Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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viridis Say, 1823, Harpalus 372

viridis Schilsky, 1888, Harpalus 372

viridis Tschitschérine, 1898, Myas [Trigonognatha] 476

viridis Jedlika, 1940, Platynus [Colpodes] 466

viridis Preller, 1862, Poecilus 482

viridis Jedlika, 1940, Pristosia [Laemostenopsis] 532



viridis Schilsky, 1888, Pterostichus [Calopterus] [HN (2)] 488

viridissimus Kraatz, 1876, Carabus 186

viriditinctus Schauberger, 1938, Stenolophus [Acupalpus] 405

viridula Mandl, 1982, Lophyra [Cicindela] 112

viridulum Kraatz, 1877, Calosoma 119

viridulus Kraatz, 1878, Carabus 152

viridulus Ragusa, 1904, Carabus 161

viridulus Jedlika, 1950, Chlaenius 352

viridulus Geoffroy, 1785, Harpalus [Buprestis] 372

viridulus Solsky, 1874, Harpalus 381

visai Dvoák, 1982, Laemostenus [Antisphodrus] 537

visitorensis J. Müller, 1936, Molops 472

vitalei J. Müller, 1929, Dyschirius 230

vitalisi Andrewes, 1920, Macrocheilus 407

vitalisi W. Horn, 1913, Therates 115

viti Pawowski, 1977, Trechus 343



vitiosa Heyden, 1885, Cicindela 105

vitiosum Gemminger & Harold, 1868, Bembidion 250

vitosensis Maan, 1933, Pterostichus 485

vitreus Paykull, 1798, Agabus [Dytiscus] 39

vitreus Dejean, 1828, Pterostichus [Feronia] 487

vittata Fischer von Waldheim, 1820, Cymindis 417

vittata Zoubkoff, 1833, Rhopalostyla [Cymindis] 430

vittatum Stephens, 1828, Bembidion [Tachys] 264

vittatus Graëlls, 1858, Acupalpus [Trechus] 399

vittatus Heyden, 1870, Acupalpus 397

vittatus Bates, 1892, Amblystomus 360

vittatus Gestro, 1875, Amblystomus 360

vittatus Bates, 1873, Badister 440

vittatus Fischer von Waldheim, 1823, Daptus 369

vittatus Gebler, 1833, Harpalus 384

vittatus Fabricius, 1775, Hydaticus [Dytiscus] 54

vittatus Sharp, 1884, Hygrotus [Coelambus] 71

vittatus Baudi di Selve, 1864, Microlestes 422

vittatus Motschulsky, 1859, Microlestes [Blechrus] 424



vittatus Sharp, 1882, Neohydrocoptus [Hydrocoptus] 34

vittatus Brullé, 1834, Polistichus 574

vittatus Motschulsky, 1850, Tachys 21, 277

vitticollis Boheman, 1848, Pseuduvarus [Hydroporus] 57



vittiger Germar, 1824, Daptus [Ditomus] 369

vittiger Gyllenhal, 1827, Ilybius [Dytiscus] 43

vittigera Dejean, 1825, Lophyra [Cicindela] 113

vittipennis J. R. Sahlberg, 1908, Microlestes 424



vittipennis Bedel, 1900, Tachyura [Tachys] 278

vittula Erichson, 1837, Hydroporus 65

vittula Fairmaire, 1859, Syntomus [Dromius] 432

vivalis Illiger, 1798, Synuchus [Carabus] 546



vivesi Jeanne, 1985, Amara [Celia] 555

vivesi Jeanne, 1985, Brachinus 212

vivesi Nègre, 1966, Calathus 526

vivesi Jeanne, 1971, Harpalus 381

vivesi Jeanne, 1996, Platyderus 523

vivesi Jeanne, 1976, Trechus 340

vividus Fabricius, 1801, Calathus [Carabus] 528



vividus Chaudoir, 1876, Poecilus 481

vixdentata Tanaka, 1959, Amara [Armatoleirides] 552



vixstriatus Bates, 1873, Tachys 276

vladimiri Obydov, 1998, Calosoma [Callisthenes] 119

vladivostokensis Lafer, 1979, Pterostichus 495

vladsimirskyi Dejean, 1830, Carabus 183



vlascanus Born, 1909, Carabus 167

vlasovi Kryzhanovskij, 1962, Amara 562



vlassinensis Matits, 1911, Molops 474

vlasuljensis Apfelbeck, 1894, Carabus 166

vlasuljensis J. Müller, 1931, Duvalius 300

vlasuljensis Ganglbauer, 1891, Molops 472



vocontianus Keith, 1995, Carabus 166

vodozi Sainte‑Claire Deville, 1906, Bembidion 246

vogelii Schaufuss, 1862, Cymindis 411

vogesiaca Bourgeois, 1898, Amara 547

vogesiacum Schuler, 1962, Bembidion [Peryphus] 259

vogti Beheim & Breuning, 1943, Carabus [Eucarabus] 134

vogtianus Beheim & Breuning, 1943, Carabus [Eucarabus] 159

voitestii Jeannel, 1930, Duvalius 301

vojnitsi Mandl, 1980, Carabus 133



volaki Kult, 1944, Ophonus [Harpalus] 391

volgense Becker, 1872, Bembidion 258

volgensis Chaudoir, 1850, Amara [Leirus] 559

volgensis Csiki, 1927, Carabus 172

volgensis Dejean, 1825, Cephalota [Cicindela] 102

volgensis Fischer von Waldheim, 1823, Scarites 232

volhynicus Lutshnik, 1927, Carabus 152

volkovi O. Berlov, 1997, Cychrus 19, 202

vollenhovenii Chaudoir, 1865, Neocollyris [Collyris] 117

volscus A. Franzini & G. Franzini, 1984, Duvalius 306



volujakianus Apfelbeck, 1894, Carabus 163

vomax Dejean, 1826, Carabus 182

voraginis Hrka & Mlejnek, 1995, Duvalius 307

voraginis Jeannel & J. Ochs, 1938, Duvalius 306

vorago Hrka & Mlejnek, 1995, Duvalius 307

vorax Andrewes, 1935, Bembidion 261



vorbringeri Csiki, 1930, Pterostichus 493

voriseki Jeanne, 1972, Leistus 82

vranense Apfelbeck, 1902, Bembidion 257

vranensis Breit, 1904, Duvalius [Trechus] 308

vranensis Apfelbeck, 1904, Molops 473

vranensis Schatzmayr, 1929, Poecilus [Pterostichus] 482

vseteckai Maan, 1936, Bembidion 253

vseteckai Maan, 1938, Nebria 18, 87

vseteckai Maan, 1944, Pterostichus [Haptotapinus] 496

vuillefroyi Gautier des Cottes, 1867, Calathus 526

vuillefroyi Chaudoir, 1866, Nebria 88

vuillefroyi Dieck, 1870, Platyderus 523

vulcanus Abeille de Perrin, 1904, Geotrechus [Trechus] 310



vulgaris Fischer von Waldheim, 1824, Broscus 236

vulgaris Tschitschérine, 1906, Trichotichnus [Amaroschesis] 395

vulgivaga Boheman, 1858, Clivina 220

vulneratum Dejean, 1831, Bembidion 263

vulneratus Klug, 1834, Cybister 51



vulneratus Putzeys, 1875, Miscelus 436

vulneripennis W. S. MacLeay, 1825, Gnathaphanus 362

vulpinus Andrewes, 1923, Andrewesius [Colpodes] 456

vulpinus K. Daniel, 1904, Harpalus [Ophonus] 382

waagenorum W. Horn, 1900, Therates 116

wadiensis Jedlika, 1964, Harpalus 384

wagae Fairmaire, 1882, Carabus 176



wagneri Tschitschérine, 1893, Bembidion 262

wagneri Küster, 1846, Carabus 181

wagneri J. Müller, 1922, Dyschirius 226

wagneri Schauberger, 1926, Harpalus 381

wagneri Tschitschérine, 1893, Pterostichus [Feronia] 506

wagneri Ganglbauer, 1906, Trechus 343



wahnschaffei Heinemann, 1911, Bembidion 252

waillyi Giordan & Raffaldi, 1982, Duvalius 306

wakamatsuensis Imura & Mizusawa, 1994, Carabus 177

wakoi Imura, 1995, Carabus 144

walkeri Branden, 1885, Nebrioporus [Deronectes] 67

walkeri C. O. Waterhouse, 1897, Protopaussus 211

walkeriana Sharp, 1913, Tachyura [Tachys] 280

wallichi Hope, 1831, Carabus 162

walterheinzi Hieke, 1988, Amara 558

walterheinzi Ledoux & Roux, 1990, Nebria 88



walteri Reitter, 1896, Carabus 179

walteri Reitter, 1889, Cymindis 415

walteri Reitter, 1883, Pterostichus [DA] 485

walteri Pawowski, 1978, Trechus 343

walteri Morvan, 1978, Xestopus [Wittmerosphodrus] 530


walteriana Ledoux & Roux, 1997, Nebria 93

waltersachi Mandl, 1984, Carabus 130

wanckowiczi Régimbart, 1883, Gyrinus 27

wangdanus Imura, Su & Osawa, 1997, Carabus 175

wanghaifengensis Deuve, 1988, Trechus [Parepaphius] 341

wangi Wewalka & Nilsson, 1994, Allopachria 73

wangi Xie & Yu, 1993, Aristochroa 471

wangi Mazzoldi, 1998, Orectochilus 29

wangorum Uéno & Ran, 1998, Sinaphaenops 320

wangxingensis Deuve & Li, 1998, Carabus 170

wankae Sokolá, 1911, Carabus 195

wansuiensis Habu, 1979, Trichotichnus 397

wanxianicus Imura, 1995, Carabus 142

wardi Andrewes, 1929, Haplogaster 232

warentzowi Tschitschérine, 1897, Poecilus [Feronia] 479

warnieri Reitter, 1907, Agonum 451

waroki Morvan, 1982, Aparupa 456

wasastjernae C. R. Sahlberg, 1824, Ilybius [Dytiscus] 43

wasastjernae J. R. Sahlberg, 1875, Pterostichus [Feronia] 508

washoeana Casey, 1924, Amara [Celia] 565

wassulandi Jedlika, 1957, Amara 560

wassulandi Jedlika, 1962, Platynus 464

watamukiensis N. Ito, 1996, Trichotichnus 397

watanabei Uéno, 1962, Epaphiopsis 309



watanabei Takizawa, 1933, Hydroporus 63

watanabei Tanaka, 1960, Pterostichus 502

watanabei Uéno, 1971, Trechiama 324

watanabeorum Uéno, 1975, Epaphiopsis 309



wauthieri David & Marchal, 1964, Ophonus 391

waziristanum Andrewes, 1932, Bembidion 257

weberi Ganglbauer, 1911, Anophthalmus [Trechus] 291

weberi Assmann, 1999, Galiciotyphlotes 448



weberi J. Müller, 1900, Haliplus 31

weberi Balkenohl, 1999, Trilophus 222

wehnckei Gerhardt, 1877, Haliplus 31

wehnckei Sharp, 1882, Laccophilus 77

weibaoensis Deuve, 1992, Carabus 188

weidneri Lassalle, 1990, Carabus [Procerus] 186

weigelii Panzer, 1801, Carabus 154

weigoldi Heller, 1923, Holosoma [Parahololius] 444

weingaertneri Winkler, 1912, Anophthalmus [Trechus] 291

weingaertneri Reitter, 1917, Nebria 86

weiperti Hieke, 1997, Amara 553

weiratheri Baliani, 1935, Amara 563

weiratheri Netolitzky, 1935, Asaphidion 243


weiratheri Netolitzky, 1930, Bembidion 257

weiratheri J. Müller, 1937, Deltomerus 283

weiratheri Scheibel, 1937, Duvalius 306

weiratheri J. Müller, 1931, Harpalus 372

weiratheri J. Müller, 1932, Laemostenus 537

weiratheri J. Müller, 1930, Molops 475

weiratheri Winkler, 1926, Neotrechus 316

weiratheri Winkler, 1933, Omphreus 446

weiratheri Maan, 1940, Platyderus 523

weiratheri J. Müller, 1932, Tapinopterus 520

weiratheri Jeannel, 1929, Trechus 343

weiratheri J. Müller, 1935, Winklerites [Scotodipnus] 240

weiratherianum Netolitzky, 1932, Bembidion 260

weiratherianus Noesske, 1928, Scotoplanetes 320

weisei Reitter, 1883, Carabus 198

weishanicola Deuve, 1999, Carabus 124

weissi Dostal, 1996, Elaphrus 207

welensii Dejean, 1828, Pterostichus [Feronia] 504

wellensii Drapiez, 1819, Pterostichus [Harpalus] 488



wellschmiedi Kirschenhofer, 1985, Pterostichus 497

wenatchee Hatch, 1950, Bembidion 262

wengshuiensis Deuve, 1994, Carabus 188

wenshuicus Font, 1998, Carabus 144

wenxianensis Deuve, 1992, Carabus 129

wenxianensis Deuve & Mourzine, 1997, Cychrus 205

wenxianicola Deuve, 1996, Carabus 189



weolseae Kwon & Lee, 1984, Carabus [Leptocarabus] 159

werchratskii Rizun & Yanitskiy, 1994, Duvalius 301



werneri Reitter, 1899, Carabus 152

werneri Reitter, 1906, Deltomerus 283

werneri Ganglbauer, 1915, Zabrus 571

westbengalensis Wiesner, 1996, Therates 116

westermanni Motschulsky, 1858, Selina 407

westwoodi Putzeys, 1866, Clivina 220

westwoodii Saunders, 1838, Platyrhopalus 211

wetterhallii Gyllenhal, 1813, Masoreus [Harpalus] 357

wewalkai Fery & Fresneda, 1988, Deronectes 59

wewalkai Fery & Nilsson, 1993, Ilybius [Agabus] 43



wewalkai Vazirani, 1975, Laccophilus 76

wewalkai Brancucci, 1982, Platambus 44



weyersi Régimbart, 1900, Laccophilus 77

weymarni J. Balfour‑Browne, 1947, Copelatus 48



weymarni J. Balfour‑Browne, 1947, Ilybius 42

whitei A. Deyrolle, 1852, Carabus 140

whitei Wollaston, 1854, Thalassophilus 346



whithillii Hope, 1838, Catascopus 435

wichmanni Jeannel, 1929, Duvalius 306

wiedemanni Ménétriés, 1836, Carabus 131

wiedemanni Chaudoir, 1850, Molops 475

wiedemanni Fischer von Waldheim, 1844, Nebria 88

wiedemanni Fairmaire, 1866, Tapinopterus [Molops] 518

wiedmanni Cristoforis & Jan, 1837, Carabus 186

wiesleri Schatzmayr, 1929, Pterostichus 504

wilkinsi Semenov, 1903, Carabus 123

wilkinsi Dokhtouroff, 1885, Cicindela 105

willbergi Reitter, 1891, Platynus 463



wimmeli Reitter, 1893, Anthracus 400

wimmeli Schulz, 1900, Carabus 136

wingelmuelleri Ganglbauer, 1904, Duvalius [Trechus] 306

winkleri Gschwendtner, 1923, Agabus [Gaurodytes] 41

winkleri Netolitzky, 1943, Bembidion 256

winkleri Mandl, 1934, Calomera [Cicindela] 101

winkleri Breuning, 1933, Carabus 182

winkleri Jeannel, 1924, Duvalius [Duvalites] 308

winkleri Schauberger, 1923, Harpalus [NP] 384

winkleri Havelka, 1952, Licinus 442

winkleri Breit, 1914, Molops 472



winkleri J. Müller, 1930, Molops [HN] 473

winkleri Bari, 1972, Nebria 93

winkleri Meggiolaro, 1959, Orotrechus 317

winkleri Mandl, 1936, Tapinopterus 520

winkleri Jeannel, 1927, Trechus 343

winkleri J. Müller, 1946, Zabrus 570

winklerianus Jeannel, 1926, Anophthalmus 293

winklerianus Jeannel, 1926, Duvalius 306

winklerianus J. Müller, 1936, Molops 472

winneguthi Apfelbeck, 1907, Duvalius [Anophthalmus] 301



winneguthi Apfelbeck, 1907, Duvalius [Anophthalmus] 301

winneguthi Scheibel, 1937, Neotrechus 315

winthemi Dejean, 1831, Anisodactylus 361

witkowskii Pawowski, 1978, Trechus 343

wittmeri Hieke, 1981, Amara 566

wittmeri Basilewsky, 1979, Bembidion [Chlorodium] 248

wittmeri Mandl, 1974, Carabus 162

wittmeri Mandl, 1975, Carabus 176

wittmeri Zamotajlov & Sciaky, 1999, Chinapenetretus 281



wittmeri Mandl, 1975, Cychropsis 202

wittmeri Wewalka, 1971, Deronectes 59

wittmeri Mateu, 1977, Dromius 421

wittmeri Brancucci, 1983, Laccophilus 77



wittmeri Morvan, 1978, Laemostenus [Pristonychus] 539

wittmeri Perrault, 1985, Leistus 81

wittmeri Morvan, 1980, Lesticus 471

wittmeri Zamotajlov, 1992, Parapenetretus 284

wittmeri Casale, 1980, Parathlibops 231

wittmeri Wewalka, 1975, Platambus 44



wittmeri Morvan, 1978, Pristosia [Laemostenopsis] 531

wittmeri Morvan, 1978, Pterostichus 510

wittmeri Uéno, 1977, Trechus 343

wittmeriana Deuve, 1983, Cychropsis [Cychrus] 19, 202

wittmerianus Deuve, Lassalle & Quéinnec, 1985, Calathus 528



wittmerianus Mandl, 1977, Carabus 176

wittmerianus Mandl, 1978, Chlaenius 349

wittmerorum Heinertz, 1978, Carabus 155

witzgalli Fery & Brancucci, 1997, Deronectes 59



wockei Born, 1898, Carabus 168

wohlberedti Emden & Schauberger, 1932, Harpalus 384

wohlberedti Winkler, 1933, Omphreus 446

wolffii Dejean, 1826, Carabus 167

wolfi Wrase, 1995, Ophonus 389

wolfianus Reitter, 1888, Carabus [Neoplectes] 132

wollastoni Sharp, 1882, Agabus 41



wollastoni Harald Lindberg, 1953, Bembidion 260

wollastoni Wrase & Jaeger, 1996, Bradycellus 400

wollastoni Cockerell, 1922, Eurygnathus 441

wollastoni Dawson, 1854, Harpalus 370

wollastoni Fauvel, 1905, Philorhizus [Dromius] 426

wollastoni Wollaston, 1854, Poecilus [Omaseus] 480

wollastoni Jeannel, 1922, Trechus 343

wollastonii Javet, 1852, Loricera 98

wolong Farka & Sciaky, 1998, Leistus 81

wolongensis Deuve, 1993, Carabus 124

woongbii Park & Kwon, 1996, Pterostichus 497

worleei Barthe, 1908, Cicindela 104

woronowi Lutshnik, 1916, Pterostichus [Platysma] 496

wraseanum Toledano, 1998, Bembidion 252

wrasei Hieke, 1988, Amara 566



wrasei Müller‑Motzfeld, 1986, Bembidion 256

wrasei Kirschenhofer, 1997, Chlaenius 354

wrasei Zamotajlov, 1999, Deltomerodes 282

wrasei Zamotajlov & Sciaky, 1996, Diplous 284

wrasei Farka & Sciaky, 1998, Leistus 81

wrasei Sciaky, 1987, Ophonus 391

wrasei Casale & Sciaky, 1999, Pseudotaphoxenus 542

wrasei Zaballos & Farinós, 1995, Typhlocharis 241



wraseiana Ledoux & Roux, 1996, Nebria 95

wroughtoni Wasmann, 1894, Paussus 211

wrzecionkoi Lassalle, 1999, Acalathus 529

wrzecionkoi Deuve, 1998, Amerizus 242

wrzecionkoi Deuve, 1996, Trechus 343

wuangxianus Kleinfeld, 1998, Carabus 129

wuduensis Deuve, 1997, Carabus 189

wuduensis Deuve, 1994, Cychrus 203

wuduensis Lassalle & Marcilhac, 1999, Reflexisphodrus 543

wudumontanus Imura, 1998, Carabus 150

wuesthoffi Schauberger, 1925, Abax 469

wufengensis Kleinfeld, 1998, Carabus 128



wui Gschwendtner, 1933, Hyphydrus 75

wui G. Ochs, 1932, Orectochilus 29



wui Gschwendtner, 1935, Peltodytes 33

wuki Matcha, 1922, Amara 549

wulanensis Zamotajlov, 1998, Diplous 284

wulffiusi A. Morawitz, 1862, Carabus 174

wuliangensis Cavazzuti, 1997, Carabus 126

wumingensis Deuve, 1992, Carabus 127

wushanling Kleinfeld, 1998, Carabus 129

wutaicola Deuve, 1988, Trechus [Parepaphius] 343

wutaishanense Kirschenhofer, 1984, Bembidion 272

wutaishanensis Shilenkov & Dostál, 1983, Nebria 89

wutaishanicus Kirschenhofer, 1986, Syntomus 433

wuyipeng Deuve, 1992, Agonotrechus 289

wuyipeng Deuve, 1992, Cychrus 206

xanthacra Chaudoir, 1850, Nebria 91

xanthacrum Chaudoir, 1850, Bembidion 251



xanthe Bates, 1892, Eucolliuris [Casnonia] 443

xanthicus Darge, 1992, Carabus 158

xanthina Casale, Giachino, Jali & Vailati, 1998, Reicheadella 222

xanthocerum Bates, 1883, Bembidion 270

xanthochiton Andrewes, 1922, Bembidion 257



xanthochrous Fuente, 1902, Stenolophus 406

xanthomerus L. Redtenbacher, 1867, Orthogonius 446

xanthomum Chaudoir, 1850, Bembidion 261

xanthomus Chaudoir, 1844, Badister 441

xanthomus Maindron, 1905, Syntomus [Metabletus] 433

xanthophanum Bates, 1888, Scalidion [Lebia] 430

xanthopleurus Chaudoir, 1856, Chlaenius 351

xanthopus Fischer von Waldheim, 1832, Cicindela 107

xanthopus Gemminger & Harold, 1868, Harpalus 384



xanthopus Stephens, 1835, Hydroporus 65

xanthopus Andrewes, 1936, Peronomerus 448



xanthopus Andrewes, 1934, Pristosia [Calathus] 532

xanthospilus Wiedemann, 1821, Callistoides [Carabus] 348



xanthotelum Bates, 1892, Bembidion 251

xanthotelus Bates, 1874, Apotomus 237

xantischius Gozis, 1911, Acilius 48

xatartii Dejean, 1828, Pterostichus [Feronia] 504



xaxarsi Born, 1917, Carabus 168

xaxarsi Zariquiey, 1919, Microtyphlus [Scotodipnus] 240

xaxarsi Schauberger, 1928, Ophonus [Harpalus] 391

xenechthum Gistel, 1857, Amara [Celia] 556

xestoides Nakane, 1962, Xestagonum [Colpodes] 469

xestum Andrewes, 1923, Bembidion 257

xestum Bates, 1883, Xestagonum [Anchomenus] 469

xiahensis Lassalle, 1999, Acalathus 529


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