duftschmid, 1812, Agonum [Carabus] 450

atratum Stephens, 1828, Agonum 453

atratum Hornung, 1844, Bembidion 249

atratus Dejean, 1829, Acupalpus 397

atratus Mannerheim, 1853, Agabus 41

atratus Dejean, 1825, Aptinus 212

atratus G. Hauser, 1921, Carabus [Coptolabrus] [HN] 141

atratus G. Hauser, 1926, Carabus [Coptolabrus] [HN] 143

atratus Heer, 1838, Carabus 138

atratus Habu, 1951, Diplous 283



atratus Ballion, 1878, Harpalus 373

atratus Latreille, 1804, Harpalus 373

atratus Gistel, 1857, Pterostichus [ND] 515

atratus Dejean, 1825, Syntomus [Dromius] 433

atratus A. Costa, 1883, Tachys 276

atrema Andrewes, 1926, Pristosia [Calathus] 530

atricapillus Linné, 1758, Demetrias [Carabus] 419

atriceps Sharp, 1882, Copelatus 47



atriceps Crotch, 1870, Hydroporus 62

atriceps Pic, 1922, Lebia 428

atriceps Fairmaire, 1869, Perigona [Nestra] 449

atriceps W. J. MacLeay, 1871, Tachys [Trechus] 275

atricolor Régimbart, 1880, Hydrovatus 70

atricolor Bates, 1883, Synuchus [Eucalathus] 545

atricomes Bates, 1873, Agonum [Colpodes] 454



atricornis Stephens, 1835, Anisodactylus [Harpalus] 361

atricornis Fairmaire & Laboulbène, 1854, Brachinus 216

atripenne De Monte, 1947, Bembidion 270

atripennis Motschulsky, 1858, Apotomus 237

atripennis Chaudoir, 1876, Brachinus 215

atripennis Motschulsky, 1864, Brachinus 213

atripensis Ballion, 1871, Brachinus 215



atripes Ganglbauer, 1900, Anisodactylus 361

atripes Motschulsky, 1844, Bembidion [Omala] 270



atripes Géhin, 1885, Carabus 195

atrocaeruleum Stephens, 1828, Bembidion [Peryphus] 245



atrocaeruleus Stephens, 1828, Harpalus 381

atrocoerulea Sturm, 1825, Amara 548

atrocoeruleus Waltl, 1838, Pachycarus [Ditomus] 367



atrocyanea Barthe, 1919, Lophyra [Cicindela] 112

atrocyaneum Netolitzky, 1943, Bembidion 254

atrocyaneus Fairmaire, 1859, Laemostenus [Sphodrus] 536

atrocyaneus Chaudoir, 1842, Ophonus 388

atrocyaneus Dieck, 1870, Pogonus 286

atroelytrata Mandl, 1981, Cylindera [Cicindela] 110



atropos Mulsant & Godart, 1860, Hydroporus 61

atropos K. Daniel, 1903, Nebria 91

atroptera Mandl, 1963, Cylindera [Cicindela] 111

atropunctatus Pic, 1919, Brachinus 212

atrorufus Strøm, 1768, Patrobus [Carabus] 285

atrosulcatus Régimbart, 1906, Copelatus 47

atrovirescens Putzeys, 1872, Amara [Leiocnemis] 555

atrox Andrewes, 1935, Bembidion 245

atrox Andrewes, 1937, Pterostichus [Feronia] 509

attemsi Apfelbeck, 1908, Nebria 85

attemsi Apfelbeck, 1904, Tapinopterus [Pterostichus] 519

attenuata Baliani, 1932, Amara 555

attenuata Herbst, 1806, Clivina [Scarites] 219

attenuata V. E. Jakovlev, 1887, Cymindis 414

attenuata Motschulsky, 1844, Nebria 88

attenuata Chaudoir, 1865, Neocollyris [Collyris] 117

attenuata L. Redtenbacher, 1844, Neocollyris [Colliuris] 117

attenuatus Fabricius, 1792, Cychrus [Carabus] 202

attenuatus Putzeys, 1866, Dyschirius 227

attenuatus Stephens, 1828, Harpalus 373



attenuatus Chevrolat, 1840, Pterostichus [Omaseus] 499

attenuatus Jeannel, 1949, Trichaphaenops 344

atticus Gautier des Cottes, 1867, Calathus 528


atticus Reitter, 1885, Leistus 83

atticus Apfelbeck, 1904, Platyderus 522

atticus Apfelbeck, 1904, Tapinopterus [Pterostichus] 519

atupus Andrewes, 1923, Chlaenius 349

aubei Perris, 1869, Agabus 38

aubei Tournier, 1868, Cymindis 411

aubei Mulsant, 1843, Deronectes [Hydroporus] 58

aubei Ganglbauer, 1891, Hyphydrus 74

aubei Saulcy, 1863, Hypotyphlus [Scotodipnus] 239

aubei Pandellé, 1867, Trechus 326

auberti Grenier, 1864, Duvalius [Anophthalmus] 301

aubryi Schuler, 1964, Amara 548

aubryi Colas, 1969, Carabus 139

aubryi Jeanne, 1974, Orthomus 476

aubryi Coiffait, 1952, Pterostichus 498

aubryi Coiffait, 1953, Trechus 326



aubryi Jeanne, 1970, Zabrus 569

auctus Wollaston, 1862, Calathus 526



auctus Semenov & Znojko, 1932, Carabus 133

audax Gistel, 1857, Amara 550

audax Tschitschérine, 1895, Pterostichus [Feronia] 501

audouini L. Dufour, 1851, Abax [Feronia] 470

audouini Brullé, 1835, Carabus 185



audouinii Barthélemy, 1835, Neolaphyra [Cicindela] 114

aufreri Colas, 1964, Carabus 195

augustae Winkler, 1926, Neotrechus 315

augusti Jeannel, 1931, Trechus 337

augustini Ledoux & Roux, 1999, Nebria [add] 89

augustus Bates, 1888, Carabus 141

aulacocnemus Semenov, 1897, Carabus 147

aulaensis Aubry, 1981, Trechus 332

aulica Panzer, 1796, Amara [Carabus] 559

aulica Dejean, 1831, Calomera [Cicindela] 99

aulicoides J. R. Sahlberg, 1913, Calomera [Cicindela] 100



aulicus Omer‑Cooper, 1931, Rhantus 46

auliensis Lapouge, 1918, Carabus 192

aumontii Lucas, 1850, Carabus 161

auraniensis J. Müller, 1903, Carabus 160

aurasius Jeannel, 1907, Graptodytes [Hydroporus] 59

aurata Fischer von Waldheim, 1828, Blethisa 206

aurata Bates, 1873, Taicona 409

auraticolle Westhoff, 1881, Agonum 451

auratoides Reitter, 1896, Carabus 194

auratus Linné, 1761, Carabus 194

auratus Heer, 1837, Pterostichus 511


aurelium Bates, 1883, Agonum [Colpodes] 452

aureocupreus Reitter, 1880, Carabus 151

aureofusca Bates, 1889, Neocollyris [Collyris] 116

aureofuscum Bates, 1883, Bembidion 244

aureolus Notman, 1920, Dyschirius 226

aureolus Faldermann, 1836, Zabrus 569

aurescens Bates, 1883, Myas [Trigonognatha] 475



aureus DeGeer, 1774, Carabus 153

aureus Sirguey, 1931, Carabus 138

aureus P. W. J. Müller, 1821, Elaphrus 206



auriceps Chaudoir, 1842, Chlaenius 355

aurichalcea Germar, 1824, Amara 553

aurichalceus Kraatz, 1879, Carabus [Megadontus] 165

aurichalceus M. F. Adams, 1817, Zabrus [Blaps] 568



auricollis Born, 1922, Carabus 156

auricollis C. O. Waterhouse, 1867, Carabus [Damaster] 148

auricollis Gené, 1839, Chlaenius 351

auricollis Tschitschérine, 1900, Lesticus 471

auriculatus Putzeys, 1872, Carabus 155

auriculatus Wollaston, 1867, Dyschirius 224

auripilis Bates, 1883, Peronomerus 448

aurocinctus Motschulsky, 1844, Carabus 149

aurolimbatus Dejean, 1829, Carabus 165

auromarginata Kraatz, 1887, Cicindela 104

auronitens Fabricius, 1792, Carabus 138

auropunctatum Herbst, 1784, Calosoma [Carabus] 120

aurosericeus Kraatz, 1880, Carabus 172

aurosplendens Born, 1910, Carabus 196

aurouxi Español, 1966, Speleotyphlus [Microtyphlus] 240

aurulenta Fabricius, 1801, Cosmodela [Cicindela] 108

auseriacum Fabricius, 1801, Agonum [Carabus] 452

ausonicus Bucciarelli & Perissinotto, 1959, Pterostichus [Omaseus] 499



ausonius Schaufuss, 1865, Laemostenus [Pristonychus] 535

austerus Cavazzuti, 1997, Carabus 127



austerus Reichensperger, 1935, Cerapterus 209

australasiae Hope, 1842, Catascopus 435

australianus Casale, 1988, Laemostenus 536

australis Fabricius, 1775, Dineutus [Gyrinus] 26

australis Uéno, 1999, Kusumia 314

australis Blackburn, 1888, Laemostenus [Pristonychus] 536

australis Fairmaire, 1859, Laemostenus [Pristonychus] 537

australis J. R. Sahlberg, 1875, Patrobus 285

australis Aubé, 1838, Rhantus [Colymbetes] 46

australofinis Nakajima, 1994, Carabus [Damaster] 122

austriaca Schrank, 1776, Cicindela 103


austriaca Ganglbauer, 1889, Oreonebria [Nebria] 95

austriacum Schweiger, 1975, Asaphidion 242



austriacus Dalla Torre, 1877, Anisodactylus 361

austriacus Sturm, 1815, Carabus 168

austriacus Sturm, 1834, Graphoderus [Hydaticus] 49

austriacus Schauberger, 1925, Leistus 83

austriacus Ganglbauer, 1889, Molops 473



austriacus Reitter, 1908, Patrobus 285

austriacus Dejean, 1831, Trechus 326



austriae Sokolá, 1907, Carabus 135

autumnalis Lutshnik, 1933, Amara 567

autumnalis Duftschmid, 1812, Harpalus [Carabus] 373

auvrayi Charet, 1992, Carabus 156

auvrayorum Deuve & Mourzine, 1993, Carabus 122

auvrayorum Deuve & Mourzine, 1997, Cychrus 203

auzati Le Moult, 1912, Carabus 140

avadcharensis Kurnakov, 1972, Carabus 132

avadcharicus Kirschenhofer, 1987, Pterostichus 504

avaricum Belousov & Sokolov, 1988, Bembidion 268

avaricus Abdurakhmanov & Kryzhanovskij, 1983, Pterostichus 500

avarus Schauberger, 1930, Harpalus 382

avenae J. R. Sahlberg, 1908, Cymindis 411



avenionensis Schaefer, 1973, Carabus 195

avenionensis Guignot, 1925, Siettitia 69

averenskii O. Berlov & E. Berlov, 1997, Pterostichus 497

aveyronensis Fauvel, 1882, Trechus 23, 327

avgolensis Belousov & Kabak, 1998, Trechus 327

avinovi Semenov & Znojko, 1932, Carabus 163

avinovi Semenov & Znojko, 1929, Colpostoma 441

avunculus Fairmaire, 1872, Hydroporus 62

awajewi A. I. Jakovlev, 1899, Hygrotus [Coelambus] 72

awajiensis Imura, Dejima & Mizusawa, 1993, Carabus 177

awakazusanus Ishikawa, 1981, Carabus 177

awanus Uéno, 1969, Ryugadous 319

awashimae Ishikawa & Takami, 1996, Carabus 176

axillare Motschulsky, 1844, Bembidion [Omala] 248

axillare Chaudoir, 1868, Omophron 208

axillaris Fabricius, 1794, Cymindis [Carabus] 411



axillaris Mannerheim, 1853, Dicheirotrichus [Acupalpus] 403

axillaris Küster, 1852, Masoreus 357

ayderense Korge, 1971, Bembidion 245

ayrdensis Breuning & Ruspoli, 1976, Carabus 201

azaleae Cavazzuti, 1986, Carabus 193

azarai Perez Arcas, 1865, Styracoderus [Pterostichus] 518

azari Deuve, 1998, Carabus 188


azbleotroades Morvan, 1996, Platynus [Batenus] 462

azilensis Jeannel, 1917, Aphoenops 294



azoricum Lindroth, 1960, Bembidion 253

azoricum Heer, 1860, Calosoma 121

azoricus Crotch, 1867, Bradycellus 401

azoricus Borges & Serrano, 1993, Cedrorum 471

azoricus Oromí & Borges, 1991, Thalassophilus 346

azrael Semenov & Znojko, 1932, Carabus 197

azrouanus Emden & Schauberger, 1932, Harpalus 378

azruensis Théry, 1933, Stictonectes [Hydroporus] 69

azruensis Antoine, 1925, Typsiharpalus 397

azumai Habu, 1973, Agonum [Platynus] 454

azumai Habu, 1973, Coleolissus 369

azumai Habu, 1968, Harpalus 384



azurea Krausse, 1910, Cicindela 104

azureola Mandl, 1981, Calomera [Lophyridia] 101

azurescens Dalla Torre, 1877, Bembidion 248

azurescens Dejean, 1826, Carabus 165

azurescens Gyllenhal, 1827, Harpalus 381

azureulus Mandl, 1983, Chlaenius 349



azureum Gebler, 1833, Bembidion [Peryphus] 247

azureus Dejean, 1831, Chlaenius 355

azureus Fabricius, 1775, Ophonus [Carabus] 388

babadagi Zamotajlov, 1988, Deltomerus 282

babai Habu, 1973, Agonum [Platynus] 450

babai Ishikawa, 1968, Carabus 159

babai Habu, 1973, Harpalus 384

babai Satô, 1990, Oreodytes 67

babai Satô, 1982, Platynectes 44

babai Uéno, 1994, Trechiama 322

babai Habu, 1978, Trephionus 546

babaianus Ishikawa, 1983, Carabus 148

babaulti Andrewes, 1924, Bembidion 256

babaulti Andrewes, 1924, Cymindis 412



babaulti Andrewes, 1924, Laemostenus [Pristonychus] 538

babaulti Andrewes, 1924, Nebria 91

babaulti Louwerens, 1953, Platynus [Colpodes] 464

babaulti Andrewes, 1924, Tachyura [Tachys] 278

babaulti Jeannel, 1935, Trechus 327

babicola Knirsch, 1925, Duvalius [Trechus] 306



babinjensis Apfelbeck, 1919, Carabus 163

babinjensis Jeannel, 1927, Trechus 327

baborensis Bruneau de Miré, 1955, Duvalius 308

baborica Bruneau de Miré, 1955, Reicheia 222

babukensis Zamotajlov, 1988, Carabus 133


babunensis Mlyná, 1979, Harpalus 383

babusarensis Casale, 1988, Sphodropsis 543

bacchusi Wewalka, 1981, Copelatus 47

bachmayeri Mlyná, 1979, Harpalus 384

bacillus Schaum, 1857, Dyschirius 224

baconi Chaudoir, 1871, Lebia [Nematopeza] 429

baconi Chaudoir, 1878, Platynus [Colpodes] 464

baconi Chaudoir, 1876, Siagona 218



baconis Benson, 1846, Paussus 210

bactrianum K. Daniel, 1902, Bembidion 245

bactrianum K. Daniel & J. Daniel, 1898, Parazuphium [Zuphium] 573

badakschana Mandl, 1955, Cicindela 108

badakschana Jedlika, 1956, Lebia 428

badakschanus Jedlika, 1956, Harpalus 373

badakshana Kryzhanovskij, 1974, Amara 559

badakshanicum Mikhailov, 1988, Bembidion 258

badakshanus Kryzhanovskij, 1972, Laemostenus 537

badenii Wehncke, 1876, Canthydrus [Hydrocanthus] 34

badenii Wehncke, 1872, Ilybius 43

baderi Krausse, 1910, Bidessus 55

baderlei Kirschenhofer, 1988, Apristus 431

baderlei Kirschenhofer, 1991, Pterostichus 487

badgleyi Fowler, 1912, Platyrhopalopsis 211

badia Minowa, 1932, Tachyura [Tachys] [DA] 278

badiola Bates, 1878, Amara [Amathitis] 551

badius Aubé, 1836, Haliplus 32

badius Clark, 1863, Hydrovatus [Hydroporus] 70

badius Omer‑Cooper, 1931, Hydrovatus 70

badius Jeannel, 1960, Trechus 327

badulini Komarov, 1989, Microlestes 422

baeckmanni Lutshnik, 1933, Amara 556

baeckmanni Semenov, 1926, Mnuphorus 358



baeckmanni Jakobson, 1907, Pterostichus [Platysma] 505

baehri Casale, 1997, Laemostenus 533

baehri Kirschenhofer, 1997, Trigonotoma 520

baehri J. Schmidt, 1996, Xestagonum 467

baenningeri Schauberger, 1927, Abax 469

baenningeri Netolitzky, 1926, Bembidion 260

baenningeri Roubal, 1926, Leistus 81

baenningeri Krausse, 1915, Omophron 208

baenningeri Jedlika, 1931, Pterostichus 488



baergi Csiki, 1932, Harpalus 378

baeri Bleuse, 1914, Carabus 195

baeri Ménétriés, 1851, Carabus 172



baerii Motschulsky, 1845, Pterostichus 509

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