Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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spinosus Morita, 1990, Apatrobus 280

spinosus Fabricius, 1781, Dineutus [Gyrinus] 26

spinosus Sharp, 1882, Methles 75

spinosus Uéno, 1980, Trechiama 323

spinulifer Uéno, 1985, Trechiama 323

spinulosus Uéno, 1982, Yamautidius 345

spississimus Lesne, 1935, Sphaerius 26



splendens Andrewes, 1926, Amara 564

splendens Andrewes, 1923, Bembidion 251

splendens Olivier, 1790, Carabus 140

splendens Gebler, 1829, Harpalus [Ophonus] 375

splendens N. Ito, 1997, Hyphaereon [Calathomimus] 387

splendens A. Morawitz, 1862, Metacolpodes [Dyscolus] 460

splendens Gené, 1836, Poecilus [Feronia] 479

splendens Gemminger & Harold, 1868, Trechus 341

splendens N. Ito, 1997, Trichotichnus 395


splendens Ganlgbauer, 1915, Zabrus 571

splendida Fischer von Waldheim, 1844, Nebria 89

splendidula Fabricius, 1801, Calleida [Carabus] 408

splendidulum Motschulsky, 1844, Agonum 450

splendidulus Semenov, 1888, Carabus 153

splendidum Heyden, 1870, Asaphidion [Tachypus] 242

splendidum Sturm, 1825, Bembidion 248

splendidus Fischer von Waldheim, 1828, Elaphrus 207



splichali Reitter, 1899, Agabus 36

splugensis Jeannel, 1927, Trechus 340

spoliatrix Tschitschérine, 1898, Pterostichus [Feronia] 514

spoliatus Jakobson, 1907, Acupalpus 398

spoliatus G. Hauser, 1919, Carabus [Coptolabrus] 141

spoliatus P. Rossi, 1792, Chlaenius [Carabus] 351

sporadicum J. R. Sahlberg, 1903, Bembidion 252

spreta Dejean, 1831, Amara 551

spretus Wollaston, 1862, Calathus 526

spretus Dejean, 1826, Carabus 187

springeri J. Müller, 1925, Abax 470

springeri J. Müller, 1931, Anophthalmus 292

springeri J. Müller, 1921, Duvalius [Trechus] 305

springeri J. Müller, 1924, Hydroporus 65

springeri J. Müller, 1928, Orotrechus 317



springeri Schatzmayr, 1929, Poecilus [Pterostichus] 480

spurcaticornis Dejean, 1829, Anisodactylus 360

spurius Kraatz, 1882, Carabus 170

spurius Semenov, 1898, Carabus 142

squamisculptus Reitter, 1908, Carabus 151

squamosus Uéno, 1997, Ishikawatrechus 312

squamosus Uéno, 1982, Yamautidius 345

srnkae W. Horn, 1894, Neocollyris [Collyris] 116

ssangnamensis Kwon & Lee, 1984, Carabus [Lipaster] 174



stabilis Andrewes, 1924, Pheropsophus 217

stackelbergi Kryzhanovskij, 1971, Carabus 153

stackelbergi Kryzhanovskij, 1987, Lebia 430

stackelbergi Kryzhanovskij, 1995, Notiophilus 97



stadleri Gschwendtner, 1922, Dytiscus 52

staehlini M. F. Adams, 1817, Carabus 181



stagnalis Geoffroy, 1785, Dytiscus 52

stagnalis Fabricius, 1787, Hydaticus [Dytiscus] 54

stagnalisaequalis Lapouge, 1916, Carabus 191

stagnatilis Gistel, 1857, Pterostichus [Omaseus] [ND] 516



stagnorum Geoffroy, 1785, Bembidion [Buprestis] 266

stammeri J. Müller, 1938, Italodytes 221



staneki Maan, 1932, Bembidion 253

staneki Štrba, 1931, Carabus [Cytilocarabus] 148

staneki Maan, 1934, Zabrus 570

stankovitchi Jeannel, 1924, Duvalius [Duvalites] 308

stanovskyi P. Moravec, 1993, Trechus 341

starcki Heyden, 1885, Carabus [Neoplectes] 133

starcki Reitter, 1888, Cychrus 19, 202

starcki Heyden, 1885, Pterostichus 501



starcki Reitter, 1887, Pterostichus [Aphaonus] 486

starcki Reitter, 1887, Pterostichus [Lyrothorax] 505

starckianus Ganglbauer, 1886, Carabus [Tribax] 133

starckianus Reitter, 1887, Pterostichus [Aphaonus] 486

starkei Assmann, 1997, Leistus 83

starkii Schaum, 1860, Bembidion 250

starygradensis Born, 1912, Carabus 179

staudacheri Scheibel, 1939, Adriaphaenops 289

staudacheri J. Müller, 1923, Anophthalmus [Trechus] 292

staudingeri Kraatz, 1883, Calomera [Cicindela] 101

staudingeri Ganglbauer, 1886, Carabus 149

staudingeri Jedlika, 1953, Harpalus 383

staudingeri Heyden, 1884, Poecilus 483

staudingeri Jedlika, 1929, Pseudotaphoxenus [Taphoxenus] 542



staudingerianus Schauberger, 1932, Harpalus 383

staurophor Netolitzky, 1930, Bembidion 255

stauropolicus Lutshnik, 1914, Dromius 420

stavropolitanum Lutshnik, 1934, Agonum 453

stearinus Kolenati, 1845, Nebrioporus [Hydroporus] 67

stecki Born, 1902, Carabus [Orinocarabus] 180

stefani Jureek, 1910, Speluncarius [Pterostichus] 517

stefaniruspolii Breuning, 1966, Carabus 154

steinbuehleri Ganglbauer, 1891, Bembidion 258

steindachneri Apfelbeck, 1908, Molops 474

steini Netolitzky, 1914, Bembidion 267



steinwehri G. Hauser, 1932, Carabus [Coptolabrus] 144

stellifer Semenov, 1898, Carabus 184

stenacrum De Monte, 1947, Bembidion 267

steno Bates, 1873, Dyschirius 230

stenocephalus Lucas, 1866, Carabus 137

stenocephalus Apfelbeck, 1918, Duvalius [Trechus] 305

stenocephalus Vereschagina, 1989, Reflexisphodrus [Taphoxenus] 543



stenodera Tschitschérine, 1895, Cymindis 412

stenoderum Chaudoir, 1850, Agonum [Anchomenus] 450

stenoderum Bates, 1873, Bembidion 247

stenoderus Motschulsky, 1866, Abacetus [Astygis] 347

stenoderus Bates, 1873, Brachinus 216

stenoderus Jeannel, 1928, Chaetoduvalius 297


stenoderus Putzeys, 1873, Dyschirius 227

stenoderus Chaudoir, 1850, Pheropsophus 217

stenoderus Chaudoir, 1846, Poecilus [Feronia] 484

stenodes Andrewes, 1935, Hexagonia 407

stenodes Andrewes, 1929, Scarites 234



stenoleptus Semenov, 1899, Pterostichus [Feronia] 485

stenomus Andrewes, 1932, Brachinus 215

stenomus Andrewes, 1936, Deltomerodes [Deltomerus] 282

stenothorax Kabak & Kataev, 1993, Dicheirotrichus [Trichocellus] 403

stenroosi Poppius, 1905, Carabus 145

stentzi A. Villa & G. B. Villa, 1833, Carabus 173

stenuiti Morvan, 1998, Agonum 455

stepanavanensis Iablokoff‑Khnzorian, 1963, Duvalius 305

stepaneki Jedlika, 1951, Lebia 429

stepaneki Jedlika, 1940, Platynus [Colpodes] 465

stepaneki Jedlika, 1952, Poecilus [Pterostichus] 479

stepaneki Maan, 1934, Tapinopterus 520

stepaneki Maan, 1948, Zabrus 571

stephani J. Müller, 1913, Orotrechus [Trechus] 317



stephaniae Schweiger, 1965, Anillidius [Nannotrechus] 290

stephanoforus Cavazzuti, 1998, Carabus 144

stephensii Crotch, 1866, Bembidion 259

stephensii Stephens, 1835, Oreodytes [Hygrotus] 68

steppensis Motschulsky, 1844, Carabus 197

steppensis Guignot, 1954, Haliplus 31



steppensis Branden, 1885, Hygrotus [Hydroporus] 72

steppensis Motschulsky, 1860, Nebrioporus [Hydroporus] 67

sterbai Jedlika, 1935, Archastes 79

sterbai Jedlika, 1965, Bembidion 271

sterbai Jedlika, 1937, Calathus [Eucalathus] 528

sterbai Breuning, 1936, Carabus 181

sterbai Jedlika, 1946, Cymindis 416

sterbai Rambousek, 1909, Deltomerus [Penetretus] 283

sterbai Jedlika, 1932, Diplous 284

sterbai Jedlika, 1940, Platynus [Colpodes] 465

sterbai Jedlika, 1937, Pristosia [Eucalathus] 531

sterbai Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 542

sterbai Jedlika, 1934, Pterostichus 497

sterbai Jedlika, 1931, Setolebia [Lebia] 430

sterilis E. Bodemeyer, 1915, Carabus 185

sternalis Seidlitz, 1887, Neohydrocoptus [Methles] 34

sternbergi Roeschke, 1898, Carabus 183



stethogramme Kolenati, 1845, Dyschirius 224

steuartii A. Deyrolle, 1852, Carabus 130

steveni Ménétriés, 1832, Carabus 201


steveni Quensel, 1806, Chlaenius [Carabus] 355

steveni Krynicki, 1832, Stenolophus 406



stevenii Dejean, 1825, Cylindera [Cicindela] 109

stevenii Dejean, 1829, Harpalus 382



stevenii Chaudoir, 1846, Pangus [Selenophorus] 392

stevenii Fischer von Waldheim, 1817, Zabrus [Pelobatus] 572

stevensi Andrewes, 1924, Brachinus 216

stevensi Andrewes, 1920, Colfax 407

stevensi Andrewes, 1920, Loricera 99

stevensi Schauberger, 1934, Siopelus [Trichotichnus] 394

stevensi Andrewes, 1925, Tachyura [Tachys] 279

stevensi Andrewes, 1923, Taridius 418

stevensianus Jedlika, 1956, Brachinus 216

stichai Jedlika, 1935, Agonum [Colpodes] 453

sticticus Linné, 1767, Eretes [Dytiscus] 53

stictopleurus Fairmaire, 1888, Pterostichus [Omaseus] 501

stictulus Belousov & Kabak, 1998, Trechus 341

stictum Stephens, 1828, Bembidion [Notaphus] 248

stictus Stephens, 1828, Ophonus 392

stierlini Heyden, 1880, Asaphidion [Tachypus] 243

stierlini Heyden, 1881, Carabus 140

stierlini Poncy, 1901, Harpalus 375

stierlini Putzeys, 1870, Perileptus 289



stierlini Gautier des Cottes, 1868, Pterostichus [Feronia] 496

stigmatica Putzeys, 1866, Clivina 220

stigmatophora Fischer von Waldheim, 1828, Cephalota [Cicindela] 102



stigmatophorus Puel, 1935, Harpalus 374

stigmula Dejean, 1826, Nebria 86

stigmulus Chaudoir, 1852, Anomotarus [Cymindis] 408

stillatus Cavazzuti & Ratti, 1998, Carabus 142

stilleri Reitter, 1913, Duvalius [Anophthalmus] 300



stipanovichi Schatzmayr, 1929, Pterostichus 512

stipraisi Barševskis, 1993, Notiophilus 97

stipraisi Belousov & Kabak, 1992, Trechus 341

stirni Pretner, 1959, Adriaphaenops [Aphaenopsis] 289

stjernvalli Mannerheim, 1830, Carabus 160

stobieckii Csiki, 1907, Duvaliopsis [Anophthalmus] 298

stocki Reitter, 1907, Agonum 449

stocki Wewalka & Biström, 1998, Glareadessus 55



stocktonensis Casey, 1924, Harpalus 377

stoeckleini J. Müller, 1937, Dyschirius 229

stoeckleini Straneo, 1941, Pterostichus [Feronia] 501



stoecklini Lopez, 1891, Carabus 130

stoetzneri G. Hauser, 1921, Carabus [Coptolabrus] 142

stoetzneri Roeschke, 1923, Cychrus 205



stoetzneri Schauberger, 1933, Harpalus 379

stoetznerianus G. Hauser, 1924, Carabus [Apotomopterus] 125

stoetznerianus Schauberger, 1932, Harpalus 382

stoicevi V. B. Guéorguiev, 1964, Pheggomisetes 318

stolfai J. Müller, 1931, Anophthalmus 291

stolfai J. Müller, 1943, Bembidion 257

stoliczkana Bates, 1878, Cicindela 105

stoliczkanus Bates, 1878, Carabus 155

stolidus Lapouge, 1925, Carabus 171

stolidus Walker, 1858, Stenolophus [Harpalus] 405

stolzi Bernau, 1914, Carabus 184

stolzi Moczarski, 1913, Molops 475



stolzi Reitter, 1905, Philorhizus [Dromius] 425

stomoides Dejean, 1831, Bembidion 268

stomoides Chaudoir, 1868, Pterostichus [Feronia] 492

stopicensis Jeannel, 1924, Duvalius [Duvalites] [DA] 307

storkani Maan, 1940, Molops 475

storkani Maan, 1939, Nebria 93

storozhenkoi Lafer, 1989, Acupalpus 399

storzicensis Daffner, 1993, Anophthalmus 291

strandi Lutshnik, 1933, Amara 568

strandi Born, 1926, Carabus [Mesocarabus] 168



strandi Lutshnik, 1936, Dicheirotrichus [Trichocellus] 403

strandi Brinck, 1943, Stictotarsus [Deronectes] 69

strandiana Lutshnik, 1934, Amara 564

strandiellus Breuning, 1934, Carabus 156

straneellus Jedlika, 1938, Pterostichus 500

straneoi Jeannel, 1931, Duvalius 305

straneoi Casale, 1988, Laemostenus 535

straneoi Sciaky & Wrase, 1997, Myas [Trigonognatha] 476

straneoi Habu, 1958, Pterostichus 513

straneoi Jeannel, 1931, Trechus 341



strasseri Fuente, 1898, Bembidion 252

strasseri Lauffer, 1905, Carabus [Chrysocarabus] 140

strasseri Reitter, 1896, Cymindis 412



strasseri Reitter, 1900, Harpalus 374

strasseri Reitter, 1898, Pterostichus 505

strasseri Ganglbauer, 1891, Trechus 341

strauchi Born, 1908, Carabus 151

straussi Netolitzky, 1910, Bembidion 262

strenua C. Zimmermann, 1832, Amara 568

strenuus Tschitschérine, 1898, Harpalus 382

strenuus Panzer, 1796, Pterostichus [Carabus] 508

strepens Fischer von Waldheim, 1828, Brachinus 214

strepitans Duftschmid, 1812, Brachinus 213


striata Schmidt‑Göbel, 1846, Dolichoctis 436

striatella Putzeys, 1875, Amara 548

striaticeps Andrewes, 1935, Bembidion 252

striatifrons Chaudoir, 1852, Lophyra [Cicindela] 113

striatipennis Lucas, 1846, Apristus [Dromius] 431

striatipennis J. Müller, 1931, Neotrechus 315

striatipennis Casale, 1988, Pseudotaphoxenus 540



striatocollis Dejean, 1828, Pterostichus [Feronia] 506

striatocollis Gautier des Cottes, 1869, Zabrus 571

striatopunctata Dejean, 1828, Amara 567

striatopunctatus Putzeys, 1846, Dyschirius 227

striatopunctatus Duftschmid, 1812, Poecilus [Carabus] 484

striatoquatermaculatus Goeze, 1777, Badister [Carabus] 440

striatoscutellata Beuthin, 1885, Cicindela 105

striatulus V. Hansen, 1944, Badister 440

striatulus Géhin, 1885, Carabus 178



striatulus Heer, 1838, Lionychus [Dromius] 432

striatulus Putzeys, 1866, Sparostes 221

striatulus Uéno, 1973, Stevensius 320

striatulus Putzeys, 1847, Trechus 341



striatum Stephens, 1828, Agonum 453

striatum Duftschmid, 1812, Bembidion [Elaphrus] 266

striatum Fabricius, 1792, Bembidion [Elaphrus] 257



striatum Paykull, 1798, Bembidion [Elaphrus] 247

striatum Reitter, 1894, Cymbionotum [Coscinia] 235

striatus Motschulsky, 1844, Apristus [Dromius] 431

striatus Fabricius, 1792, Aulonogyrus [Gyrinus] 26

striatus Ballion, 1878, Carabus 178

striatus Chaudoir, 1869, Carabus 177

striatus DeGeer, 1774, Carabus 179

striatus Semenov, 1887, Carabus 129

striatus Linné, 1758, Colymbetes [Dytiscus] 45



striatus Sharp, 1869, Haliplus 30

striatus Depoli, 1930, Laccophilus 77

striatus Hänel, 1940, Molops 475

striatus G. R. Waterhouse, 1833, Notiophilus 96

striatus Lagar Mascaró, 1965, Percus 479

striatus Zeng & Z. Pu, 1992, Platambus [Hydronebrius] 44

striatus P. Rossi, 1792, Pterostichus [Carabus] 509

striatus Dejean, 1825, Scarites 233

stricta Ledoux & Roux, 1991, Nebria 90

stricticaudis Kurnakov, 1961, Lindrothius 529

stricticeps Baliani, 1932, Amara 567

stricticolle Jeannel, 1955, Bembidion [Notaphus] 258

stricticollis Fairmaire, 1886, Andrewesius [Colpodes] 456


stricticollis Baudi di Selve, 1864, Microlestes 423

stricticollis Tschitschérine, 1893, Ophonus 391

stricticollis Casale, 1988, Pseudotaphoxenus 542

stricticollis Solsky, 1874, Pterostichus [Argutor] 508

stricticollis Jeannel, 1927, Trechus 341

strictum Schuler, 1962, Bembidion [Peryphus] 21, 258

strictus Morvan, 1971, Brachinus 216

strictus Dejean, 1828, Percus [Feronia] 478

strictus Baudi di Selve, 1891, Rhegmatobius [Dichropterus] 240

strigata Dejean, 1825, Calomera [Cicindela] 99

strigatus Roberts, 1913, Haliplus 30

strigiceps Reitter, 1905, Paradromius [Dromius] 424

strigicollis R. F. Sahlberg, 1844, Stereocerus [Feronia] 517

strigifrons Fairmaire, 1874, Clivinopsis [Dyschirius] 223



strigifrons Motschulsky, 1860, Dytiscus 52

strigifrons Baudi di Selve, 1864, Notiophilus 96

strigifrons Reitter, 1894, Paradromius [Dromius] 424

strigifrons Wollaston, 1865, Paradromius [Dromius] 424

strigilatera Reitter, 1894, Paradromius [Dromius] 424

strigipennis Fairmaire, 1888, Pristosia [Calathus] 531

strigipennis Kiesenwetter, 1861, Trechus 341

strigitarsis Jedlika, 1958, Pseudotaphoxenus [Taphoxenus] 542



strigosus Ghiliani, 1887, Aulonogyrus [Gyrinus] 26

strigosus Reiche, 1862, Carterus 364

strigosus Sciaky & Wrase, 1997, Pterostichus 514



strigulosus Sharp, 1882, Copelatus 48

strigulosus Régimbart, 1892, Gyrinus 27

strinatii Coiffait, 1956, Turkanillus 238



striola Fabricius, 1792, Abax [Carabus] 470

striola Gyllenhal, 1826, Hydroporus [Hyphydrus] 65



striolata Putzeys, 1866, Amara [Curtonotus] 561

striolata Illiger, 1800, Lophyra [Cicindela] 113

striolatus Zoubkoff, 1833, Acinopus 368

striolatus Gyllenhal, 1808, Agabus [Dytiscus] 41



striolatus Fischer von Waldheim, 1823, Carabus 197

striolatus Fabricius, 1801, Molops [Carabus] 475



striolatus Reitter, 1894, Tachys 276

stristiceps Chaudoir, 1864, Tricondyla 118

stroganowi Zoubkoff, 1837, Carabus 165

strohmi C. G. Thomson, 1874, Laccophilus 76

stronczynskii Weissberg, 1924, Dyschirius 226

strophium Fischer von Waldheim, 1842, Carabus 153

strouhali Mandl, 1955, Carabus 146

strumosus Erichson, 1837, Dyschirius 224

strupii Meschnigg, 1944, Anillus [Anillus] 238


strupii Scheibel, 1937, Duvalius 305

strupii Meschnigg, 1944, Microtyphlus [Scotodipnus] [DA] 240

strupii Winkler, 1933, Omphreus 446

strupii J. Müller, 1935, Pterostichus 485

stscheglowi Mannerheim, 1827, Carabus 137

stschegolewi Poppius, 1906, Carabus 165

stschukini Fischer von Waldheim, 1844, Carabus 183

stschukini Ménétriés, 1837, Chlaenius 354

stschurowskii Solsky, 1874, Carabus 201

stuebeli Heyden, 1892, Phloeozetus 438

stulta Lutshnik, 1935, Amara 552

stultus L. Dufour, 1820, Percus [Broscus] 479

stummuis Morvan, 1995, Platynus [Colpodes] 465

stupaensis Cavazzuti, 1996, Carabus 188

sturanicus Breuning & Ruspoli, 1979, Carabus 158

sturanii Raynaud & Marchal, 1967, Calosoma [Campalita] 121

sturanyi Apfelbeck, 1904, Duvalius [Trechus] 305

sturanyi Apfelbeck, 1906, Molops 474

sturanyi Apfelbeck, 1906, Nebria 85



sturensis Born, 1899, Carabus 180

sturensis J. Daniel, 1903, Pterostichus 504

sturmi Ménétriés, 1832, Calomera [Cicindela] 101



sturmi Schilsky, 1889, Carabus 170

sturmi Babington, 1835, Philorhizus [Dromius] 426

sturmii Gyllenhal, 1808, Agabus [Dytiscus] 37



sturmii Duftschmid, 1812, Bembidion [Elaphrus] 254

sturmii Panzer, 1803, Bembidion [Carabus] 270

sturmii Dejean, 1829, Harpalus 377

sturmii Schaufuss, 1865, Laemostenus [Cryptotrichus] 536

sturmii Gené, 1836, Lionychus [Dromius] 432



sturmii Kraatz, 1875, Molops 474

sturmii Duftschmid, 1812, Olisthopus [Carabus] 461


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