Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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sexpunctatus Sharp, 1882, Canthydrus 34

sexpunctatus Schaufuss, 1882, Carabus 158

sexpunctatus Drapiez, 1819, Hydaticus [Dyticus] 54

sexpunctatus Sirguey, 1929, Notiophilus 97

sexpunctatus Chaudoir, 1843, Scarites 233

sexpunctatus Motschulsky, 1861, Syntomus [Apristomorphus] 433

sexpustulatus Fabricius, 1777, Hydroporus [Dytiscus] 64

sexstigmatum Korge, 1964, Agonum 455

sexstriata Duftschmid, 1812, Tachyura [Elaphrus] 279


sexualis Fairmaire, 1881, Abax 470

sexualis Reiche, 1857, Agabus 39

sexualis K. Daniel & J. Daniel, 1898, Platynus 463

sexualis A. Zimmermann, 1919, Rhantus 46

seychellarum Scott, 1912, Polyderis [Tachys] 274

shaanxianus Sciaky & Wrase, 1997, Stomis 517

shaanxicus Deuve & Imura, 1990, Carabus 198

shaanxiensis Deuve, 1991, Carabus 149

shaanxiensis P. Moravec & Wrase, 1998, Trechus 340

shachgireii Zamotajlov, 1991, Carabus 131

shaheshang Imura, 1993, Carabus 149

shahinei Schatzmayr & Koch, 1934, Porotachys [Tachys] [DA] 275

shahristana Kryzhanovskij & Mikhailov, 1987, Amara [Curtonotus] 561

shaid Belousov & Kabak, 1998, Trechus 340

shakhensis Belousov, 1998, Porocimmerites 318

shakhensis Belousov, 1987, Trechus 340

shamaevi Imura, 1996, Carabus 129

shaman Shilenkov, 1996, Carabus 172



shamiliana Lutshnik, 1933, Amara 565

shanghaiensis Kult, 1949, Dyschirius 229

shankoucola Deuve, 1994, Cychrus 203

shanminicus Deuve, 1999, Carabus 128

shansutiaensis Lassalle & Remond, 1998, Carabus 129



shantarensis Lafer, 1989, Dyschirius 227

shantungensis Born, 1910, Carabus [Coptolabrus] 144

shantungensis Mandl, 1971, Cicindela 105

shaovuensis Jedlika, 1956, Pterostichus 495

shapingensis Xie & Yu, 1993, Parena 434

sharanicus Deuve & Letellier, 1980, Carabus 154

sharanus Straneo, 1984, Pterostichus [Amolopsa] 494

shardaghensis Apfelbeck, 1918, Carabus 165

sharovae O. Berlov, 1997, Cychrus 19, 202

sharovae Gryuntal, 1984, Dyschirius 224

sharpi Régimbart, 1895, Bidessus 55

sharpi Joy, 1912, Bradycellus 401

sharpi Wehncke, 1875, Dytiscus 52

sharpi Wehncke, 1880, Haliplus 32

sharpi Peschet, 1917, Hydaticus 53

sharpi Reitter, 1887, Hydroscapha 25

sharpi Jakobson, 1908, Ilybius [Agabus] 43

sharpi Régimbart, 1889, Laccophilus 77



sharpi A. I. Jakovlev, 1896, Rhantus 47

sharpi Jeannel, 1921, Trechus 340



sharplaninae Korge, 1964, Bembidion 20, 269

shatrovskyi Belousov & Kabak, 1994, Trechus 340



shavrini O. Berlov & Anistschenko, 1999, Pterostichus 508

shavsheticus Zaitzev, 1930, Carabus 185

shchurovi Belousov & Zamotajlov, 1997, Caucasorites 297

shchurovi Belousov & Kabak, 1996, Trechus 340

shebbearei Andrewes, 1930, Platynus [Colpodes] 465



sheeti Ali, 1967, Amara 552

shennongjia Imura, 1996, Cychrus 206

shennongjianus Imura, 1996, Carabus 129

shenseensis Perrault, 1991, Leistus 82



shermani Casey, 1913, Pelophila 98

sherpa J. Schmidt, 1998, Rupa 466



sheshnagensis Ledoux, 1980, Carabus 155

shessiensis Imura & Zamotajlov, 1993, Carabus 133

shibanaii Uéno, 1955, Nebria 89

shibatai Uéno, 1964, Agonum [Altagonum] 449

shibatai Habu, 1973, Chydaeus 362

shibatai N. Ito, 1987, Coleolissus 369

shibatai Uéno, 1989, Epaphiopsis 309

shibatai Habu, 1969, Harpalomimetes [Anisodactylus] 363

shibatai N. Ito, 1990, Hyphaereon [Calathomimus] 387

shibatai N. Ito, 1985, Psychristus [Bradycellus] 404

shibatai Ishida, 1961, Pterostichus 502

shibatai Habu, 1978, Synuchus 546

shibatai Habu, 1973, Trichotichnus 397

shibataianum Habu, 1974, Agonum [Platynus] 454

shibataianus Habu, 1978, Trephionus 547

shigaraki Hiura & Katsura, 1971, Carabus [Apotomopterus] 177

shihi Uéno & Saito, 1991, Jujiroa 459

shiibanus Habu, 1958, Pterostichus 513

shiita Lutshnik, 1933, Amara 555

shikokuana Oike, 1939, Cicindela 105

shikokuense Habu, 1974, Agonum [Platynus] 452

shikokuense Morita, 1996, Bembidion [Ocydromus] 266

shikokuensis Nakane, 1961, Carabus 159

shikokuensis Habu, 1973, Chydaeus 362

shikokuensis Habu, 1978, Synuchus 545

shikokuensis Kasahara & Y. Itô, 1995, Trichotichnus 397

shilenkovi Morita, 1989, Bembidion 266

shilenkovi O. Berlov, 1989, Carabus 170

shilenkovi Farka, nom. nov., Leistus 18, 83

shilenkovi Erjiomin & Kabak, 1991, Pterostichus 492

shilenkovi Belousov & Kabak, 1992, Trechus 341

shilinensis Zamotajlov, 1999, Naxipenetretus 284

shima Ishikawa & Kubota, 1995, Carabus 178

shimaga Jedlika, 1959, Chlaenius 349


shimenensis Deuve, 1997, Carabus 123

shimenicus Cavazzuti, 1999, Carabus 182

shimentaiensis Deuve & Tian, 1999, Carabus 127

shimizui Imura, 1994, Carabus 122

shimizuianus Imura, 1994, Carabus 127

shimoheiensis Ishikawa, 1992, Carabus [Leptocarabus] 159

shimomurai Morvan, 1995, Skoeda 466

shimomurai Morvan, 1996, Xestagonum 468

shimomuraianus Morvan, 1995, Platynus [Colpodes] 465

shimonoyai Kurosawa & Kudo, 1980, Carabus [Damaster] 143

shimosae Tanaka, 1956, Tachys 276



shimoyamai Habu, 1974, Agonum [Platynus] 450

shimoyamai Habu, 1978, Bembidion 252



shimoyamai Jedlika, 1962, Brachinus 216

shimoyamai Jedlika, 1962, Pterostichus 495

shinanensis Habu, 1953, Amara [Curtonotus] 561

shinanensis Born, 1922, Carabus 159

shinganensis Shilenkov, 1998, Trechus 341

shingarevi Lafer, 1979, Pterostichus 495

shinnai Nakajima, 1993, Carabus [Acoptolabrus] 122

shirahatai Habu, 1954, Agonum 454

shirahatai Habu & Baba, 1960, Apatrobus [Patrobus] 280

shirahatai Nakane, 1961, Carabus 149

shirahatai Habu, 1947, Nebria 94

shirahatai Habu, 1954, Synuchus [Crepidactyla] 545

shirakii W. Horn, 1927, Cylindera [Cicindela] 108

shirakii Habu, 1973, Stenolophus 405

shiranensis Kanda, 1928, Carabus [Damaster] 148

shirazi Jedlika, 1968, Tachyura [Tachys] 280

shiretokoana Nakane, 1960, Nebria 89

shiroumanus Nakane, 1959, Agabus [Gaurodytes] 36

shirozui Habu, 1954, Hikosanoagonum [Agonum] 458

shirozui Habu, 1985, Pterostichus 513

shirtalensis Gottwald, 1990, Carabus 123

shismatica Antoine, 1957, Amara [Amathitis] 552



shiuni Nakajima, 1993, Carabus [Acoptolabrus] 122

shivalensis Belousov & Kabak, 1998, Trechus 341



shizuokaensis Nakane, 1953, Carabus [Apotomopterus] 177

shkodrensis Scheibel, 1936, Neotrechus 315



shljebensis Mandl, 1985, Carabus 165

shogulaensis Hieke, 1997, Amara 564

shokalskii Semenov & Breuning, 1931, Carabus 160

shoorengensis Habu & Baba, 1962, Apatrobus [Patrobus] 280

shoriensis Obydov, 1999, Carabus 174

shotaroi Morita, 1987, Pterostichus 502



shtchurovi Belousov & Zamotajlov, 1993, Carabus 132

shuamaluko Deuve, 1991, Carabus 189

shuamaluko Farka, 1995, Leistus 81

shun Deuve, 1995, Carabus 127

shunichii Habu, 1973, Bembidion 271

shushensis Belousov & Kabak, 1994, Trechus 23, 325

shuten Uéno, 1978, Trechiama 323

siagonicus Ménétriés, 1849, Pterostichus [Platysma] 485

siagonoides Brullé, 1832, Carterus [Ditomus] 363

sibericus Sharp, 1882, Ilybius [Agabus] 43

sibiriacus Csiki, 1930, Poecilus [Pterostichus] 482

sibirica Csiki, 1929, Amara 556

sibirica Dokhtouroff, 1883, Cicindela [HN] 107

sibirica Fischer von Waldheim, 1821, Cicindela 104

sibirica Fischer von Waldheim, 1821, Cicindela 107

sibirica V. E. Jakovlev, 1891, Cymindis 24, 411

sibirica Csiki, 1902, Nebria 94

sibiricola Lutshnik, 1934, Amara 557

sibiricum Gebler, 1847, Agonum 454

sibiricum Dejean, 1831, Bembidion 250



sibiricum Motschulsky, 1844, Bembidion [Trachypachus] 250

sibiricum Motschulsky, 1850, Bembidion 244

sibiricum Motschulsky, 1844, Calosoma 121

sibiricus J. R. Sahlberg, 1880, Agabus [Gaurodytes] 35

sibiricus Reitter, 1900, Badister 441

sibiricus Motschulsky, 1864, Brachinus 216

sibiricus Gebler, 1841, Calathus 528

sibiricus Fischer von Waldheim, 1820, Carabus 200



sibiricus A. Morawitz, 1862, Demetrias 418

sibiricus Motschulsky, 1844, Diplous [Patrobus] 284

sibiricus Motschulsky, 1844, Elaphrus 207

sibiricus Motschulsky, 1860, Haliplus 31

sibiricus Csiki, 1932, Harpalus 379

sibiricus J. R. Sahlberg, 1880, Hydroporus 65

sibiricus Motschulsky, 1844, Notiophilus 97

sibiricus J. R. Sahlberg, 1880, Olisthopus 461

sibiricus Motschulsky, 1860, Orectochilus 28

sibiricus Gebler, 1829, Pterostichus [Molops] 485

sibiricus Mäklin, 1881, Suphrodytes [Hydroporus] 70

sibiricus Motschulsky, 1844, Syntomus [Dromius] 433

sibylla Holdhaus, 1942, Amara 563

sibyllae Klapperich, 1950, Tenomerga [Cupes] 25

sicanus Csiki, 1927, Carabus 161

sicanus Fairmaire, 1858, Platyderus [Feronia] 522

sicardi Fagniez, 1923, Duvalius 305



sicardi Bedel, 1918, Mesolestes [Microlestes] 421

sicelidis Reiche, 1868, Cymindis 412

sichemita Reiche & Saulcy, 1855, Brachinus 214

sichotana Lafer, 1978, Amara 557

sichotanus Lafer, 1989, Trechiama [Leptepaphiama] 321

sichotensis Born, 1914, Carabus 133

sichuanensis Kataev & Wrase, 1996, Dicheirotrichus 403

sichuanensis Hendrich & Mazzoldi, 1995, Nebrioporus 66

sichuanensis Naviaux, 1994, Neocollyris 116

sichuanensis Morvan, 1998, Paranchodemus [Kenserzh] 462

sichuanensis Wrase & Jaeger, 1995, Psychristus 404

sichuanensis Kirschenhofer, 1997, Synuchus 546

sichuanicola Deuve, 1989, Agonotrechus [Trechus] 289

sichuanicola Deuve, 1989, Carabus 176

sichuanicus Deuve, 1995, Duvalioblemus 298

sichuanum Deuve, 1990, Broscosoma 235

sichuanus Sciaky, 1994, Leistus 81

sichuanus Sciaky & Facchini, 1997, Microzargus 440

sichuanus Deuve, 1988, Trechus 341



siciliana Porta, 1923, Cicindela 104

siciliensis W. Horn, 1891, Cylindera [Cicindela] 111

sicula Dejean, 1831, Amara 566

sicula Gistel, 1837, Cassolaia [Cicindela] 101

sicula Preudhomme de Borre & Ragusa, 1882, Nebria 92

siculorum Schilder, 1953, Cicindela 104

siculum Dejean, 1831, Bembidion 256

siculus Patti, 1844, Brachinus 213

siculus Ragusa, 1882, Canthydrus [Hydrocanthus] 33

siculus Ragusa, 1881, Carabus 195

siculus Baudi di Selve, 1882, Duvalius [Anophthalmus] 305

siculus J. Müller, 1922, Dyschirius 224

siculus Fery, 1995, Graptodytes 60



siculus Régimbart, 1882, Gyrinus 28

siculus Wehncke, 1883, Haliplus 32

siculus Dejean, 1829, Harpalus 382

siculus Dejean, 1826, Licinus 442

siculus Levrat, 1857, Pedius [Argutor] 478



siculus Dejean, 1828, Percus [Feronia] 478

siculus Motschulsky, 1865, Sphodrus [Sphodrus] 543

siderea Bates, 1892, Drypta 359

sidonia Reitter, 1893, Bronislavia 363

sidonicus Chaudoir, 1873, Orthomus 477

sidzhakensis Deuve, 1991, Carabus 147

siebkei Sparre‑Schneider, 1910, Bembidion 262

sienla Jedlika, 1937, Pristosia [Eucalathus] 531



sierranus Piochard de la Brûlerie, 1873, Eocarterus [Ditomus] 365

sieversi Ganglbauer, 1887, Carabus [Sphodristocarabus] 194

sieversi J. R. Sahlberg, 1910, Hydroporus 63

sifanica Tschitschérine, 1894, Amara 561

sifanica Semenov & Znojko, 1928, Nebria 89

sifanicus Semenov, 1898, Carabus 146



sigma Lindroth, 1940, Bembidion 271

sigma P. Rossi, 1790, Philorhizus [Carabus] 426

sigma Uéno, 1980, Trechiama 323

signata W. Horn, 1902, Neocollyris [Collyris] 117

signatellus Klug, 1834, Hydroglyphus [Hydroporus] 56

signaticornis Chaudoir, 1863, Calathus 525

signaticornis Duftschmid, 1812, Harpalus [Carabus] 386

signatipenne Jacquelin du Val, 1852, Bembidion 257



signatipennis Laporte, 1835, Rhantaticus [Hydaticus] 49

signatum Semenov, 1889, Cymbionotum [Coscinia] 235

signatus Grimmer, 1841, Agabus [Colymbetes] 40

signatus Panzer, 1796, Anisodactylus [Carabus] 361



signatus Faldermann, 1836, Cychrus 19, 202

signatus Wollaston, 1857, Trechus 341



signifer Reitter, 1884, Metadromius [Metabletus] 421

signifer Schmidt‑Göbel, 1846, Peliocypas 434



signifera Bates, 1873, Anchista [Paraphaea] 408

siguniangensis Deuve, 1997, Carabus 147

siguniangensis Deuve, 1994, Cychrus 203

sihamensis Britton, 1948, Coryza 221

sikhimensis Mandl, 1982, Cylindera [Cicindela] 112

sikhotealinensis Lafer, 1988, Agabus [Gaurodytes] 40

sikhotealinus Uéno & Lafer, 1994, Trechus 341

sikkimensis Andrewes, 1930, Acupalpus 399

sikkimensis Andrewes, 1930, Amara 554

sikkimensis Deuve & Ledoux, 1987, Apatrobus [Patrobus] [DA] 281

sikkimensis Andrewes, 1921, Callistomimus 348

sikkimensis Mandl, 1982, Carabus 162

sikkimensis Fairmaire, 1901, Cychropsis [Cychrus] 19, 202

sikkimensis Morvan, 1992, Deltocolpodes 457

sikkimensis Jedlika, 1955, Dirotus 458

sikkimensis Brancucci, 1989, Lacconectus 48

sikkimensis Perrault, 1984, Leistus 84

sikkimensis Andrewes, 1940, Oodes 445

sikkimensis Kirschenhofer, 1992, Oxycentrus 392

sikkimensis Andrewes, 1923, Platynus [Colpodes] 465

sikkimensis Régimbart, 1899, Rhantus 46

sikkimensis Morvan, 1999, Skouedhirraad 467

sikkimensis Andrewes, 1934, Synuchus [Calathus] 546


sikkimensis Morvan, 1998, Xestagonum 468

silaceus Dejean, 1831, Porotachys [Bembidium] 274

silaensis Haury, 1881, Carabus 138

silemi Jedlika, 1957, Amara 548

silensis A. Fiori, l899, Syntomus [Metabletus] 433



silesiacus Letzner, 1844, Agabus 38

silesiacus Reitter, 1896, Carabus 181

silesiacus Gautier des Cottes, 1872, Pterostichus [Orites] 491

silfverbergi Hieke, 1996, Amara 552



silicicola Antoine, 1925, Ophonus [Harpalus] 391

silicicola Uéno, 1981, Trechiama 323

sillemi Netolitzky, 1935, Bembidion 262

silphoides Ponza, 1805, Deronectes [Dytiscus] [add, syn of opatrinus (Germar)] 58

silphoides P. Rossi, 1790, Licinus [Carabus] 442



silphoides C. Zimmermann, 1831, Zabrus 568

silphoides Dejean, 1828, Zabrus 569



silvalis Motschulsky, 1865, Synuchus [Taphria] 546

silvanae Monzini, 1995, Boldoriella 296

silvanoi F. Battoni, 1982, Pristosia 531

silvanoides Dieck, 1869, Typhlocharis 241

silvanus Andrewes, 1942, Abacetus 347

silvanus F. Battoni, 1987, Calathus 526

silvatica Deuve, 1983, Aparupa 456

silvatica Mikhailov, 1977, Merizomena 409

silvatica P. Moravec & Wrase, 1998, Ushijimaella 345

silvaticoides W. Horn, 1937, Cicindela 106

silvaticus Jeannel, 1941, Aphoenops 295



silvaticus Coiffait, 1956, Laemostenus [Pristonychus] 539

silvaticus Winkler, 1926, Neotrechus 315



silvaticus Preller, 1862, Poecilus 483

silveiranus Lompe, 1997, Trechus 341



silvester Habu, 1955, Synuchus [Calathus] 545

silvestrii Baliani, 1937, Amara 551

silvestrii Gestro, 1896, Duvalius [Anophthalmus] 305

silvestrii Dvoák, 1993, Odotoncarus [Carterus] 366

silvestrii Gridelli, 1930, Odotoncarus [Carterus] 366

silvestris Morita, 1997, Trichotichnus 397

silvicola C. Zimmermann, 1832, Amara 565

silvicola Jeannel, 1928, Anophthalmus 292

silvicola Löbl, 1995, Sphaerius [Microsporus] 26

silviphilus Deuve, 1993, Carabus 189

simardianus Deuve, 1990, Carabus [ND] 201

simbuaensis Deuve & Hodebert, 1991, Trechus 341

similaris Xie & Yu, 1992, Catascopus 435

similata Gyllenhal, 1810, Amara [Harpalus] 551


similatoides Jedlika, 1934, Amara 550

similatus Betta, 1847, Dyschirius 230

similatus Gautier des Cottes, 1870, Pterostichus [Omaseus] [ND] 516

similis Dostal & Zettel, 1999, Axonya 235



similis Dejean, 1829, Bradycellus [Acupalpus] 401

similis Westhoff, 1881, Cicindela 107

similis Belousov, 1998, Cimmerites 297



similis Régimbart, 1899, Cybister 50

similis Zamotajlov, 1999, Deltomerodes 282



similis Petri, 1891, Dyschirius 225

similis Kirby, 1837, Hygrotus [Hydroporus] 72

similis C. G. Thomson, 1856, Ilybius 43



similis Régimbart, 1889, Laccophilus 77

similis Morvan, 1997, Lepcha 459



similis Chaudoir, 1837, Oodes 445

similis Dejean, 1829, Ophonus [Harpalus] 389



similis W. Horn, 1892, Prothyma [Euryoda] 115

similis Mannerheim, 1852, Pterostichus 492



similis Kirby, 1837, Stereocerus 517

similissimus Deuve, 1988, Carabus 176

simillima Wollaston, 1865, Cymindis [Tarus] 417

simillimus Deuve, 1997, Cychrus 204

simillimus Fairmaire, 1886, Pterostichus [ND] 516

simonae Cavazzuti, 1984, Carabus 194

simoni Heyden, 1879, Carabus 154

simoni Abeille de Perrin, 1881, Duvalius [Trechus] 305

simoni Ganglbauer, 1900, Typhlocharis 241

simonianus Breuning & Ruspoli, 1970, Carabus 193

simonis C. A. Dohrn, 1886, Euplatyrhopalus [Platyrhopalus] 211

simonyi Ganglbauer, 1891, Nebria 86



simonyi Ganglbauer, 1891, Trechus 331

simplex Dejean, 1828, Amara 565



simplex Walker, 1871, Colymbetes 45

simplex Chaudoir, 1876, Coryza 221

simplex Zoubkoff, 1833, Cymindis 416



simplex Chaudoir, 1850, Dyschirius 227

simplex Clark, 1863, Haliplus 31



simplex Duftschmid, 1812, Harpalus [Carabus] 375

simplex Sharp, 1882, Hydrovatus 70

simplex Chaudoir, 1868, Molops 475

simplex Ledoux & Roux, 1996, Nebria 95

simplex Raffray, 1886, Platytarus 410

simplex Semenov, 1891, Synuchus 546

simplicens Belousov & Kabak, 1993, Trechus 341

simplicicollis Wollaston, 1862, Calathus 528



simplicidens A. Morawitz, 1863, Amara 554

simplicidens Schauberger, 1929, Harpalus 385



simplicifrons Apfelbeck, 1904, Dyschirius 227

simplicior Antoine, 1963, Dromius 420



simplicipennis Dejean, 1826, Carabus 197

simplicipes Sharp, 1884, Nebrioporus [Deronectes] 66



simpsoni Stephens, 1828, Agonum 453

simulacrum Rapuzzi, 1997, Carabus 188


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