Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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treulandi J. Müller, 1909, Aphaenopidius [Trechus] 293

treutikum Morvan, 1996, Xestagonum 469

triangularis Reitter, 1897, Cymindis 415



triangularis Semenov, 1889, Paradromius [Dromius] 424

triangularis Nietner, 1858, Tachys [Bembidium] 275

triangulatus Schilsky, 1888, Dolichus 530

triangulus Ledoux & Roux, 1995, Archastes 79

tribilensis Meggiolaro, 1959, Anophthalmus 293

tribulatus Cavazzuti, 1995, Carabus 125

trichophorus Tschitschérine, 1897, Harpalus 383

trichothorax Bezina & Imura, 1997, Carabus 135



tricolor Fabricius, 1801, Bembidion [Carabus] 246

tricolor Reitter, 1887, Calathus 527

tricolor M. F. Adams, 1817, Cicindela 104

tricolor Gebler, 1845, Merizomena [Cymindis] 24, 409

tricolor Naviaux, 1991, Neocollyris 117

tricolor Schaufuss, 1862, Pheropsophus 217

tricolor Fabricius, 1798, Poecilus [Carabus] 483

tricostatus Chaudoir, 1868, Lesticus 471

tricuspidata Dejean, 1831, Amara 568

tricuspidatus A. Morawitz, 1863, Anisodactylus 361

tricuspidatus Fabricius, 1792, Ditomus [Scaurus] 365

tricuspius Andrewes, 1929, Dyschirius 228



tridens A. Morawitz, 1862, Amara 568

tridens A. Morawitz, 1862, Harpalus 386

tridentatus Fischer von Waldheim, 1821, Harpalus 387

tridentinus Depoli, 1939, Abax 470

tridentinus Bertolini, 1887, Carabus 196

tridentinus J. Müller, 1951, Laemostenus 535

tridentinus J. Müller, 1918, Molops 474



trifasciatus Wollaston, 1846, Bidessus [Hydroporus] 55

trifasciatus Chaudoir, 1876, Tetragonoderus 357

trifidus Panzer, 1795, Hydroglyphus [Dytiscus] 56

trigintaduo Cavazzuti & Ratti, 1998, Carabus 127



trigintiduosulcatus Lapouge, 1898, Carabus 164

trigintistriatus Mandl, 1974, Carabus 162

triglyphicus J. Ochs, 1965, Carabus 197

triguttata Herbst, 1806, Cylindera [Cicindela] 112

triimpressum R. F. Sahlberg, 1844, Agonum [Bembidium] 453



trilineatus Haller, 1885, Carabus 172

trilineatus Feng, 1936, Hygrotus [Coelambus] 72

trilobatus Cavazzuti, 1994, Carabus 124

trimaculata Gebler, 1829, Lebia 429

trimaculata Villers, 1789, Lebia [Carabus] 429

trimaculatum Andrewes, 1927, Bembidion 245

trimaculatus Drapiez, 1820, Haliplus 33

trimaculatus Olivier, 1790, Macrocheilus [Carabus] 407

trimaculatus Dejean, 1826, Panagaeus 447

trinii Fischer von Waldheim, 1817, Zabrus [Pelobatus] 572

trinotatus LaFerté‑Sénectère, 1851, Callistoides [Chlaenius] 348

triops Uéno, 1994, Trechiama 324



tripolitana Schulz, 1909, Calomera [Cicindela] 101

tripolitanum Schatzmayr, 1937, Bembidion 252

tripunctata Bates, 1873, Parena [Bothynoptera] 434

triramosa Kollar, 1836, Cylindera [Cicindela] 110

triseriatus Kirby, 1837, Colymbetes 45

triseriatus Putzeys, 1878, Deltomerus 283

triseriatus A. Fleischer, 1897, Harpalus 383

triseriatus Gebler, 1847, Pterostichus [Platysma] 506

trisetifer Jeannel, 1926, Duvalius 307

trisetosus Zamotajlov & Sciaky, 1996, Naxipenetretus [Deltomerodes] 284

trisignata Dejean, 1822, Cylindera [Cicindela] 111

trisignata Ménétriés, 1848, Lebia 429

triste Dejean, 1828, Agonum 450

triste Andrewes, 1935, Asaphidion 243



triste Fabricius, 1792, Bembidion [Carabus] 251

triste Schilsky, 1888, Bembidion 248

tristicula Csiki, 1932, Galerita 359

tristicula Reitter, 1888, Nebria 87

tristiculus Kraatz, 1878, Carabus 171

tristiculus Semenov, 1887, Carabus 178

tristiculus K. Daniel & J. Daniel, 1898, Trechus 342



tristis Latreille, 1804, Acinopus [Harpalus] 368

tristis Aubé, 1838, Agabus 41



tristis Baliani, 1937, Amara 550

tristis Putzeys, 1866, Amara [Curtonotus] 560

tristis Motschulsky, 1850, Carabus 134

tristis Schaller, 1783, Chlaenius [Carabus] 350



tristis V. E. Jakovlev, 1887, Cymindis 416

tristis Stephens, 1827, Dyschirius 227

tristis Chaudoir, 1843, Galerita 359

tristis Lutshnik, 1922, Harpalus [Smirnovia] 378

tristis Tschitschérine, 1898, Harpalus 375

tristis Paykull, 1798, Hydroporus [Dytiscus] 65

tristis Uéno, 1989, Masuzonoblemus 314

tristis Lindroth, 1956, Synuchus 546

tristis Duftschmid, 1812, Trechus [Carabus] 336

tristis Schönherr, 1806, Trechus [Carabus] 340

tristis Stephens, 1828, Trechus 337

trisulcatum Germar, 1840, Clinidium [Rhysodes] 78

triton Fall, 1922, Agabus 37

tritus Casey, 1920, Patrobus 22, 285

tritus Bates, 1889, Platymetopus 393

triumphalis W. Horn, 1902, Prothyma [Cicindela] 115

triumphans Cavazzuti, 1987, Carabus 158



triumphator Schaufuss, 1882, Carabus 158

trivialis Duftschmid, 1812, Amara [Carabus] 550

trivialis Gyllenhal, 1810, Amara [Harpalus] 547

trivialis Tschitschérine, 1898, Harpalus 383

trivialis Stephens, 1828, Hydroporus 63

troberti Kraatz, 1860, Carabus 140

trochanteratus Emden, 1954, Taphoxenus 544

trochanteribus Mateu, 1990, Sphodrus 543

troglavensis J. Müller, 1917, Molops 472

troglodytes Erichson, 1842, Distichus [Scarites] 231

troglodytes Uéno, 1955, Jujiroa 459

troglodytes Schaufuss, 1863, Platyderus 523

troglodytes Jeannel, 1920, Trechus 331

troglodytes Jeannel, 1924, Zariquieya [Molops] 521

troglomontanus Absolon & Maan, 1947, Neotrechus 315



troglophilus Csiki, 1907, Duvalius [Anophthalmus] 299

troglophilus Mateu, 1997, Pterostichus [Tinautius] 514

troglophilus Uéno, 1953, Tachys 276

trogossitoides L. Dufour, 1820, Dixus [Aristes] 365

trojanus Dejean, 1826, Carabus 182



tromsoensis Sparre‑Schneider, 1888, Dicheirotrichus [Bradycellus] 402

tronqueti Ledoux & Roux, 1995, Archastes 79

tronqueti Deuve, 1995, Kozlovites 313

tronquetiana Deuve, 1995, Epaphiopsis [Pseudepaphius] 309



trophina Gistel, 1857, Amara [Celia] 556

trophonius Abeille de Perrin, 1872, Geotrechus [Anophthalmus] 311



tropidurus Bates, 1873, Pterostichus 493

trossulus Semenov, 1904, Elaphrus 207



troyeri Born, 1911, Carabus 151

trsiensis Zamotajlov, 1996, Deltomerus 282

trubilini Zamotajlov, 1997, Pterostichus 486

truilheti Coiffait, 1952, Trechus 337

truncatellus Linné, 1761, Syntomus [Carabus] 433

truncaticollis Eschscholtz, 1833, Carabus 134

truncaticollis Heinz & Ledoux, 1990, Nebria 90

truncaticollis Uéno, 1970, Rakantrechus 319

truncatodentatus J. Müller, 1936, Molops 473

truncatus Coiffait, 1959, Aphoenops 294



truncatus Fabricius, 1792, Dromius [Carabus] 419

truncatus Rosenhauer, 1842, Harpalus 381

truncatus Nietner, 1858, Polyderis [Bembidium] 274

truncatus Dejean, 1828, Pterostichus [Feronia] 509

truncatus Park & Kwon, 1996, Pterostichus [HN, repl. in vol. 2] 502

truncatus Habu, 1978, Synuchus 546

truncatus Rambur, 1837, Trymosternus [Cymindis] 410

truquii Guérin‑Méneville, 1855, Neolaphyra [Cicindela] 114

trybomi J. R. Sahlberg, 1880, Amara 554

trypalsi Breuning, 1975, Carabus 168

tryznai Farka, 1999, Leistus 81

tryznaiana Ledoux & Roux, 1998, Nebria (as traznaiana) 89

tsaidamensis Fedorenko, 1993, Dyschirius 227



tschalikovi Samko, 1932, Bembidion 257

tschapecki Géhin, 1885, Carabus 184

tschegeti Gottwald, 1983, Carabus 138

tscheliensis Lapouge, 1918, Carabus 199

tscheresovae Komarov, 1995, Dicheirotrichus [Trichocellus] 403

tscherkessicus Heinz & Korge, 1964, Carabus 200

tscherkessicus Reitter, 1896, Pterostichus 504



tschernyschovi Obydov, 1998, Carabus 191

tschetschenicus Semenov, 1897, Carabus 169

tschetschenicus Tschitchérine, 1897, Pterostichus [Feronia] 496

tschikatunovi Mikhailov, 1977, Cymindis 418



tschiliensis G. Hauser, 1921, Carabus [Coptolabrus] 144

tschiliensis Schauberger, 1929, Harpalus 385

tschitaensis Jedlika, 1957, Amara 556

tschitscherinella Hieke, 1990, Amara 561

tschitscherini Semenov, 1894, Agonum [Anchomenus] 452

tschitscherini Lutshnik, 1935, Amara 553

tschitscherini Jakobson, 1906, Bembidion 250

tschitscherini Semenov, 1897, Carabus 137

tschitscherini Semenov, 1903, Chilotomus 364

tschitscherini A. I. Jakovlev, 1896, Colymbetes [Cymatopterus] 45

tschitscherini Semenov & Znojko, 1927, Coryza 221

tschitscherini Semenov, 1895, Cymindis 415


tschitscherini Reitter, 1899, Dicheirotrichus [Trichocellus] 403

tschitscherini Semenov, 1895, Discoptera 356

tschitscherini Semenov, 1898, Elaphrus 207

tschitscherini Semenov, 1909, Laemostenus 533

tschitscherini Semenov, 1906, Leistus 84

tschitscherini Semenov, 1900, Merizomena [Glycia] 409



tschitscherini Glasunov, 1902, Nebria 91

tschitscherini Semenov, 1891, Poecilus [Pterostichus] 485



tschitscherini Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 540

tschitscherini Semenov, 1904, Stomis 518



tschitscherini Iablokoff‑Khnzorian, 1962, Tachys 21, 275

tschitscherini Schauberger, 1936, Trichotichnus 394

tschitscherinianus Jakobson, 1907, Pterostichus [Platysma] 494

tschuchtschorum J. R. Sahlberg, 1887, Pterostichus [Feronia] 497

tsekuricus Belousov, 1992, Carabus 131

tsengjialiae Šustek, 1996, Dischissus 447

tshabanorum Kabak, 1998, Poecilus 484

tsharynensis Kabak, 1994, Carabus 146

tshatkalica Kabak, 1999, Cymindis 416

tshatkalicus Kabak, 1997, Poecilus 485

tshchaltensis Novotný & Voíšek, 1988, Carabus 200

tshemalensis Gebert, 1995, Cicindela 106

tshetshenicus Zamotajlov, 1992, Deltomerus 283

tshildebaevi Belousov & Kabak, 1992, Trechus 342

tshitscherini Semenov & Znojko, 1929, Colpostoma 441

tshitsherini Zaitzev, 1916, Notiophilus 98

tshitsherini Belousov, 1987, Trechus 342

tshuiliensis Kabak, 1994, Carabus 145

tsingtaoriensis Cheo, 1934, Gyrinus 28



tsingtauensis G. Hauser, 1913, Carabus [Coptolabrus] 144

tsogoensis Deuve, 1997, Carabus 176

tsuchimae Breuning, 1932, Carabus 177

tsudai Uéno, 1960, Kusumia 314

tsugaru Nakane, 1993, Hyphydrus 74

tsukubanus Takami & Ishikawa, 1997, Carabus 176

tsukubasanus Kasahara, 1988, Pterostichus 499

tsukushiensis Yoshimura, 1932, Haliplus 32

tsurugaensis Ishida, 1959, Pterostichus 513

tsurugaensis Uéno, 1980, Trechiama 324

tsurugiensis Habu, 1976, Apatrobus [Patrobus] 280

tsurugiyamanus Habu, 1959, Pterostichus 507

tsurugiyamanus Habu, 1959, Trichotichnus 397

tsushimae Habu, 1981, Nebria 94

tsushimana Habu, 1961, Coptodera 436

tsushimanum Habu & Baba, 1959, Agonum 453



tsushimanus Habu & Baba, 1968, Harpalus 386

tsutsuii Uéno, 1954, Cillenus [Armatocillenus] 272



tsylbulskii Deuve & Dolin, 1991, Carabus 134

tuberatus A. Morawitz, 1862, Carabus 191

tuberculata Motschulsky, 1844, Blethisa 206



tuberculatus Dejean, 1826, Carabus 195

tuberculatus Fischer von Waldheim, 1828, Carabus 153

tuberculatus Gryuntal, 1984, Dyschirius 226

tuberculatus Mäklin, 1878, Elaphrus 207



tuberculifer J. Müller, 1922, Dyschirius 225

tuberculifer Koenig, 1906, Pterostichus 496

tuberculiger Tschitschérine, 1897, Pterostichus [Feronia] 494

tuberculipennis Mandl, 1974, Carabus 162

tuberculipennis Mandl, 1987, Cychropsis 202

tuberculosus Dejean, 1829, Carabus 153



tuberculosus Géhin, 1885, Carabus [Procerus] [HN] 185

tuberosus J.‑P. Nicolas, 1973, Pterostichus 490



tuerkheimi Harold, 1877, Carabus 137

tuerki Beuthin, 1886, Calomera [Cicindela] 100

tuiukensis Obydov, 1998, Carabus 145

tujngoli Jedlika, 1966, Bembidion 268

tukchensis Straneo, 1983, Pterostichus 510

tulatensis O. Berlov & Shilenkov, 1996, Carabus 170

tuleschkovi Maan, 1933, Zabrus 571

tulgoeti Mateu, 1972, Pterostichus 503

tumida A. Morawitz, 1862, Amara 561

tumidiceps Munster, 1924, Elaphrus 207

tumidipennis Uéno, 1997, Junaphaenops 313



tumidum Gemminger & Harold, 1868, Bembidion 258

tumidus Jeannel, 1921, Trechus 342

tumidus Reiche & Saulcy, 1855, Zabrus 572

tundrae Tschitschérine, 1894, Pterostichus [Feronia] 506

tunetanum De Monte, 1947, Bembidion 256

tungchengensis Li, 1993, Carabus [Coptolabrus] 143

tungus Obydov, 1999, Carabus 174

tungus Zaitzev, 1910, Hydroporus 63

tungusicus Poppius, 1906, Pterostichus [Cryobius] 493

tunisensis Jedlika, 1958, Laemostenus 539

tunisius J. Müller, 1924, Dyschirius 230

tunkunensis A. Zimmermann, 1928, Agabus [Gaurodytes] 36

tunkunensis Hieke, 1990, Amara 561

turanensis Jedlika, 1959, Pterostichus 506

turanica Jedlika, 1957, Amara 561

turanicum Csiki, 1928, Bembidion 262



turanicus Reitter, 1894, Dicheirotrichus [Trichocellus] 403

turanicus Znojko, 1928, Dyschirius 226

turbinata Champion, 1925, Hydroscapha 25

turca Seidlitz, 1887, Nebrioporus [Deronectes] 67

turcati Giordan & Raffaldi, 1983, Duvalius 305

turcestanicus Gschwendtner, 1934, Ilybius 42

turcica Hieke, 1976, Amara 566

turcica Schaum, 1859, Cephalota [Cicindela] 101

turcica Fabricius, 1787, Lebia [Carabus] 429

turcica Chaudoir, 1843, Nebria 87



turcicum Ganglbauer, 1891, Bembidion 252

turcicum Gemminger & Harold, 1868, Bembidion 253

turcicus Jaeger, 1992, Acupalpus 399

turcicus J. Frivaldszky, 1880, Anillidius [Anophthalmus] 290



turcicus Schweiger, 1969, Bradycellus 400

turcicus Motschulsky, 1866, Carabus 131

turcicus Jedlika, 1958, Harpalus 383

turcicus Casale, 1988, Laemostenus 536

turcicus Coiffait, 1956, Parvocaecus [Winklerites] 240

turcicus I. Frivaldszky von Frivald, 1835, Paussus 211

turcicus Jedlika, 1963, Phloeozetus 438

turcicus Apfelbeck, 1904, Platynus 463

turcicus Brandmayr & Drioli, 1978, Pterostichus [Platysma] [HN!] 511



turcicus Ménétriés, 1836, Pterostichus [Abax] 495

turcmenica Tschitschérine, 1894, Amara 562



turcmenicus Motschulsky, 1865, Chlaenius [Epomis] 353

turcmenus Guignot, 1957, Agabus 37

turcomanicum Reitter, 1908, Parazuphium [Zuphium] 574

turcomannicum Motschulsky, 1844, Calosoma 120

turcomannicus Motschulsky, 1850, Carabus 197

turcomannicus Motschulsky, 1850, Pterostichus 509

turcomanorum Thieme, 1881, Carabus [Pantophyrtus] 183

turcosinensis Mandl, 1955, Carabus 146

turcosinensis Mandl, 1938, Cephalota [Cicindela] 103



turculus Bates, 1878, Harpalus 377

turfanus Jedlika, 1969, Pterostichus 485

turgenicus Belousov & Kabak, 1994, Trechus 342



turgidus Erichson, 1843, Herophydrus [Hyphydrus] 71

turinensis Jeannel, 1922, Trechus 329

turkestana Kryzhanovskij, 1974, Amara [Curtonotus] 563



turkestanica Csiki, 1929, Amara 548

turkestanica Beuthin, 1892, Calomera [Cicindela] 100

turkestanica Ballion, 1871, Cicindela 108



turkestanica Tschitschérine, 1896, Cymindis 415

turkestanica Jedlika, 1966, Lebia 429

turkestanicoides W. Horn, 1938, Cicindela 108


turkestanicum Fassati, 1957, Bembidion 261

turkestanicum Jedlika, 1960, Colpostoma 441

turkestanicus Ballion, 1871, Anchomenus 456

turkestanicus Liebke, 1928, Brachinus 216



turkestanicus Heyden, 1880, Carabus 192

turkestanicus Csiki, 1932, Harpalus 381

turkestanicus Semenov, 1891, Laemostenus 539

turkestanicus Reitter, 1891, Poecilus 484

turkestanicus Lutshnik, 1935, Pogonus 286



turkestanicus Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 542

turkestanicus Heyden, 1884, Scarites 233

turkestanicus Csiki, 1928, Tachys 276

turkestanicus Belousov & Kabak, 1992, Trechus 342

turkestanus Breuning, 1928, Carabus 146

turkmenicus Kirschenhofer, 1988, Apristus 431



turkmenicus Fedorenko, 1989, Dyschirius 224

turkmenicus Komarov, 1996, Olegius 286

turkmenika Putchkov, 1993, Cephalota [Cicindela] 102

turnai Deuve, 1998, Amerizus 242

turnai Toledano, 1998, Bembidion 252

turnai Deuve, 1994, Carabus 191

turnai Deuve, 1994, Cychrus 206

turnai Deuve & Quéinnec, 1993, Trechus 342

turnaianus Deuve, 1995, Carabus 135

turnaianus Deuve, 1993, Trechus 342

turnaioides Deuve, 1998, Trechus 342

turukensis Belousov & Kabak, 1992, Trechus 343



tusnadensis Csiki, 1908, Poecilus [Pterostichus] 484

tutancamon Schatzmayr, 1936, Clivina 220

tuvensis Shilenkov, 1996, Carabus 165

tuxeni Mandl, 1979, Carabus 127

tuxeni Jeannel, 1957, Trechus 343

tvrtkovici Casale & Jali, 1999, Croatotrechus 297

tychus Jeannel, 1960, Trechus 343

typhlops Uéno, 1996, Dimitshydrus 73



typhonius Gistel, 1857, Carabus 164

typhonius Gistel, 1857, Harpalus 387



tyrannous Li, 1993, Carabus [Coptolabrus] 143

tyrannus Lapouge, 1911, Carabus [Coptolabrus] 144

tyrolense Schweiger, 1975, Asaphidion 242



tyrolensis Kraatz, 1876, Carabus 180

tyrrhenicum Jeannel, 1941, Bembidion [Peryphus] 261

tyrrhenicus Csiki, 1930, Poecilus [Pterostichus] 484

tyrrhenicus Jeannel, 1927, Trechus 343

tyschkanica Kryzhanovskij & Shilenkov, 1976, Nebria 89


tyshkanensis Deuve & Quéinnec, 1992, Trechus 343

ubachi Español, 1965, Geotrechus 310

ubinensis Puisségur, 1955, Carabus [Cechenus] 155

ubinensis Bruneau de Miré, 1964, Nebria 87

ubykh Belousov, 1998, Porocimmerites 318

uchiyamai Morita, 1987, Pterostichus 502

ucrainicus Lazorko, 1951, Carabus 184

ucrainicus Lazorko, 1954, Leistus 84

udege Nilsson, 1994, Agabus 41

udege Lafer, 1989, Harpalus 383



udensis Jedlika, 1957, Amara 548

uedaorum Morita & Hirasawa, 1996, Pterostichus 498

uenoi Habu, 1971, Agonum 452

uenoi Habu, 1972, Amara 565

uenoi Jedlika, 1965, Bembidion 265

uenoi Habu, 1973, Broscosoma 235

uenoi Ishikawa, 1960, Carabus [Apotomopterus] 177

uenoi Habu, 1975, Chydaeus 362

uenoi Imura, 1995, Cychrus 206

uenoi Nakane, 1963, Hydroporus 65

uenoi Satô, 1972, Microdytes 75

uenoi Habu, 1972, Minypatrobus [Patrobus] 284

uenoi Habu, 1967, Mochtherus [Dolichoctis] 437


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