Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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taphrioides Motschulsky, 1844, Harpalus 377

tappesi Gautier des Cottes, 1867, Calathus 527

taprobanicus Sharp, 1890, Rhantus 47



taqueti Lapouge, 1924, Calosoma [Callipara] 119

taradakensis Kasahara & Ohtani, 1988, Pterostichus 513

tarantsha Belousov & Kabak, 1994, Trechus 335

tarasovi Deuve, 1996, Carabus 150

tarbagataica Kabak, 1999, Cymindis 416

tarbagataicus Kraatz, 1878, Carabus 174

tarbagataicus Belousov & Kabak, 1992, Trechus 342

tarcentinus Meggiolaro, 1961, Orotrechus 317



tarda Herbst, 1779, Hygrobia [Dytiscus] 35

tardoides Jedlika, 1965, Harpalus 376

tardoides V. Hansen, 1940, Harpalus 382

tardokijanensis Lafer, 1989, Trechus 342



tardokiyanensis E. Berlov & O. Berlov, 1996, Carabus 165

tardus Panzer, 1796, Harpalus [Carabus] 383



tareli Pic, 1923, Cicindela 105

tareumiut G. E. Ball, 1962, Pterostichus 492

targionii Della Torre, 1881, Orotrechus [Anophthalmus] 317

tarimensis Tschitschérine, 1896, Poecilus [Feronia] 481



tarnensis Géhin, 1885, Carabus 195

tarrieri J. Ochs, 1965, Carabus [Chrysocarabus] 140

tarsalis J. R. Sahlberg, 1875, Calathus 527

tarsalis Mannerheim, 1825, Harpalus 383



tarsalis Apfelbeck, 1904, Pterostichus 508

tarsata Chaudoir, 1876, Drypta 358

tarsatum Zetterstedt, 1828, Agonum [Harpalus] 450

tarsatus Zetterstedt, 1838, Agabus [Dyticus] 40

tarsicum Peyron, 1858, Bembidion 267



tarsicus Jedlika, 1958, Harpalus 375

tartariae Bates, 1878, Amara [Leiocnemis] 558



tartarica Mannerheim, 1837, Cicindela 104

tartarus Bates, 1878, Syntomus [Metabletus] 433

taschkensis Jedlika, 1961, Agatus 408

taschkensis Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 542

taschketensis Jedlika, 1958, Microderes [Harpalus] 387

tashkenticus Kirschenhofer, 1986, Brachinus 213

tasmantus Morvan, 1996, Platynus [Batenus] 463

tassii Vigna Taglianti, 1995, Ocys 273

tastavensis Belousov & Kabak, 1994, Trechus 343

tatarica Motschulsky, 1844, Cicindela 104

tataricus Rost, 1890, Carabus [Plectes] 201

tataricus Gebler, 1829, Cybister [Dytiscus] 51

tataricus Ménétriés, 1849, Parophonus [Ophonus] 392

tatarorum Csiki, 1930, Pterostichus 505


tatianae E. Berlov, 1996, Pterostichus 506

tatrica Schulz, 1909, Cicindela 107

tatrica L. Miller, 1859, Nebria 87



tatricus Roubal, 1938, Agabus 40

tatricus Reitter, 1896, Carabus 181

tatricus L. Miller, 1859, Deltomerus [Patrobus] 283

tatricus Jeannel, 1928, Duvalius 307

tatricus Kinel, 1949, Hydroporus 61

tatricus Kult, 1947, Pterostichus 492

tatsienlui Breuning, 1934, Carabus 176

tatsienluioides Deuve, 1992, Carabus 189

tatzaopin Deuve, 1996, Cychrus 205

tauernensis Mandl, 1984, Carabus 179

taunicus Heyden, 1871, Carabus 172

taurica Motschulsky, 1844, Amara [Celia] 565



taurica Mannerheim, 1837, Cicindela 104

taurica Motschulsky, 1844, Cymindis 412

tauricum J. Müller, 1918, Bembidion 257



tauricum Motschulsky, 1850, Calosoma [Callisoma] 120

tauricus Jeannel, 1930, Anillidius [Nannotrechus] 290

tauricus Bonelli, 1810, Carabus 186

tauricus Ganglbauer, 1888, Carabus [Procrustes] 187

tauricus Roeschke, 1897, Carabus 197

tauricus Motschulsky, 1850, Harpalus [Pseudoophonus] 382

tauricus Motschulsky, 1860, Hygrotus [Hydroporus] 71

tauricus Dejean, 1828, Laemostenus [Pristonychus] 539



tauricus Heyden, 1883, Poecilus [Pedius] 479

tauricus Pliginskiy, 1912, Pseudaphaenops [Trechus] 318

tauricus Winkler, 1912, Pseudaphaenops [Trechus] 318



tauricus Chaudoir, 1837, Scarites 232

tauricus Motschulsky, 1850, Tachys 276

tauricus Chaudoir, 1844, Zabrus 572

taurinensis Baudi di Selve, 1871, Scotodipnus 240

taurulus Andrewes, 1927, Broscus 236

taxi Breit, 1911, Duvalius [Anophthalmus] 300

taxianus Breit, 1912, Duvalius [Anophthalmus] 298

taygetana Rottenberg, 1874, Nebria 87

taygetanum Maan, 1935, Bembidion 254

taygetanus Apfelbeck, 1904, Calathus 524

taygetanus Casale, 1979, Duvalius 305

taygetanus Pic, 1911, Penthophonus 393



taygetanus Heyden, 1883, Zabrus 572

tazekensis Antoine, 1941, Cymindis 413

tazekensis Antoine, 1959, Eocarterus 365

tazekensis Antoine, 1941, Orthomus [Platysma] 477



tazieffi Breuning & Ruspoli, 1970, Carabus 148

teberdanus Jeannel, 1960, Trechus 342

teberdensis Belousov, 1998, Alanorites 290

teberdensis Zolotarev, 1913, Carabus [Tribax] 200



teberdensis Lutshnik, 1928, Pterostichus [Platysma] 501

tectimundi Netolitzky, 1943, Bembidion 270

tectum Motschulsky, 1844, Calosoma 121

tecuciensis Marcu, 1932, Chlaenius 351

tedeschii Sciaky, 1977, Boldoriella [Speotrechus] 296



teicherti Mandl, 1963, Carabus 174

tekesiana Emetz, 1972, Cymindis 416

telavensis Tschitschérine, 1897, Pterostichus [Feronia] 496



telekiana Csiki, 1940, Nebria 87

telekii Csiki, 1937, Carabus 173

telemachus G. Hauser, 1925, Carabus [Apotomopterus] 127

telemus Ragusa, 1892, Bembidion 249

teletskianus Belousov & Kabak, 1994, Trechus 342

telezkianus Shilenkov, 1996, Carabus 172

telionis J. Ochs, 1949, Duvalius 302

tellicus Bruneau de Miré, 1958, Laemostenus [Antisphodrus] 534

telluris Bates, 1883, Carabus 136



telonensis Bonelli, 1813, Scarites 232

teluetica Antoine, 1939, Cymindis 413

temirensis Fedorenko, 1994, Dyschirius 228

temperatus H. Kolbe, 1886, Harpalus [Selenophorus] 380

temperei Aubry, 1974, Agonum [Europhilus] 453

temperei Jeannel, 1947, Pterostichus [Haptoderus] 489

tempestivus Panzer, 1799, Trechus [Carabus] 339

temporalis J. Müller, 1913, Anophthalmus [Trechus] 22, 291

temporalis Bedel, 1909, Penetretus 285



temporalis Apfelbeck, 1902, Trechus 338

tenasserimus Guignot, 1956, Neptosternus 77

tenax Casey, 1918, Bembidion 244

tendanus Born, 1898, Carabus [Orinocarabus] 180

tenebricosus Kurnakov, 1962, Carabus 131



tenebricosus Chaudoir, 1843, Penthus 367

tenebricosus Westhoff, 1881, Poecilus [Feronia] 484

tenebrioides Duftschmid, 1812, Acinopus [Carabus] 368

tenebrioides Waltl, 1838, Penthus [Ditomus] 367

tenebrioides Goeze, 1777, Zabrus [Carabus] 573

tenebrius Heinz & Staven, 1989, Carabus 160

tenebrosa Sloane, 1890, Pseudozaena 208

tenebrosum Motschulsky, 1844, Bembidion [Notaphus] 254

tenebrosus Régimbart, 1880, Copelatus 48

tenebrosus Dejean, 1829, Harpalus 370


tenebrosus Fabricius, 1801, Zabrus [Blaps] 573

tenella Motschulsky, 1850, Nebria 87

tenellum Erichson, 1837, Bembidion 249

tenellus Uéno, 1959, Allotrechiama [Rakantrechus] 290

tenellus Klug, 1832, Chlaenius 355

tenellus Motschulsky, 1862, Polyderis 274

tenenbaumi Jedlika, 1930, Pterostichus 487

tenenbaumianus Jedlika, 1930, Pterostichus 487



tener Csiki, 1927, Carabus 142

tener Coquerel, 1859, Philorhizus [Dromius] 425

tener Uéno, 1989, Trechiama 324

tenerifae Machado, 1984, Orthomus [Wollastonia] 477

tenerifensis Jedlika, 1958, Dicrodontus [Cymindis] 573

tenerifensis Franz, 1981, Trechus 342

teneriffae Wollaston, 1864, Harpalus 370

teneriffensis Jedlika, 1953, Chlaenius 354

tenganensis G. Hauser, 1921, Carabus [Coptolabrus] 141

tengchongicola Deuve, 1999, Carabus 127

tengrensis Jedlika, 1960, Poecilus [Pterostichus] 481

tengrensis Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 542

tenoensis Israelson & Palm, 1979, Trechus 342



tenoinus Enderlein, 1929, Carabus 176

tenryuense Habu, 1954, Asaphidion 243



tensicollis Marseul, 1880, Dyschirius 224

tentek Belousov & Kabak, 1996, Trechus 342

tentiobtusus Morvan, 1979, Laemostenus [Pristonychus] 539

tenuelimbatus Reitter, 1900, Harpalus 374

tenuescens Andrewes, 1929, Dyschirius 230



tenuesculptus Solsky, 1874, Dixus [Aristus] 365

tenuestriata Baliani, 1943, Amara 561

tenuestriatum Fairmaire, 1876, Bembidion 264

tenuestriatus Fairmaire, 1884, Zabrus 568

tenuicervix Imura, 1999, Carabus 188

tenuicollis Uéno, 1986, Agonotrechus 289



tenuicollis Mandl, 1970, Carabus 162

tenuifascia Fischer von Waldheim, 1828, Cicindela 104

tenuiformis Bates, 1883, Carabus 159

tenuiformis Uéno, 1980, Trechiama 324

tenuilimbatus Ballion, 1871, Chlaenius 350

tenuilimbatus K. Daniel & J. Daniel, 1898, Trechus 342

tenuimanus Deuve & Imura, 1990, Carabus 124

tenuimanus Bates, 1873, Dicheirotrichus 403

tenuimarginatum Leoni, 1907, Agonum 449

tenuimarginatus Chaudoir, 1868, Pterostichus [Feronia] 494



tenuipes Lapouge, 1925, Carabus 145

tenuis Fischer von Waldheim, 1828, Cylindera [Cicindela] 109

tenuis Uéno, 1962, Epaphiopsis 309

tenuis Andrewes, 1937, Laemostenus [Pristonychus] 539

tenuis Mateu, 1984, Microlestes 424

tenuis Heinz & Ledoux, 1990, Nebria 90



tenuis LeConte, 1850, Patrobus [Pterostichus] 285

tenuis Marsham, 1802, Stomis [Carabus] 517

tenuis Uéno, 1983, Trechiama 324

tenuissima Bänninger, 1925, Nebria 93

tenuistriata Fairmaire, 1889, Pristosia [Calathus] 531



tenuistriatus Krynicki, 1832, Chlaenius 351

tenuistriatus J. Müller, 1934, Dyschirius 227

tenuistriatus Wollaston, 1864, Platyderus 522

tenuitarsis Kraatz, 1877, Carabus [Ischnocarabus] 156

teodoroi Gridelli, 1930, Microdaccus 422

teradai Habu, 1978, Coleolissus [Tenuistilus] 369

teradai Habu, 1978, Parabroscus 544

teradai Uéno, 1971, Trechiama 324

teradai Habu, 1980, Trichotichnus 397

terahci Charet, 1993, Carabus 153

teranishii Kamiya, 1938, Copelatus 48

teranishii Kamiya, 1933, Orectochilus 28

terceiranus Machado, 1988, Trechus 342



teres Notman, 1921, Amara 549

teres Sharp, 1882, Hydroporus 65



teres Habu, 1958, Nippononebria 95

teretis Park & Kwon, 1996, Pterostichus 497

tergestinus J. Müller, 1905, Typhlotrechus [Anophthalmus] 345

tergluense Netolitzky, 1918, Bembidion 250

teriolensis Schauberger, 1921, Abax 470

teriolensis K. Daniel & J. Daniel, 1898, Platynus 463

terminale Heer, 1841, Bembidion 268

terminale Motschulsky, 1850, Bembidion 253

terminalis Sciaky, 1987, Anophthalmus 291



terminalis Chaudoir, 1865, Neocollyris [Collyris] 117

terminans Gemminger & Harold, 1868, Bembidion 253

terminata Kirby, 1825, Hexagonia 407

terminatus Dejean, 1826, Chlaenius 350

terminatus Panzer, 1793, Leistus [Carabus] 83

ternovensis J. Müller, 1931, Anophthalmus 293

terraenovae Uéno, 1988, Trechiama 324

terrenus Knirsch, 1929, Neotrechus 316

terrestris LeConte, 1848, Amara [Isopleurus] 565

terrestris Motschulsky, 1844, Harpalus [Pseudoophonus] 383

terricola Herbst, 1784, Laemostenus [Carabus] 539


terricola Bonelli, 1813, Scarites 232

terskeiensis Belousov & Kabak, 1992, Leistus 18, 84

terskeiensis Belousov & Kabak, 1992, Trechus 342

teruyai Habu, 1975, Agonum [Platynus] 454



tesari Jedlika, 1952, Diplous 284

tesari Jedlika, 1951, Parena [Bothynoptera] 434



tescicola C. Zimmermann, 1832, Amara 565

tescorum Arnoldi & Kryzhanovskij, 1964, Lymnastis 274



tescorum Tschitschérine, 1899, Pterostichus [Feronia] 509

tesinensis Bernau, 1913, Carabus 184

tesnensis Belousov & Kabak, 1999, Trechus 342



tesselatum Dejean, 1826, Omophron 208

tesselatus Geoffroy, 1785, Agabus [Dyticus] 40

tesselatus Aubé, 1838, Nebrioporus [Hydroporus] 67

tessellatus Drapiez, 1819, Hydroporus [Dytiscus] [NP] 65

tessellatus Dejean, 1831, Platytarus [Cymindis] 410

testacea Baliani, 1943, Amara 566

testacea Olivier, 1811, Nebria 93

testacea Chaudoir, 1872, Parena [Crossoglossa] 434

testaceipenne Ménétriés, 1832, Bembidion 249

testaceoides Fassati, 1947, Bembidion 249

testaceum Duftschmid, 1812, Bembidion [Elaphrus] 250

testaceum Klug, 1832, Zuphium 574

testaceus Dejean, 1825, Apotomus 237

testaceus Rambur, 1837, Brachinus 213

testaceus Boheman, 1858, Canthydrus [Hydrocanthus] 34



testaceus Erichson, 1837, Dromius 420

testaceus Aubé, 1837, Laccophilus 76



testaceus Frölich, 1799, Leistus 82

testaceus Lucas, 1846, Masoreus 357

testaceus Putzeys, 1870, Perileptus 289

testaceus Rambur, 1838, Platyderus [Argutor] 523

testaceus Dejean, 1828, Pogonistes [Pogonus] 286

testaceus Jedlika, 1940, Synuchus [Parcalathus] 546

testaceus Motschulsky, 1850, Tachys 276

testaceus Duftschmid, 1812, Trechus [Carabus] 338

testaceus Fabricius, 1792, Trechus [Carabus] 325

testadilatata Morvan, 1974, Nebria 91



testudinarius Gautier des Cottes, 1867, Calathus 525

testudinea Putzeys, 1865, Amara [Leiocnemis] 555

tethys Netolitzky, 1926, Bembidion 265

tetracoderus Gemminger & Harold, 1868, Trechus 342

tetracolum Say, 1823, Bembidion 263

tetradicus Puel, 1935, Harpalus 374

tetraglochis Andrewes, 1929, Metacolpodes [Colpodes] 460



tetragonoderus Chaudoir, 1876, Chlaenius 353

tetragrammum Chaudoir, 1846, Bembidion 253



tetragrammus Hochhuth, 1846, Hydroglyphus [Hydroporus] 56

tetragrapha Reitter, 1884, Tachyura [Tachys] 279

tetraphacus Bedel, 1896, Tachys 277

tetraporum Bates, 1883, Bembidion 265

tetrasemum Chaudoir, 1846, Bembidion 253



tetrasemus Dejean, 1831, Mochtherus [Thyreopterus] 437

tetrasetosum Habu, 1973, Xestagonum [Agonum] 469

tetraspila Solsky, 1874, Tachyura [Tachys] 21, 280

tetraspilota Chaudoir, 1852, Rhytidophaena [Euryoda] 115

tetraspilotum Stephens, 1835, Bembidion [Peryphus] 263

tetraspilotus W. S. MacLeay, 1825, Mochtherus [Dromius] 437

tetraspilus Solsky, 1874, Mnuphorus [Lebia] 358

tetrasticta Ferrer y Dalmau, 1911, Cicindela 106

tetrastigma Chaudoir, 1850, Bembidion 253

tetrastigma Chaudoir, 1869, Dolichoctis 436

tetsuoi Uéno, 1982, Yamautidius 345

tetungi Shilenkov, 1982, Nebria 95

teunni Nakajima, 1993, Carabus [Acoptolabrus] 122

teutonoides Preudhomme de Borre, 1882, Badister 440

teutonus Schrank, 1781, Stenolophus [Carabus] 406

tev Morvan, 1998, Tostkar 467

tev Morvan, 1999, Xestagonum 468

tewoensis Deuve, 1992, Carabus 154

tewoicus Deuve, 1996, Carabus 121



tewuensis Marcilhac, 1994, Myas [Trigonognatha] 476

textus Lapouge, 1921, Carabus 192

thais Heinz, 1997, Carabus 191

thais Bedel, 1897, Lebia 429

thalassophila Machado, 1987, Orzolina 273

thaleri Franz, 1991, Trechus 342

thalia Andrewes, 1926, Amara 558

tharepatiense Habu, 1973, Xestagonum [Agonum] 469

theanus Reitter, 1895, Carabus 201

thebana Doesburgh, 1895, Cylindera [Cicindela] 111

thebaudi Colas, 1971, Carabus 136

theeli Mäklin, 1878, Pterostichus [Feronia] 492

theodoridesi Mateu, 1961, Microlestes 424

theophilei E. Deyrolle, 1869, Carabus 193

theresiae Casale, M. Etonti & Giachino, 1992, Orotrechus 317

thermalis Uéno, 1995, Accoella 289

thermalis Germar, 1838, Hydroglyphus [Hydroporus] 56

thermarum Motschulsky, 1844, Bembidion 255

thermarum Motschulsky, 1850, Carabus 157


thermarum Steven, 1806, Mastax [Brachinus] 217

thermoides Nakane, 1963, Bembidion 259

thermonectoides Sharp, 1884, Hydaticus 54



therondi Puel, 1929, Dyschirius 228

theryi Antoine, 1953, Amara [Celia] 556

theryi Peyerimhoff, 1925, Deronectes [Hydroporus] 59

theryi Alluaud, 1930, Platydela 114

theryi Guillebeau, 1897, Pseudanillus [Microtyphlus] 238

theseus Briansky, 1910, Carabus 133

thessalicus Casale, 1982, Laemostenus 533

thessalicus J. Müller, 1930, Molops 475

thessalicus Reitter, 1886, Tapinopterus 520

thessalicus Meixner, 1928, Trechus 342



thessalonicensis Apfelbeck, 1904, Carabus 182

thessalonicensis Schatzmayr, 1909, Oodes 445

thessalonicus Schatzmayr, 1943, Pterostichus 485

thessalonicus Jeannel, 1930, Trechus 342



thessalus Putzeys, 1873, Calathus 525

theveneti Chevrolat, 1874, Zabrus 569



thianschanensis Breuning, 1934, Carabus 160

thianschanicus Lapouge, 1925, Carabus 178

thianschanicus Gschwendtner, 1923, Dytiscus [Macrodytes] 52

thianshanskii Breuning, 1932, Carabus 123

thibali Coiffait, 1958, Aphoenops 294

thibetana Tschitschérine, 1894, Amara 553



thibetana Blanchard, 1871, Cicindela 105

thibetanum Fairmaire, 1887, Calosoma 120

thibetanus Breuning, 1950, Carabus 163

thibetanus Fairmaire, 1893, Cychrus 205

thibetanus Dvoák, 1994, Leistus 81

thibetanus Morvan, 1998, Paranchodemus [Kenserzh] 462

thibetanus Casale, 1988, Pseudotaphoxenus 542

thibetanus Régimbart, 1899, Rhantus 47

thibetanus Jeannel, 1928, Trechus 342

thibeticola Breuning, 1964, Carabus 176

thibeticum Fassati, 1957, Bembidion 262

thielei Brandmayr & Zetto Brandmayr, 1984, Oxypselaphus [Anchus] 462

thierryi Ledoux & Roux, 1995, Archastes 79

thilliezi Deuve, 1994, Carabus 127

thirkii Chaudoir, 1857, Carabus [Procrustes] 186



thisbe Antoine, 1951, Amara 547

thomsoni C. Roth, 1897, Acupalpus [Stenolophus] 399

thomsoni J. R. Sahlberg, 1871, Agabus [Gaurodytes] 37



thomsoni Fairmaire, 1858, Carabus 161

thomsoni Sharp, 1882, Colymbetes 45

thomsoni Zaitzev, 1908, Gyrinus 27

thomsonianus Géhin, 1876, Carabus 192

thomsonii Seidlitz, 1887, Graphoderus 49

thomsonii Reiche, 1860, Paussus 211



thongdii Fleutiaux, 1919, Heptodonta 115

thonitida Ledoux & Roux, 1990, Nebria 87


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