Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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sturmii Dejean, 1831, Pterostichus [Feronia] 491

stussineri Géhin, 1885, Carabus 152

stussineri Ganglbauer, 1897 (as 1896), Laemostenus [Aechmites] 539



stuxbergi Mäklin, 1878, Pterostichus [Feronia] 492

stygia Tschitschérine, 1895, Cymindis 416

stygicus Putzeys, 1875, Miscelus 437

stygicus Chaudoir, 1871, Patrobus 285

stygius Drioli, 1982, Anophthalmus 293

stygius Ganglbauer, 1890, Carabus 160

stygius Jeannel, 1919, Geotrechus 311

stygius Stephens, 1828, Harpalus 382

stygius Régimbart, 1899, Platambus [Agabus] 44

styranus Schauberger, 1921, Pterostichus [Calopterus] 488


styriaca Schaum, 1856, Nebria 85

styriacus Sharp, 1882, Agabus 40

styriacus Winkler, 1933, Arctaphaenops 296



styriacus Kraatz, 1887, Carabus 173

styriacus Seidlitz, 1887, Hydroporus 64

styriacus Schmid, 1970, Laemostenus [Antisphodrus] 535

styriacus Chaudoir, 1871, Patrobus 285



styriacus Grimmer, 1841, Trechus 338

styx Apfelbeck, 1904, Duvalius [Trechus] 308

styx Andrewes, 1937, Pterostichus [Feronia] 516

suan Belousov & Kabak, 1994, Trechus 341

suaneticus Reitter, 1877, Troglocimmerites [Anophthalmus] 344

suavis Ledoux & Roux, 1996, Nebria 95

suavis Sharp, 1882, Nebrioporus [Deronectes] 67

suavissimum Bates, 1883, Agonum [Anchomenus] 451

subachilles Deuve, 1994, Carabus 125

subacuminata Chaudoir, 1843, Nebria 89

subacuminatus A. Fleischer, 1898, Trechus 341



subaenea Sturm, 1825, Amara 556

subaeneum Dejean, 1828, Agonum 452

subaeneus Chaudoir, 1846, Apristus 431



subaeneus Kraatz, 1886, Carabus 171

subaeneus Wollaston, 1864, Dyschirius 225

subaeneus Erichson, 1837, Ilybius 43

subaeneus Bates, 1883, Leistus 84

subaeneus Chaudoir, 1850, Pterostichus [Feronia] 506



subaeneus Motschulsky, 1860, Pterostichus [Orthomus] [HN] 487

subaeneus Motschulsky, 1844, Syntomus [Dromius] 432

subaequalis Schaufuss, 1865, Laemostenus [Pristonychus] 539

subaerea Bänninger, 1921, Nebria 89



subalbanicus Puel, 1938, Harpalus 382

subalpina Glasunov, 1902, Nebria 91

subalpinus Baudi di Selve, 1871, Binaghites [Scotodipnus] 239



subalpinus Bernau, 1913, Carabus 184

subalpinus C. G. Thomson, 1871, Hydroporus 63

subangulata Chaudoir, 1843, Aspidoglossa [Dyschirius] 219

subangulatus Reitter, 1900, Harpalus 373

subapicalis Beuthin, 1899, Cicindela 104

subapicalis Putzeys, 1877, Coptodera 436

subapicalis G. Ochs, 1930, Orectochilus 29

subapicaloides Jedlika, 1956, Coptodera 436



subaquilus Gozis, 1912, Agabus 38

subarctica Kanô, 1933, Cymindis 418

subarcticus Lindroth, 1961, Dyschirius 227

subatratum De Monte, 1949, Bembidion 264


subauratus Kraatz, 1886, Carabus 171

subcaeruleus Stephens, 1828, Harpalus 23, 372

subcallosum Wollaston, 1864, Bembidion 253



subcatenulatus Westhoff, 1881, Carabus 139

subcavicola Uéno, 1955, Trephionus 547



subchalibaeus Reitter, 1900, Harpalus 382

subcoarctatus Hänel, 1940, Pterostichus 493

subcoeruleus Quensel, 1806, Poecilus [Carabus] 484



subcoeruleus Schaum, 1858, Poecilus [Pterostichus] [HN] 484

subconnexum De Monte, 1953, Bembidion 252



subconstrictus Reitter, 1900, Dixus [Aristus] 24, 365

subcontemptus Puel, 1935, Harpalus 376

subcontracta K. Daniel & J. Daniel, 1891, Nebria 87

subconvexa Putzeys, 1865, Amara 551

subconvexum K. Daniel & J. Daniel, 1902, Bembidion 255

subcorax Semenov, 1897, Carabus 192

subcordatus Poppius, 1908, Harpalus 372

subcordatus Dejean, 1829, Ophonus [Harpalus] 391

subcordatus Chaudoir, 1842, Pterostichus 509

subcordatus Chaudoir, 1846, Trechus 341

subcordicollis Bates, 1873, Pentagonica 448

subcoriaceus Géhin, 1885, Carabus 192

subcoriaceus Wollaston, 1861, Eretes [Eunectes] 53

subcostatum Motschulsky, 1850, Bembidion [Peryphus] 263



subcostatus Motschulsky, 1850, Carabus 169

subcostatus Dejean, 1829, Gnathaphanus [Harpalus] 362

subcostatus Gerhardt, 1909, Hydroporus 61

subcostatus Ménétriés, 1837, Pseudotaphoxenus [Sphodrus] 542



subcrenatus Géhin, 1885, Carabus 166

subcrenatus Chaudoir, 1866, Platyderus 523



subcyanea Motschulsky, 1850, Cymindis 415

subcyaneus Rey, 1886, Anisodactylus 361

subcyaneus Kraatz, 1878, Carabus [Sphodristus] 192



subcyaneus Illiger, 1801, Laemostenus [Carabus] 539

subcyaneus Preller, 1862, Poecilus 484

subcylindrica Peyron, 1858, Clivina 220

subcylindricum Reitter, 1892, Bembidion 257

subcylindricus Belousov, 1998, Cimmerites 297

subcylindricus Motschulsky, 1849, Dyschirius 227

subcylindricus Dejean, 1829, Harpalus 382

subcylindricus Semenov, 1891, Pseudotaphoxenus [Taphoxenus] 542

subcylindricus Chaudoir, 1843, Scarites 232

subdepressa Putzeys, 1866, Amara [Leiocnemis] 562

subdepressus N. Ito, 1994, Oxycentrus 392

subdiaphanus Wollaston, 1861, Eretes [Eunectes] 53


subdilatata Motschulsky, 1844, Nebria 89

subdiscola Morvan, 1997, Lepcha 459

subditus Lewis, 1879, Bradycellus [Tachycellus] 402

subelongata Motschulsky, 1849, Amara [Celia] 547

subelongatus Stephens, 1835, Hydroporus 62

subenormis Casey, 1924, Harpalus 377

subexaratus Reitter, 1889, Carabus 164

subfasciatum Chaudoir, 1850, Bembidion 268



subfasciatus A. Zimmermann, 1919, Copelatus 47

subfasciatus Putzeys, 1875, Elaphropus [Tachys] 273

subfasciatus Motschulsky, 1862, Tachys 275

subfastuosus Lie, 1996, Carabus 151

subfenestratum Stephens, 1828, Bembidion [Philochthus] 263

subferrugatus Pater, 1938, Ophonus [Metophonus] 391

subfestivus Oberthür, 1885, Carabus 139

subformosus Semenov, 1887, Carabus 142



subfulgens Jeanne, 1969, Carabus 165

subfuliginosum Habu, 1978, Agonum 452

subfusculus Csiki, 1930, Pterostichus 508

subfuscus Wollaston, 1865, Calathus 528



subfuscus A. Morawitz, 1862, Pterostichus 508

subgibbus Motschulsky, 1860, Pterostichus [Pseudocryobius] 492

subglaber Uéno, 1982, Rakantrechus 319

subglaber Andrewes, 1937, Vachinius [Pristonychus] 356



subglabratus Reitter, 1896, Carabus 179

subglobosum P. Rossi, 1792, Bembidion [Carabus] 244

subglobosum Paykull, 1798, Bembidion [Carabus] 270

subglobosus Uéno, 1987, Taiwanotrechus 321



subgraniferus Semenov, 1898, Carabus 189

subgraniger Reitter, 1896, Carabus 196

subguttatus Crotch, 1873, Haliplus 32

subhamatus Chaudoir, 1856, Chlaenius 348

subiasi Ortuño & Zaballos, 1992, Pterostichus [Haptoderus] 492

subidolon Deuve, 1994, Carabus 121

subimpressicollis Capra, 1940, Pterostichus 513

subimpressum Kirschenhofer, 1989, Bembidion 263



subimpressus Motschulsky, 1845, Acilius 48

subindigestus Deuve, 1994, Carabus 191



subinflatum Motschulsky, 1859, Bembidion [Peryphus] 262

subinterrupta Beuthin, 1899, Cicindela 107

subisraelita Puel, 1938, Scarites 232

subitus Csiki, 1930, Pterostichus 508

sublacerata Solsky, 1874, Cylindera [Cicindela] 111

sublaevicollis Ganglbauer, 1905, Poecilus [Pterostichus] 482

sublaevigatus Tschitschérine, 1898, Harpalus 380


sublaevis Vuillefroy, 1866, Calathus 525

sublaevis Drapiez, 1820, Carabus 166

sublaevis Rey, 1887, Deronectes [Hydroporus] 58

sublaevis Poppius, 1908, Harpalus [Ophonus] 375

sublaevis Falkenström, 1922, Hydroporus 61

sublaevis Solsky, 1873, Notiophilus 98

sublaevis J. R. Sahlberg, 1880, Pterostichus [Feronia] 499

sublaevis Ménétriés, 1836, Zabrus 572

sublimbatus Uéno, 1996, Phreatodytes 35

sublimis Andrewes, 1930, Amara 564

sublinearis Escalera, 1914, Paradromius 424

sublineatus Géhin, 1885, Carabus 187

sublineatus Sharp, 1882, Laccophilus 77

sublivida Semenov, 1889, Nebria 91

sublucida Motschulsky, 1865, Cymindis [Tarus] 417

sublucidus Poppius, 1906, Pterostichus [Cryobius] 492

sublustris Tschitschérine, 1898, Amara 561

submaculatum Poppius, 1906, Bembidion 262

submaculatus Wollaston, 1861, Syntomus [Dromius] 433



submarginatus Stephens, 1828, Dytiscus 52

submarginatus J. R. Sahlberg, 1903, Licinus 442

submarginatus Reitter, 1908, Pogonus 287

submeridionalis Breuning, 1975, Carabus 146



submetallescens Poppius, 1906, Pterostichus 506

submetallicus Nietner, 1858, Abacetus [Distrigus] 347



submetallicus Chaudoir, 1850, Pterostichus [Feronia] 486

submicans Reitter, 1888, Carabus 151

subminimus P. Machard, 1995, Carabus 130

submutatum Netolitzky, 1911, Bembidion 252

submuticus C. G. Thomson, 1874, Hydroporus 65

subnaevulus Reitter, 1903, Trechus 339

subnebulosus Stephens, 1828, Agabus [Colymbetes] 39

subnigrinus Heinz, 1970, Carabus 157

subnigritulus Heinz, 1990, Carabus 157

subnitens Reitter, 1896, Carabus 153

subnitens Chaudoir, 1855, Scarites 232



subnitidus Uyttenboogaart, 1930, Calathus 526

subnivalis Schweiger, 1964, Carabus 131

subnotatus Chaudoir, 1844, Brachinus 215

subnotatus Dejean, 1831, Trechus 341

subnubilis Babington, 1836, Haliplus 32

subolivaceus LaFerté‑Sénectère, 1851, Brachyodes 444



subopacus Wollaston, 1864, Dixus [Aristus] 365

subopacus Chaudoir, 1852, Notiophilus 98

subopacus Mannerheim, 1853, Rhantus [Agabus] 47

subovalis Motschulsky, 1864, Cymindis [Psammastus] 414

subovata Motschulsky, 1864, Lebia [Lebida] 24, 427

subovatus Machado, 1992, Lymnastis 274

subovatus Putzeys, 1875, Platynus [Anchomenus] 464

subovatus Motschulsky, 1860, Pterostichus [Platysma] 487

subpannonicus Daffner, 1994, Orotrechus 317

subpappianus Lassalle, 1999, Carabus 133

subparallelus Ballion, 1878, Carabus [Lipaster] 146

subparallelus Abeille de Perrin, 1904, Geotrechus [Trechus] 310

subparallelus Straneo, 1955, Pterostichus 506

subparallelus Scholz, 1927, Rhantus 46

subparallelus Uéno, 1980, Trechiama 323

subplagiatum J. R. Sahlberg, 1908, Bembidion 264

subplanata Putzeys, 1866, Amara [Amathitis] 552



subpolaris LeConte, 1863, Diacheila 206

subponticus Casale, 1988, Laemostenus 538

subpraestans Kurnakov, 1961, Lindrothius 529

subproductus Chaudoir, 1880, Scarites 232

subpubescens Chaudoir, 1877, Anchista 408

subpunctatus Dejean, 1828, Abax 470

subpunctatus Stephens, 1828, Ophonus 391

subpunctatus Chaudoir, 1866, Platyderus 523

subpunctulus Roeschke, 1907, Carabus [Imaibius] 155



subpurpurascens Eidam, 1941, Carabus 166

subpurpureus Reitter, 1901, Chlaenius 351

subpyrenaeus Barthe, 1923, Carabus 171

subpyrenaeus Jeanne & Zaballos, 1986, Zabrus 569

subquadratus Brullé, 1832, Acinopus 368

subquadratus Motschulsky, 1860, Agabus [Colymbetes] 38

subquadratus Ledoux & Roux, 1998, Archastes 79

subquadratus Dejean, 1829, Ophonus [Harpalus] 389

subreductus Breuning, 1977, Carabus 198

subregularis Semenov, 1897, Carabus 192

subreticulatus G. Hauser, 1921, Carabus [Damaster] 148

subrhodopensis Blumenthal, 1976, Carabus 138

subriparia Schilder, 1953, Cicindela 107

subrobustus P. Machard, 1994, Carabus 157

subrotundatus Motschulsky, 1859, Hydrovatus 71



subrufulus O. Schneider, 1898, Bidessus 55

subrugosus Kraatz, 1877, Carabus [Procrustes] 186

subrugosus Straneo, 1955, Pterostichus 497



subsaharensis Antoine, 1954, Laemostenus [Sphodroides] 539

subschartowi Lapouge, 1898, Carabus 171

subscutellaris Reitter, 1902, Lebia 429

subseriatus J. R. Sahlberg, 1910, Hydroporus 61

subsericatus Régimbart, 1899, Agabus 39

subsericatus Fairmaire, 1886, Andrewesius [Colpodes] 456



subsignata Pic, 1905, Trichis 439

subsimilis Gautier des Cottes, 1870, Calathus 524

subsimilis Chaudoir, 1846, Harpalus 374

subsimilis Tschitschérine, 1893, Poecilus [Feronia] 480



subsimulator Hänel, 1940, Pterostichus 493

subsinuaticollis Puel, 1934, Acinopus 369

subsinuatus Duftschmid, 1812, Harpalus [Carabus] 373

subsinuatus Rey, 1886, Ophonus 391

subsinuatus Dejean, 1828, Pterostichus [Feronia] 492

subspinosus Klug, 1834, Dineutus [Gyrinus] 26

substriatum Andrewes, 1925, Asaphidion 243

substriatum Chaudoir, 1846, Bembidion 252

substriatum Motschulsky, 1859, Calosoma [Callisthenes] 119

substriatus Duftschmid, 1812, Dyschirius [Scarites] 230

substriatus Stephens, 1828, Gyrinus 28

substriatus G. R Waterhouse, 1833, Notiophilus 98

substriatus Ballion, 1871, Pseudotaphoxenus [Sphodrus] 542

substricta Morvan, 1975, Cymindis 413

substrictum LeConte, 1848, Bembidion [Peryphus] 262

substrigatus Sharp, 1882, Colymbetes 45

substrigosa Champion, 1920, Hydroscapha 25

subsulcata J. R. Sahlberg, 1880, Amara 559

subsulcatus Motschulsky, 1848, Harpalus [Bioderus] 372

subsulcatus d'Amore Fracassi, 1903, Percus 478

subsulcipes Uéno, 1983, Oroblemus 316



subsuturalis Souverbie, 1855, Cylindera [Cicindela] 111

subterranea Decou, Nitzu & Juberthie, 1994, Clivina 220

subterranea Putzeys, 1866, Reicheia 222

subterraneus L. Miller, 1868, Duvalius [Trechus] 307



subterraneus Dejean, 1831, Laemostenus [Pristonychus] 539

subterraneus Uéno, 1958, Rakantrechus 319

subterraneus Machado, 1989, Thalassophilus 346

subtile Motschulsky, 1844, Agonum [Anchomenus] 450

subtile Breit, 1912, Asaphidion 243

subtilesculpta W. Horn, 1912, Lophyra [Cicindela] 112

subtilesignata Mandl, 1970, Cylindera [Cicindela] 112

subtilestriatum Mandl, 1954, Calosoma 119

subtilis Motschulsky, 1850, Acupalpus [Stenolophus] 398

subtilis Casey, 1924, Amara [Curtonotus] 559

subtilis Sharp, 1882, Hydrovatus 71

subtilis Erichson, 1837, Ilybius [Agabus] 43

subtilis Uéno, 1957, Ishikawatrechus 312

subtilis Tschitschérine, 1898, Microderes [Neopangus] 387


subtilis Kraatz, 1875, Molops 473

subtilis Schaum, 1857, Ocys [Bembidion] 273

subtilis Zamotajlov & Heinz, 1998, Parapenetretus 284

subtilis Motschulsky, 1844, Pterostichus [Steropus] 493

subtilis R. F. Sahlberg, 1844, Pterostichus [Feronia] 490

subtilis Schaum, 1862, Zabrus 571

subtilissimus Casale & Sciaky, 1999, Laemostenus 540

subtilissimus Sciaky, 1996, Pterostichus 488

subtilistriatus Hauser, 1922, Carabus 160

subtonsus LeConte, 1855, Hydroporus 65

subtrapezicollis Hänel, 1940, Pterostichus 493

subtruncata Chaudoir, 1842, Cylindera [Cicindela] 109

subtruncatum Motschulsky, 1860, Agonum [Tanystola] 453

subtruncatus Chaudoir, 1846, Harpalus 383

subtruncatus Chaudoir, 1843, Molops 474

subtruncatus Fairmaire, 1876, Nebrioporus [Hydroporus] 66

subtypicum Schuler, 1958, Bembidion [Peryphus] 260

subuliformis Farka, 1999, Leistus 81

subvirens Haury, 1885, Carabus 151

subvirens Chaudoir, 1846, Harpalus 376

subvirescens Motschulsky, 1866, Carabus 151

subviridulus Mandl, 1978, Chlaenius 351



subviridulus Mandl, 1992, Chlaenius 351

subvittata Kraatz, 1890, Cephalota [Cicindela] 102

subvittatus Bates, 1892, Syntomus [Metabletus] 433

subvittulus Motschulsky, 1859, Neohydrocoptus [Hydrocoptus] 34

sucainus J. Müller, 1924, Anophthalmus [Trechus] 292

successor Reitter, 1896, Carabus 178

succinctus Klug, 1834, Eretes [Eunectes] 53

succisus Lapouge, 1916, Carabus 171

succyanipennis Imura, 1996, Carabus 127



suchebaatori Jedlika, 1967, Harpalus 380

sudai Morita, 1991, Pterostichus 494



sudanensis Omer‑Cooper, 1974, Bidessus 55

sudanensis Schatzmayr & Koch, 1934, Tachyura [DA] 280



sudeticus Schulz, 1901, Carabus 196

sudeticus Gerhardt, 1909, Pterostichus 511

sudeticus Pawowski, 1975, Trechus 338

sudrei Sainte‑Claire Deville, 1922, Pterostichus 498



sudresi Lapouge, 1898, Carabus 195

suensoni Hieke, 1990, Amara 559

suensoni Mandl, 1981, Anchomenus [Chlaeniomimus] 455

suensoni Zamotajlov & Kryzhanovskij, 1990, Archipatrobus [Patrobus] 281

suensoni Kirschenhofer, 1984, Bembidion 257

suensoni Kirschenhofer, 1986, Brachinus 216


suensoni Kirschenhofer, 1986, Calleida 408

suensoni Mandl, 1981, Calosoma 120

suensoni Mandl, 1979, Carabus 171

suensoni Mandl, 1992, Chlaenius 352

suensoni Kirschenhofer, 1986, Cymindis 24, 415

suensoni Kataev, 1997, Harpalus 386

suensoni Kirschenhofer, 1990, Jujiroa 459

suensoni Kirschenhofer, 1991, Lissopogonus 280

suensoni Mandl, 1981, Myriochila [Cicindela] 114

suensoni Shilenkov & Dostál, 1983, Nebria 90

suensoni Kirschenhofer, 1986, Peliocypas [Risophilus] 434

suensoni Schauberger, 1923, Pheropsophus 218

suensoni Lindroth, 1956, Pristosia 531

suensoni Lindroth, 1956, Synuchus 546

suensoni Jeannel, 1957, Trechus 341

suensonianus Mandl, 1979, Carabus 156

suffriani Westhoff, 1881, Carabus 166

suffriani Loew, 1843, Cicindela 104

suffriani W. Scriba, 1855, Gyrinus 28

suffuscus Andrewes, 1934, Calathus 528



suffusus Wollaston, 1863, Haliplus 33

suffusus Crotch, 1873, Ilybius 43

sugai Ishikawa, 1966, Carabus 134

sugai Nakane, 1987, Elaphrus 207

sugai Uéno, 1984, Trechus 325



suganoe Nakajima, 1993, Carabus [Acoptolabrus] 122

sugillatus Erichson, 1834, Cybister 51

sugimotoi Habu, 1975, Lymnastis 274

sugimotoi Habu, 1975, Trichotichnus 395

suhensis Daffner, 1996, Anophthalmus 291

sui Imura & Zhou, 1998, Carabus 197



sulcalus Rambur, 1837, Platytarus [Cymindis] 410

sulcata Motschulsky, 1851, Tachyura [Tachys] 278

sulcatellus Coiffait, 1958, Anillus 238


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