Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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sulcatissimus Lapouge, 1898, Carabus 194

sulcatissimus Reitter, 1908, Poecilus 483



sulcatoaffinis Hummel, 1822, Acilius [Dyticus] 48

sulcatopunctata Putzeys, 1875, Tachyura [Tachys] 278

sulcatulum Chaudoir, 1846, Bembidion 251

sulcatulus Motschulsky, 1850, Brachinus 213

sulcatulus Faldermann, 1836, Harpalus [NO] 384

sulcatum LeConte, 1848, Bembidion [Ochthedromus] 248

sulcatus A. Fiori, 1899, Abax 469

sulcatus Linné, 1758, Acilius [Dytiscus] 49

sulcatus Belousov, 1998, Alanorites 290



sulcatus DeGeer, 1774, Carabus 194

sulcatus Fabricius, 1792, Dixus [Scaurus] 365

sulcatus Chaudoir, 1876, Eremolestes [Lionychus] 431

sulcatus Coiffait, 1959, Geotrechus 310

sulcatus Stephens, 1835, Pterostichus [Omaseus] 500

sulcatus Fabricius, 1787, Rhysodes [Cucujus] 78

sulcatus Putzeys, 1861, Scapterus 231

sulcatus Olivier, 1795, Scarites 234



sulcibasis Straneo, 1989, Steropanus 517

sulcicolle J. R. Sahlberg, 1880, Bembidion 263

sulcicollis Paykull, 1798, Chlaenius [Carabus] 348

sulcicollis Joseph, 1870, Laemostenus [Sphodrus] 535

sulcicollis Lewis, 1888, Omoglymmius [Rhysodes] 78

sulcicollis Habu, 1978, Oodes 445

sulcicollis Chaudoir, 1843, Siagona 218

sulcifer Tschitschérine, 1902, Ophonus [Harpalus] 390

sulcifrons Chaudoir, 1850, Tachys 275

sulcigera Putzeys, 1866, Clivina 220

sulcipenne J. R. Sahlberg, 1880, Bembidion 266

sulcipenne Reitter, 1889, Bembidion 268

sulcipennis C. R. Sahlberg, 1824, Acilius [Dyticus] (as Zetterstedt, 1824) 48

sulcipennis Mulsant, 1853, Clinidium [Rhysodes] 78

sulcipennis Joseph, 1870, Laemostenus [Sphodrus] 535

sulcipennis Régimbart, 1892, Orectochilus 29

sulcipennis Fairmaire, 1889, Pristosia [Pristonychus] 531

sulcipennis J. R. Sahlberg, 1887, Pterostichus [Feronia] 492

sulcitarsis A. Morawitz, 1862, Pterostichus 486



sulculata Putzeys, 1875, Tachyura [Tachys] 278

sulculatus Cavazzuti, 1999, Carabus 126

sulinensis Born, 1903, Carabus 197

sulphuricola Zaitzev, 1951, Nebrioporus [Deronectes] 67

sulphuripes Germar, 1824, Harpalus 383

suluk Belousov & Kabak, 1996, Trechus 341



sulzeri Brahm, 1790, Callistus [Carabus] 347

sumakovi Poppius, 1912, Hydroporus 60

sumaoi Morita, 1981, Cillenus [Bembidion] 272

sumatrae Régimbart, 1880, Hyphydrus 75

sumatranus C. A. Dohrn, 1891, Dischissus [Panagaeus] 447

sumatrense Louwerens, 1953, Bembidion 264

sumatrensis Herbst, 1806, Calomera [Cicindela] 99

sumavica Obenberger, 1922, Oreonebria [Nebria] 96

sumbarica Putchkov, 1993, Cicindela 103

sumptuosus Csiki, 1927, Carabus 143

sumptuosus A. Morawitz, 1862, Poecilus [Pterostichus] 480

sundarense J. Schmidt, 1995, Xestagonum 468



sundukovi O. Berlov & E. Berlov, 1999, Pterostichus 493

sundukovi P. Moravec & Wrase, 1997, Trechus 341

sungariensis Lafer, 1979, Pterostichus 495

sunicum Iablokoff‑Khnzorian, 1970, Bembidion 263

sunpanensis Semenov, 1898, Carabus 142

sunwukong Imura, 1993, Carabus 158

superans Wollaston, 1854, Amara 565



superbulus Deuve & Li, 1994, Carabus 149

superbus Kraatz, 1878, Carabus 151

superbus Schaufuss, 1872, Carabus 174

superior Sokolá, 1907, Carabus 196

superlita Bates, 1888, Metacolpodes [Colpodes] 460

superna Andrewes, 1923, Nebria 94

superstes Jeannel, 1926, Geotrechus 310

supramontanus Cerruti & Henrot, 1956, Sardaphaenops 319

supremus Schauberger, 1927, Ophonus [Harpalus] 388

surakovi Obydov, 1995, Carabus 164



suramensis Kraatz, 1877, Carabus 201

suramensis Reitter, 1885, Laemostenus [Antisphodrus] 533

surcoufi Antoine, 1933, Pterostichus 499

surdipennis Mateu & Deuve, 1979, Trechus 341



surdum Andrewes, 1935, Bembidion 255

surejae Csiki, 1927, Carabus 186

surigaensis Jedlika, 1935, Chlaenius 353

surinamensis Fabricius, 1792, Harpalus [Carabus] 378

surugana Nakane, 1957, Nebria 91

suruganus Nakane & Straneo, 1979, Pterostichus 513

susamyrensis Hieke, 1988, Amara 561

susamyrensis Belousov & Kabak, 1992, Trechus 341

susanneae Gebert, 1994, Cephalota 102

sushenicus Kataev, 1990, Harpalus 383



sushkini Lutshnik, 1934, Harpalus 380

susica Antoine, 1939, Cymindis 414

susicus Antoine, 1941, Carabus 137



susicus Bruneau de Miré, 1953, Dyschirius 230

susicus Antoine, 1941, Laemostenus 539



susicus Bruneau de Miré, 1949, Poecilus [Pseudopedius] 479

suslovi Lutshnik, 1915, Poecilus [Platysma] 483

sussamyrensis Breuning, 1926, Carabus 192

susterai Jedlika, 1951, Lebia 429

susterai Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 542

susterai Maan, 1943, Tapinopterus [Pterostichus] 520

sutinensis Winkler, 1926, Duvalius 306

sutomorensis Reitter, 1885, Carabus 129

sutomorensis Reitter, 1905, Leistus 84


sutschanense Jedlika, 1958, Agonum 451

sutschanense Jedlika, 1936, Bembidion 265

sutschanensis Jedlika, 1957, Amara 551

sutschanensis Schauberger, 1929, Harpalus 385

sutschanensis Jedlika, 1962, Pterostichus 506

suturale Motschulsky, 1850, Bembidion 257

suturalis Dejean, 1829, Acupalpus [HN] 399



suturalis Stephens, 1828, Acupalpus [Trechus] 398

suturalis Stephens, 1827, Badister 440

suturalis Dejean, 1825, Brachinus 214

suturalis Dejean, 1825, Cymindis 414



suturalis Puel, 1929, Dyschirius 227

suturalis P. W. J. Müller, 1821, Graptodytes [Hyphydrus] 59

suturalis Schaufuss, 1864, Neotrechus [Anophthalmus] 316

suturalis Motschulsky, 1844, Paradromius [Dromius] 425

suturalis Chaudoir, 1846, Parophonus [Ophonus] 393

suturalis Schmidt‑Göbel, 1846, Peliocypas 434

suturalis Schaum, 1863, Pentagonica [Rhombodera] 448

suturalis Westwood, 1845, Platyrhopalus 211

suturalis Lacordaire, 1835, Rhantus 46

suturalis W. S. MacLeay, 1825, Rhantus [Colymbetes] 46

suturalis Putzeys, 1870, Trechus 341

suturangulus Reitter, 1887, Harpalus 383



suturatus Geoffroy, 1785, Acupalpus [Buprestis] 398

suturellus Chaudoir, 1876, Brachinus 216



suturellus Motschulsky, 1844, Microlestes [Dromius] 422

suturellus Harris, 1828, Rhantus [Colymbetes] 47



suturifer Reitter, 1884, Parophonus [Ophonus] 393

suturifer Reitter, 1884, Tachys [Tachys] [DA] 276

suvorovi Semenov, 1912, Chlaenius [Eochlaenius] 353

suvorovi Shilenkov, 1976, Nebria 89



suworowi Semenov, 1907, Carabus 145

suwortzewi Tschitschérine, 1899, Amara 565

suzukaensis Uéno, 1980, Trechiama 323

suzukanus Ishikawa & Kubota, 1994, Carabus 177

suzukii Uéno, 1989, Trechiama 324



svanetica Jureek, 1922, Nebria 91

svanicus Lutshnik, 1922, Pterostichus [Platysma] 501

svanicus Belousov, 1989, Trechus 341

svenhedini Falkenström, 1932, Agabus [Gaurodytes] 41

svetlanae Nilsson, 1989, Agabetes 75

svilajensis J. Müller, 1917, Molops 473

svilajensis Winkler, 1926, Neotrechus 315

svircevi J. Müller, 1925, Neotrechus [Trechus] 315

swaneticus Reitter, 1883, Carabus 181


swaneticus Reitter, 1883, Pterostichus [Feronia] 501

swatensis Heinertz, 1979, Carabus 162

swaticus Casale, 1982, Laemostenus [Pristonychus] 538

swinhoei Bates, 1878, Abroscelis [Cicindela] 99

swinhoei Bates, 1870, Carabus 137

swinhoei Oberthür, 1883, Carabus 148

swinhoei Bates, 1866, Chlaenius 355

sycophanta Fairmaire, 1886, Andrewesius [Colpodes] 456

sycophanta Linné, 1758, Calosoma [Carabus] 120

sycophanta Fairmaire, 1871, Pterostichus [Feronia] 499

sydowi Jeannel, 1930, Duvalius 308

syleus Kirschenhofer, 1997, Pterostichus 502

sylphides Habu, 1975, Agonum [Platynus] 453

sylphis Bates, 1873, Agonum [Colpodes] 453

sylvarum Bates, 1883, Lebia 430



sylvatica Gistel, 1857, Amara [Celia] 565

sylvatica Linné, 1758, Cicindela 107

sylvaticus Dejean, 1826, Carabus 136

sylvaticus C. G. Thomson, 1857, Dyschirius 225

sylvaticus L. Gaudin, 1925, Trechus 327

sylvestris Panzer, 1793, Carabus 181

sylviae Merceron, 1998, Pterostichus 503

sylvicola Dejean, 1822, Cicindela 107

sylvicola K. Daniel & J. Daniel, 1898, Trechus 341

sylvinus Gistel, 1857, Carabus 181

symbolum Kolenati, 1845, Hydroglyphus [Hydroporus] 56

symmetricus Straneo, 1955, Pterostichus 502



synallactes Ganglbauer, 1887, Carabus [Tribax] 132

syriaca Putzeys, 1866, Amara [Leiocnemis] 555

syriaca Trobert, 1844, Calomera [Cicindela] 100

syriaca J. R. Sahlberg, 1908, Clivina 220

syriaca Pic, 1901, Lebia 429

syriaca Gistel, 1837, Megacephala 118

syriaca Csiki, 1928, Tachyura [Tachys] 21, 280

syriacum De Monte, 1953, Bembidion 260

syriacum Chaudoir, 1861, Zuphium 574

syriacus Jedlika, 1961, Apotomus 237

syriacus Chaudoir, 1863, Calathus 526

syriacus Gautier des Cottes, 1866, Calathus [HN] 526

syriacus Pic, 1893, Callistus 347

syriacus L. Redtenbacher, 1843, Carabus 186

syriacus Chaudoir, 1876, Chlaenius 350

syriacus Wewalka, 1971, Deronectes 59

syriacus Putzeys, 1866, Dyschirius 229



syriacus Wehncke, 1880, Haliplus 32

syriacus Brisout de Barneville, 1885, Microlestes [Blechrus] 424

syriacus Dejean, 1829, Ophonus [Harpalus] 391

syriacus Régimbart, 1884, Orectochilus 28

syriacus Chaudoir, 1871, Pogonus 287



syriacus Putzeys, 1870, Trechus 339

syriensis Breuning, 1943, Carabus [Tomocarabus] 198



syriensis Jedlika, 1958, Harpalus 374

syriensis Jedlika, 1957, Poecilus [Pterostichus] 480



syrmiensis Mandl, 1955, Carabus 139

syrum Lapouge, 1930, Calosoma [Campalita] 120

syrus Roeschke, 1898, Carabus 158

szaboi Csiki, 1914, Duvalius [Anophthalmus] 307

szalayi Csiki, 1912, Duvalius [Anophthalmus] 300

szalliesi Kataev & Wrase, 1995, Harpalus 383

szechuanensis G. Ochs, 1929, Gyrinus 28

szechwanensis G. Hauser, 1931, Carabus [Coptolabrus] 144

szechwanensis Falkenström, 1936, Cybister 50

szekessyi Jedlika, 1953, Amara 566



szekessyi Fassati, 1955, Bembidion 244

szekessyi Jedlika, 1961, Bembidion 267

szekessyi Jedlika, 1952, Catascopus 435

szekessyi Jedlika, 1966, Cymindis 411

szekessyi Kult, 1949, Dyschirius 224

szekessyi Jedlika, 1956, Orthomus [Pterostichus] 477

szekessyi Jedlika, 1960, Pristosia [Kanoldia] 531

szekessyi Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 541

szekessyi Jedlika, 1962, Pterostichus 516

szekessyi Jedlika, 1954, Trichotichnus [Iridessus] 397

szekessyi Jedlika, 1960, Xestagonum [Colpodes] 469

szekessyianus Sciaky, 1996, Pterostichus 516

szelenyii Horvatovich, 1974, Nebria 85

szeli Deuve, 1994, Carabus 183

szepligetii Csiki, 1908, Poecilus [Pterostichus] 484

szetschuana Jedlika, 1934, Amara 553

szetschuana Jedlika, 1933, Lebia 430

szetschuana Jedlika, 1934, Pentagonica 448

szetschuana Jedlika, 1932, Pristosia [Eucalathus] 531

szetschuanense Jedlika, 1934, Xestagonum [Colpodes] 469

szetschuanensis Jedlika, 1963, Brachinus 216

szetschuanensis Schauberger, 1930, Harpalus 385

szetschuanensis Tschitschérine, 1889, Pterostichus [Haptoderus] 501

szetschuanus Jedlika, 1932, Andrewesius 456

szetschuanus Breuning, 1932, Carabus 127

szetschuanus Schauberger, 1932, Chydaeus 361

szetschuanus Jedlika, 1932, Diplous 284



szetschuanus Schauberger, 1930, Harpalus 385

szetschuanus Jedlika, 1959, Parapenetretus [Deltomerus] 284

szetschuanus Jedlika, 1962, Poecilus [Pterostichus] 483

szetschuanus Jedlika, 1935, Stenolophus [Anoplogenius] 406



szetschwana Heller, 1923, Galerita 359

szetshuanus Breuning, 1931, Cychrus 205

sziladyi Csiki, 1904, Duvalius [Anophthalmus] 300

szilagyii Csiki, 1912, Anophthalmus 293

szilicensis Bokor, 1922, Duvalius [Duvalites] 307

sziliczensis Csiki, 1912, Duvalius [Anophthalmus] 307



szobroniensis Géhin, 1885, Carabus 196

szoerenyensis Csiki, 1908, Carabus 172

szujeckii Pawowski, 1972, Trechus 342

tabackuriensis Novotný & Voíšek, 1988, Carabus 160

tabalpuri Jedlika, 1965, Orthotrichus 461

tabellatum Wollaston, 1854, Bembidion 246

tabida Reiche, 1855, Cymindis 411

taborskyi Maan, 1947, Carabus [Procrustes] 186

taborskyi Maan, 1939, Tapinopterus 519

tachongi Jedlika, 1936, Pterostichus 516

tachyoides Bates, 1883, Perigona 449

tachypoda Tschitschérine, 1898, Amara 562

tachysoides Antoine, 1933, Ocys [Bembidion] 273

taciturnus Dejean, 1829, Harpalus 383

tacitus Dvoák, 1982, Laemostenus [Antisphodrus] 537

tacoensis Comotti, 1990, Allegrettia 290

tadzhikistanus Kryzhanovskij, 1968, Carabus 153

tadzhikistanus Kryzhanovskij, 1968, Platyderus 523

taebaegsanus Nemoto, 1988, Pterostichus 497

taebaeksaniculus Deuve, 1991, Carabus 149

taebeagsanensis Ishikawa & Kim, 1983, Carabus [Damaster] 142

taebeagsanus Ishikawa & Kim, 1983, Carabus [Lipaster] 174



taeniatus P. Rossi, 1792, Graphoderus [Dytiscus] 49

taeniatus Panzer, 1796, Oxypselaphus [Carabus] 462

taeniatus Wiedemann, 1823, Tetragonoderus [Bembidion] 357



tagawai Nakajima, 1993, Carabus [Acoptolabrus] 122

tagax Andrewes, 1925, Tachyura [Tachys] 280

taghizadehi Morvan, 1974, Platyderus 523

taghizadehi Morvan, 1974, Trechus 342



tagongensis Deuve, 1992, Carabus 146

tahirai Kasahara, 1992, Pterostichus 502



tahodiensis Jeannel, 1936, Trechus 330

tahomae Casey, 1924, Amara [Celia] 565

taibaiensis Zamotajlov & Sciaky, 1999, Chinapenetretus 281

taibaimontanus Imura, 1994, Carabus 198


taibaishanicus Deuve, 1989, Carabus 189

taichii N. Ito, 1992, Chydaeus 362

taichii Kataev & N. Ito, 1999, Trichotichnus 394

taikinus Ishikawa, 1993, Carabus [Procrustes] 164

tailoensis Deuve, 1995, Carabus 123



tainei Lassalle, 1998, Carabus 188

taishi Belousov & Kabak, 1996, Trechus 343

taitii Magrini, 1986, Trechus 330

taiwana Satô, 1990, Allopachria [Microdytes] 73

taiwana Nakane, 1956, Coptodera [Coptoderina] 436

taiwanensis Nilsson & Wewalka, 1994, Agabus 41

taiwanensis Brancucci, 1983, Laccophilus 77

taiwanensis Perrault, 1986, Leistus 81

taiwanensis Hendrich & Balke, 1997, Neptosternus 77

taiwanica Hieke, 1999, Amara 567



taiwanicus Kryzhanovskij, 1956, Carabus 143

taiwanum Netolitzky, 1939, Bembidion 271

taiwanus Habu, 1978, Synuchus 546

taiwanus Habu, 1975, Trichotichnus 395

taiyuanense Kirschenhofer, 1984, Bembidion 263

takabai Habu, 1962, Platynus [Agonum] 463

takachihoi Habu, 1954, Nipponagonum [Agonum] 461

takahashii Miwa, 1935, Hydroscapha 25

takahashii Ishida, 1958, Pterostichus 507

takahasii Uéno, 1952, Kusumia [Ryugadous] 314

takakurai Satô, 1985, Copelatus 48

takakurai Habu, 1954, Trephionus 547

takaosanus Habu, 1958, Pterostichus 513

takasagonis Habu, 1973, Bembidion 20, 266

takashimai Deuve & Imura, 1993, Carabus 150

taketoi Habu, 1962, Nebria 94

takeuchii Habu, 1955, Synuchus [Calathus] 546

takiharensis Katsura & Tominaga, 1978, Carabus [Ohomopterus] 177

taksangensis Straneo, 1977, Pterostichus 510

taksonyis Csiki, 1930, Pterostichus 508



talasensis Grundmann, 1956, Chlaenius 349

talassica Shilenkov, 1982, Nebria 91

talassicus Kabak, 1994, Poecilus 485

talassicus Belousov & Kabak, 1992, Trechus 342



talastauensis Hieke, 1988, Amara 560

talgarensis Matjíek, 1991, Carabus 150

talgarensis Jeannel, 1927, Trechus 342



talianus G. Hauser, 1919, Carabus [Apotomopterus] 124

talibani Savich, 1999, Pterostichus 486

taliensis Fairmaire, 1886, Carabus [Coptolabrus] 188


taliensis Fairmaire, 1886, Lophyra [Cicindela] 113

talpa L. Redtenbacher, 1850, Dixus [Ditomus] 365

talychensis Chaudoir, 1846, Cicindela 107

talyschensis Motschulsky, 1850, Brachinus 216

talyschensis Ménétriés, 1832, Carabus 188

talyschensis Reitter, 1887, Platyderus 523

talyschensis Jedlika, 1952, Taphoxenus 544

tamaensis Yoshida & Nomura, 1960, Trechiama 324

tamang Deuve, 1983, Carabus 162

tamangi J. Schmidt, 1998, Trechus 342

tamarae Wrase & Kirschenhofer, 1991, Pterostichus 505

tamaranus Machado, 1992, Olisthopus 461

tamaranus Machado, 1992, Paradromius 424

tambra Andrewes, 1923, Bembidion 271

tamerlanum Zaitzev, 1918, Calosoma 118

tamgaicus Belousov & Kabak, 1992, Trechus 342

tamsi Ménétriés, 1832, Carabus 182

tamsii Dejean, 1831, Pterostichus [Feronia] 496

tamurensis Deuve & Hodebert, 1991, Trechus 342

tanaicensis Motschulsky, 1844, Dyschirius 224

tanaiticus Lutshnik, 1933, Dixus [Ditomus] 24, 365

tanakai Straneo, 1961, Abacetus 347

tanakai Uéno, 1974, Kurasawatrechus 313

tanakai Uéno, 1999, Kusumia 314

tanakai Ishida, 1964, Pterostichus 498

tanakaorum Morita & Ohkura, 1988, Pterostichus 507



tangeriana Lallemant, 1868, Cicindela 106

tanggorensis Deuve, 1997, Carabus 146

tangjelaensis Shilenkov, 1998, Nebria (as tangjelaenswis) 90

tangkosis Morvan, 1982, Carabus 162

tangkosis Morvan, 1982, Pterostichus 510

tangonis Uéno, 1985, Trechiama 324

tanguticus Semenov, 1887, Carabus 176

tangutorum Kataev, 1993, Harpalus 383

tangweii Li, 1992, Hydroglyphus [Guignotus] 56

tantillus Motschulsky, 1859, Microlestes 423

tantillus Fairmaire, 1888, Pterostichus [Abax] [ND] 516



tantillus Reitter, 1896, Pterostichus 504

tanypedilus A. Morawitz, 1886, Carabus 183

tanzawaensis Ishikawa, 1986, Carabus [Leptocarabus] 149

tanzawanus Habu, 1955, Synuchus [Calathus] 546

tanzawanus Uéno & Sone, 1994, Trechiama 324

taoi Kasahara, 1993, Pterostichus 502

taoi Kasahara, 1989, Stenolophus 406



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