Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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uenoi Habu, 1978, Myas 476

uenoi Nakane, 1963, Nebria 94

uenoi Paik & Lafer, 1995, Onycholabis 461

uenoi Straneo, 1955, Pterostichus 507

uenoi Habu, 1976, Rupa 466

uenoi Habu, 1975, Stenolophus 405

uenoi Lindroth, 1956, Synuchus 546

uenoi Tanaka, 1960, Tachys [DA] 275

uenoi Habu, 1969, Trichotichnus 395

uenoi P. Moravec & Wrase, 1998, Ushijimaella 345

uenoiana Habu, 1972, Nebria 91

uenoianum Morita, 1996, Bembidion [Ocydromus] 256

uenoianus Imura, 1995, Carabus 192

uenoianus Habu, 1974, Tachys 276

uenorum Kasahara & Y. Itô, 1995, Trichotichnus 397



uenoshiba Jedlika, 1965, Bembidion 247

uhagoni Bolívar y Pieltain, 1912, Broscus 236



uhagoni Gautier des Cottes, 1870, Calathus 525

uhagonii Seidlitz, 1887, Hygrotus 73

uhagonii Crotch, 1869, Trechus 332

uhligi Holdhaus, 1904, Amara 563

ukokensis Belousov & Kabak, 1994, Carabus 150



ukokensis Belousov & Kabak, 1996, Trechus 342

ulani Jedlika, 1968, Pterostichus 490

ulcerosus Lapouge, 1902, Carabus 138

ulgeni Gottwald, 1982, Carabus 200

uliginosus Linné, 1761, Agabus [Dytiscus] 37

uliginosus O. F. Müller, 1776, Agabus [Dyticus] 37

uliginosus Hochhuth, 1871, Chlaenius 350

uliginosus Putzeys, 1846, Dyschirius 227

uliginosus Fabricius, 1792, Elaphrus 207



uliginosus Erichson, 1837, Platynus [Anchomenus] 464

ulkei G. H. Horn, 1870, Pelophila 98

ullrichii W. Redtenbacher, 1842, Elaphrus 207



ulrichhoffmanni Lie, 1989, Carabus 172

ulrichi Heinz, 1965, Poecilus [Pterostichus] 482

ulrichi Pawowski, 1976, Trechus 343

ulrichii Germar, 1824, Carabus 152

ultima Ledoux & Roux, 1998, Nebria 93

ultimus Obydov, 1999, Carabus 170

ulubeyensis Kirschenhofer, 1981, Pterostichus 492

uludagensis Hieke, 1984, Amara 563

uludagensis Schweiger, 1966, Amara [Bradytus] 553

uludagensis Schweiger, 1965, Anillidius [Nannotrechus] 290

uluderensis Shilenkov & Heinz, 1984, Nebria 87

umbilicatus Ledoux, 1984, Ledouxius [Penetretus] 284

umbralis Gistel, 1857, Pterostichus [ND] 516

umbraticus Uéno, 1983, Trechiama 324

umbratulosa W. Horn, 1930, Cylindera [Cicindela] 109

umbratus Wollaston, 1865, Philorhizus [Dromius] 426

umbratus Ménétriés, 1832, Platyderus [Feronia] 523

umbricola Gistel, 1857, Pterostichus [ND] 516

umbricola Wollaston, 1854, Trechus 343

umbrigerum Chaudoir, 1876, Zuphium 574

umbrina Heer, 1837, Oreonebria [Nebria] 95

umbrinus Motschulsky, 1855, Philodytes [Laccophilus] 78

umbrosa Motschulsky, 1851, Tachyta [Tachys] 277

umbrosus A. Zimmermann, 1926, Herophydrus 71

umbrosus Gyllenhal, 1808, Hydroporus [Hyphydrus] 65



umbrosus Gschwendtner, 1930, Methles 75

umeayi Habu, 1959, Bembidion 256

umekii Imura & Mizusawa, 1999, Carabus 177

umeyai Habu, 1959, Bembidion 272

umiatense Lindroth, 1963, Bembidion 244

umpyrensis Gottwald, 1985, Carabus 131



unanensis Born, 1909, Carabus 173

uncatus Uéno, 1986, Trechiama 324



uncinata Baehr, 1998, Gunvorita 573

uncinata Kasahara, 1994, Rupa 466

unctulatus Duftschmid, 1812, Pterostichus [Carabus] 493

unctulum Antoine, 1941, Bembidion 20, 249

unctulus Andrewes, 1932, Paropisthius 98

unctus Lapouge, 1908, Carabus 130

uncus Guignot, 1959, Hydrovatus 71

undata Motschulsky, 1850, Cicindela 106

undecimlineellus Fairmaire, 1877, Nebrioporus [Hydroporus] 67

undulata Dejean, 1825, Myriochila [Cicindela] 114



undulatum Sturm, 1825, Bembidion 250

undulatus Schrank, 1776, Agabus [Dytiscus] 37

undulatus Gebler, 1841, Microderes [Ophonus] 387

undulifer Motschulsky, 1859, Laccophilus 77

ungensis Csiki, 1906, Carabus 196

unguicularis C. G. Thomson, 1867, Agabus [Eriglenus] 41

unguicularis Crotch, 1874, Hygrotus [Hydroporus] 72

unguicularis J. R. Sahlberg, 1880, Hygrotus [Coelambus] 72

unguinosa Ledoux, Roux & Sciaky, 1994, Nebria 88



ungularis LeConte, 1862, Ilybius [Colymbetes] 41

unicolor Dejean, 1828, Agonum 452

unicolor Chaudoir, 1850, Bembidion 264

unicolor J. Müller, 1918, Bembidion 270

unicolor Kraatz, 1882, Carabus 170

unicolor Brullé, 1834, Dicrodontus [Polistichus] 573

unicolor Motschulsky, 1844, Dyschirius 227

unicolor Chaudoir, 1848, Glycia 409



unicolor A. Fiori, 1914, Haliplus 32

unicolor Munster, 1922, Haliplus [HN] 32

unicolor Motschulsky, 1844, Harpalus 383

unicolor Schilsky, 1888, Hygrotus 73

unicolor A. Fleischer, 1900, Lionychus 431

unicolor Schilsky, 1888, Lionychus 432

unicolor Putzeys, 1846, Miscelus 437



unicolor Motschulsky, 1866, Nebria 94

unicolor Germar, 1839, Parazuphium [Zuphium] 574

unicolor Jedlika, 1940, Platynus [Colpodes] 465

unicolor Westwood, 1833, Platyrhopalus 211

unicolor Ragusa, 1887, Tachyura [Tachys] 278

unicoloripes Poppius, 1910, Pterostichus 499

unicus Minowa, 1932, Scarites 234

unidentatus Aubé, 1838, Dineutus [Dineutes] 26

unidentatus Uéno, 1969, Stygiotrechus 320



unifasciatus Panzer, 1796, Blemus [Carabus] 296

unifasciatus Dejean, 1831, Somotrichus [Lebia] 439



uniformis Motschulsky, 1844, Harpalus [Pseudoophonus] 383

uniformis Feng, 1936, Laccophilus 76

uniformis Motschulsky, 1859, Laccophilus 77



uniformis Reitter, 1905, Philorhizus [Dromius] 425

unilineatus Reitter, 1888, Elaphropus [Tachys] 273

unilineatus Schrank, 1781, Graptodytes [Dytiscus] 59

unilineatus Báguena Corella, 1935, Nebrioporus [Hydroporus] 66

uninotata Andrewes, 1935, Hexagonia 407



unipunctata Schauberger, 1932, Acinopus 369

unipunctata Dokhtouroff, 1885, Cicindela 105

unipunctatus Jacquelin du Val, 1857, Badister 440

unipunctatus Germar, 1824, Demetrias [Dromius] 419

unipustulatus Bonelli, 1813, Badister 440



uniseriata Reitter, 1923, Cymindis 416

uniseriatus Vuillefroy, 1866, Calathus 526

unisetiger Uéno, 1997, Trechus 343

unistriatus Goeze, 1777, Bidessus [Dytiscus] 55



unistriatus Schrank, 1781, Bidessus [Dytiscus] 55

unistriatus Putzeys, 1875, Elaphropus [Tachys] 273

unistriatus Reitter, 1887, Elaphropus [Tachys] 273

unistriatus Apfelbeck, 1904, Pterostichus 495

univestis Jedlika, 1940, Dromius [Lebidromius] 421

unzenensis Jeannel, 1930, Epaphiopsis [Trechus] 309

uozumii Uéno, 1951, Ishikawatrechus [Nipponotrechus] 312

uozumii Uéno, 1955, Ryugadous [Yuadorgus] 319

uozumii Uéno, 1982, Yamautidius 345



upsaliensis Gmelin, 1790, Chlaenius [Carabus] 350

uralense Fassati, 1944, Bembidion 263



uralensis Motschulsky, 1844, Amara [Acrodon] 553

uralensis Motschulsky, 1850, Amara [Leirus] 560

uralensis Semenov, 1902, Dromius 419

uralensis Glasunov, 1901, Nebria 89

uralensis Motschulsky, 1850, Pterostichus [Eurypessus] 506

urartaeus Pavesi & Sciaky, 1994, Trechus 343

urbanus Minowa, 1932, Scarites 234

urbionensis Arribas, 1991, Nebria 88

urbionensis Jeanne, 1970, Platyderus 522

urbionensis Jeanne, 1970, Zabrus 569



urcitana Mateu, 1952, Tachyura 278

urcitanus Mateu, 1963, Trymosternus 410

urengaicus Jureek, 1924, Pterostichus 506

urgens Tschitschérine, 1898, Poecilus [Feronia] 481

urgensis A. I. Jakovlev, 1899, Hygrotus [Coelambus] 73

uriensis Schauberger, 1933, Harpalus 377

urinator Illiger, 1807, Gyrinus 28


ursuius Lapouge, 1911, Carabus 140

ursuius Jeannel, 1937, Laemostenus [Ceuthosphodrus] 533

ursulae J. Schmidt, 1998, Xestagonum 469

ursus J. Schmidt, 1998, Xestagonum 469

uruktensis Kabak, 1990, Carabus 178

urumbashi Kabak & Putchkov, 1996, Nebria 90

usgentense Solsky, 1874, Calosoma [Callisthenes] 119

usgentensis Schauberger, 1932, Chilotomus 364

usgentensis Heyden, 1884, Eocarterus [Sabienus] 366

usgentensis Belousov & Kabak, 1998, Trechus 343

ushishirensis Obydov & Saldaitis, 1996, Carabus 164

uskokensis Scheibel, 1934, Anophthalmus 291



uskutdaghensis P. Machard, 1991, Carabus 201

usslaubi Saulcy, 1870, Typhloreicheia [Reicheia] 223



ussuricus Born, 1914, Carabus 144

ussuricus Mlyná, 1979, Harpalus 383

ussuricus Kataev & Dudko, 1997, Stenolophus 406

ussuriense Jedlika, 1965, Asaphidion 243

ussuriensis Lafer, 1989, Acupalpus 399

ussuriensis Lutshnik, 1935, Amara 551

ussuriensis Jedlika, 1937, Badister 441

ussuriensis Lafer, 1989, Bradycellus 401

ussuriensis Born, 1914, Carabus 136

ussuriensis Jedlika, 1963, Drypta 359

ussuriensis Fedorenko, 1991, Dyschirius [Dyschiriodes] 229

ussuriensis Chaudoir, 1863, Harpalus 386



ussuriensis Tschitschérine, 1898, Harpalus 383

ussuriensis Lafer, 1989, Masuzoa 314



ussuriensis A. Morawitz, 1862, Nebria 89

ussuriensis Nilsson, 1997, Platambus [Agabus] 44



ussuriensis Jedlika, 1953, Pseudotaphoxenus [Taphoxenus] 540

ussuriensis Tschitschérine, 1897, Pterostichus [Feronia] 508

ustulatus Dejean, 1829, Dicheirotrichus [Harpalus] 402

ustum Quensel, 1806, Bembidion [Carabus] 244



ustum Stephens, 1828, Bembidion [Peryphus] 243

ustus Andrewes, 1927, Platynus [Anchomenus] 463

usun Belousov & Kabak, 1992, Trechus 341

usurpator Cavazzuti, 1987, Carabus 158

utschderensis Reitter, 1890, Trechus 343

uvidum Andrewes, 1924, Bembidion 260

uygur Kabak, 1997, Poecilus 484

uygur Deuve, 1993, Trechus 343

uygurorum Belousov & Kabak, 1994, Trechus 343

uyttenboogaarti Klynstra, 1937, Dyschirius 226

uyttenboogaarti Emden, 1929, Nesacinopus 388



uyttenboogaarti Jeannel, 1936, Trechus 343

uzbek Deuve & Kaláb, 1992, Carabus 147

uzbekorum Zaitzev, 1953, Hygrotus [Coelambus] 73

uzokensis Obenberger, 1917, Nebria 92

vaccai Gestro, 1885, Duvalius [Anophthalmus] 303



vachoni Jeannel, 1937, Oreonebria 96

vadoni Colas, 1962, Carabus 195

vafer Csiki, 1927, Carabus 181

vafer Duftschmid, 1812, Olisthopus [Carabus] 461

vafra Lutshnik, 1935, Amara 564

vafra Andrewes, 1935, Tachyura [Tachys] 280



vagabunda Duftschmid, 1812, Amara [Carabus] 548

vagabunda Andrewes, 1935, Tachyura [Tachys] 280

vagabundus Belousov, 1998, Cimmerites 297

vagabundus Antoine, 1961, Laemostenus [Pristonychus] 535

vagans Tschitschérine, 1897, Amara 558

vagans Olivier, 1795, Carabus 197

vagans Sharp, 1882, Colymbetes 45

vagans Jeannel, 1960, Trechus 335

vagelineatus A. Zimmermann, 1922, Laccophilus 77

vagemaculata Breit, 1914, Cymindis 414

vagepictus Fairmaire & Laboulbène, 1855, Hydroporus 65

vagepictus Fairmaire, 1875, Paradromius [Dromius] 424

vagepunctatus Heer, 1837, Pterostichus 513

vaghezzae Ghidini, 1937, Duvalius 301

vagus Brinck, 1943, Hyphydrus 74

vaillanti Schuler, 1956, Bembidion [Peryphus] 272



vaitoiani Strasser, 1900, Carabus 131

valadolensis Gautier des Cottes, 1866, Syntomus [Metabletus] 433

valassinae Monguzzi, 1982, Boldoriella 296

valbonensis Jeannel, 1927, Trechus 343

valcarceli Wrase, Riuz‑Tapiador & Zaballos, 1998, Leistus 83

valdanii Guérin‑Méneville, 1859, Graphipterus 360

valdenbergi Mandl, 1981, Cylindera 110



valderiensis Capra, 1940, Pterostichus 513

valens Cavazzuti & Ratti, 1999, Carabus 124



valens LeConte, 1863, Plochionus 409

valentinae Obydov, 1997, Calosoma [Callisthenes] 118

valentinae Lafer, 1996, Chaetapatrobus 281

valentinei Suenson, 1957, Trechoblemus 324

valeriae Fancello, 1988, Typhloreicheia 223

valesiacus Ganglbauer, 1891, Pterostichus 504

valida Fairmaire, 1859, Amara 567

valida Ledoux & Roux, 1996, Nebria 90

validiceps Reitter, 1887, Pterostichus 505


validicollis Sciaky & Pavesi, 1995, Trechus 343

validicornis Baudi di Selve, 1891, Typhloreicheia 223



validior Rey, 1886, Harpalus 379

validior Alluaud, 1939, Polyderis [Tachys] 274

validior Tschitschérine, 1889, Pterostichus 514

validipes Tschitschérine, 1888, Amara [Pterostichus] 561

validipes K. Daniel, 1902, Trechus 343

validiusculus Piochard de la Brûlerie, 1873, Carterus 364

validiusculus Chaudoir, 1859, Pterostichus [Feronia] 503

validula Tschitschérine, 1898, Amara 554

validulus Tschitschérine, 1895, Acalathus 529

validum Netolitzky, 1920, Bembidion 269

validus Csiki, 1906, Carabus 173

validus Kraatz, 1884, Carabus [Pseudotribax] 189

validus Chaudoir, 1846, Deltomerus [Cardiomera] 283

validus Régimbart, 1899, Dytiscus 52

validus Dejean, 1828, Pterostichus [Feronia] 514

validus Schaum, 1862, Zabrus 572

valikhanovi Kabak, 1990, Carabus 150

valikhanovi Belousov & Kabak, 1993, Trechus 343

vallestris K. Daniel & J. Daniel, 1898, Trechus 343

valleti Casale & Genest, 1986, Aphoenops 294

vallierensis Colas, 1965, Carabus 156



vallierensis Jeanne, 1969, Carabus 139

vallierensis Coiffait, 1952, Pterostichus [Haptoderus] 490

vallierensis Coiffait, 1974, Trechus 330

valliger Hellén, 1929, Hydroporus 63

vallombrosus G. Rasetti & F. Rasetti, 1920, Duvalius [Trechus] 305

valonensis J. Müller, 1936, Molops 475

valonensis Apfelbeck, 1904, Nebria 92

valpellicis Casale, 1980, Doderotrechus 298



valstronae Focarile, 1949, Trechus 341

valyianus Bokor, 1922, Duvalius [Duvalites] 306



vandalitiae Reiche, 1862, Percus [Feronia] 479

vandeli Fourès, 1954, Aphoenops 295

vandeli Colas & Mateu, 1958, Calathus 528

vandeli Coiffait, 1959, Geotrechus 310

vandeli Coiffait, 1969, Hydraphaenops 311

vandeli Mateu & Colas, 1954, Tachys 275

vandeli Coiffait, 1952, Trechus 338

vanderberghi Perreau & Quéinnec, 1987, Geotrechus 310

vanemdeni Schauberger, 1932, Harpalus 383

vanensis P. Machard, 1985, Carabus 186

vannii Magrini & Sclano, 1998, Duvalius 305

vanvolxemi Putzeys, 1875, Carabus 137


vanvolxemi Putzeys, 1874, Nebria 93

vanvolxemi Putzeys, 1874, Pterostichus 504

vaporariorum Linné, 1758, Cymindis [Carabus] 418

vaquesensis Giordan, 1988, Duvalius 302

vardonensis J. Ochs, 1949, Speotrechus 320

vareillesi Giordan & Raffaldi, 1983, Duvalius 304

varendorffi Sainte‑Claire Deville, 1903, Trechus 343

variabile J. Müller, 1902, Bembidion 258

variabile Waltl, 1835, Bembidion 252

variabilis Ballion, 1878, Carabus 178

variabilis Andrewes, 1929, Dyschirius 229

variabilis Lucas, 1842, Nebria 92

variabilis Sirguey, 1929, Notiophilus 97

variabilis Clark, 1863, Peltodytes [Haliplus] 33

variabilis Ménétriés, 1832, Pterostichus [Feronia] 501

variabilis Balkenohl, 1999, Trilophus 222

varians Chaudoir, 1882, Brachyodes [Oodes] 444

varians Fischer von Waldheim, 1823, Carabus 194



varians LeConte, 1850, Hydroporus 65

varians Schaufuss, 1862, Platyderus 523

varians Uéno, 1981, Trechiama 324

varicolor Heer, 1838, Amara 568

varicolor Fabricius, 1803, Bembidion [Carabus] 246



varicornis LeConte, 1848, Harpalus 377

varicornis Newman, 1833, Nebria 92

variefucatus Font, 1997, Carabus 143

variegatum Boheman, 1848, Bembidion 258

variegatum Olivier, 1811, Omophron 208



variegatus Geoffroy, 1785, Chlaenius [Buprestis] 350

variegatus Fabricius, 1781, Graphipterus [Carabus] 359

variegatus Sturm, 1834, Haliplus 32



variegatus Geoffroy, 1785, Hydroporus [Dyticus] 64

variegatus Aubé, 1838, Hyphydrus 74

variegatus Stephens, 1828, Hyphydrus 75

variegatus Germar & Kaulfuss, 1816, Laccophilus [Dytiscus] 77

variegatus Aubé, 1838, Nebrioporus [Hydroporus] 67

variicornis A. Morawitz, 1863, Chlaenius 348

variicornis Chaudoir, 1865, Neocollyris [Collyris] 116

variipes Bates, 1883, Harpalus 374

variipes Chaudoir, 1850, Heptodonta [Aenictomorpha] 115

variipes Chaudoir, 1868, Pterostichus [Feronia] 506

variitarsis Chaudoir, 1860, Neocollyris [Collyris] 116



varinii Gautier des Cottes, 1866, Orthomus [Feronia] 477

variola Netolitzky, 1910, Bembidion 267



variolaris Ménétriés, 1849, Carabus 182

variolatum Mikhailov, 1983, Bembidion 267

variolatus O. G. Costa, 1839, Carabus 181

variolatus Dejean, 1828, Pterostichus [Feronia] 504

variolosa Fabricius, 1794, Cymindis [Carabus] 415

variolosus Fabricius, 1787, Carabus 154



variolosus Herbst, 1784, Laccophilus [Dytiscus] 77

variolosus Graëlls, 1858, Pterostichus [Corax] 489

varipes Stephens, 1828, Acilius 49

varipes Motschulsky, 1850, Asaphidion [Tachypus] 242

variseta Straneo, 1984, Pterostichus [Amolopsa] 494

varistriatus Kraatz, 1878, Carabus 173

varium Olivier, 1795, Bembidion [Carabus] 254

varius Aubé, 1838, Graptodytes [Hydroporus] 60

varius Bold, 1868, Haliplus [HN] 30

varius Nicolai, 1822, Haliplus 30



varius Fabricius, 1801, Hydaticus [Dytiscus] 53

varius Csiki, 1930, Pterostichus 492

variventris Schaufuss, 1862, Brachinus 214

varsavianorum Pawowski, 1974, Tachys 276

vartashensis Belousov, 1989, Duvalius 305

vartianorum Mandl, 1967, Cephalota [Cicindela] 103

vartianorum Mandl, 1989, Chlaenius 351

vartianorum Mandl, 1973, Cymindis [Petrovitziella] 417

vartianorum Jedlika, 1967, Hexagonia 407



varum Vauloger de Beaupré, 1898, Parazuphium [Zuphium] 573

varvasi Solier, 1835, Carabus 152

varvasi Laporte, 1834, Chlaenius 355

vasconicus Jeannel, 1913, Hydraphaenops [Aphaenops] 312

vasconicus Jeannel, 1937, Laemostenus [Ceuthosphodrus] 535

vasconicus Aubry, 1963, Pterostichus [Haptoderus] 490



vasconicus Jeannel, 1920, Trechus 331

vasconicus Uhagón, 1904, Zabrus 569



vasiceki A. A. Fleischer, 1916, Neotrechus [Anophthalmus] 315

vasilinini Lutshnik, 1916, Harpalus 380


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