Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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xiahensis Lassalle, 1993, Pseudotaphoxenus 543

xiangchengicus Deuve, 1993, Carabus 191

xiaoxiangensis Deuve, 1995, Carabus 127

xiaoxianglingensis Cavazzuti, 1996, Carabus 126

xiaoxinganensis Li & Liang, 1992, Nippononebria 95

xiei Deuve, 1992, Carabus 191

xiei Deuve, 1989, Cychrus 206

xiei Deuve, 1992, Trechus 343

xilinensis Zamotajlov & Wrase, 1997, Robustopenetretus [Chinapenetretus] 286

xingshanensis Imura, 1996, Carabus 129



xingziensis Kleinfeld, 1996, Carabus 156

xinjiangensis Huang, Hu & Sun, 1994, Harpalus 386

xinjiangensis Deuve, 1992, Trechus 343

xinlongicus Deuve, 1995, Carabus 190

xinyingensis Kleinfeld, 1998, Carabus 156

xiuyanensis Deuve & Li, 1998, Carabus 183

xiwuensis Deuve, 1996, Trechus 343

xueshanicola Deuve, 1992, Carabus 188

xui Deuve, 1996, Carabus 191

xunyangbanus Imura, 1998, Carabus 159

xupuensis Kleinfeld, 1998, Carabus 129

yaanensis Deuve & Tian, 1999, Carabus 125

yaconinus Bates, 1873, Carabus 178



yadongensis Huang, 1998, Harpalus 386

yagii Kitayama, Mori & Matsui, 1993, Hydrovatus 71

yajiangensis Deuve, 1994, Carabus 175

yajiangensis Deuve & Mourzine, 1998, Cychrus 205

yajiangensis Farka, 1995, Leistus 81

yak Deuve, 1991, Carabus 189

yakalamakus Lassalle, 1992, Carabus 193

yakuense Habu, 1974, Hikosanoagonum [Platynus] 458



yakushimana Nakane, 1963, Cicindela 108

yakushimanus Nakane & Ishida, 1961, Pterostichus 502

yakushimanus Nakane, 1955, Therates 115

yakutiae Nilsson & Larson, 1990, Agabus 41

yakutiae Nilsson, 1990, Hydroporus 65

yamaguchii Kasahara, 1991, Platynus [Colpodes] 466

yamajii Kasahara, 1995, Apatrobus 280

yamajii Kasahara, 1993, Pterostichus 513

yamaokai Ishikawa & Kubota, 1994, Carabus 178

yamato Nakane, 1953, Carabus [Apotomopterus] 178

yamato Miyatake, 1985, Tenomerga 25

yamatonis Habu, 1975, Agonum [Platynus] 454

yamauchii Takami & Ishikawa, 1997, Carabus 176

yamauchii Uéno, 1993, Oroblemus 316

yamauchii Morita, 1992, Pterostichus 485

yamauchii Uéno, 1982, Yamautidius 345

yamizonis Uéno, 1988, Kurasawatrechus 313

yanfoueri Morvan, 1996, Xestagonum [Colpodes] 469

yangmingensis Deuve, 1995, Carabus 156

yangminshanicus Font, 1997, Carabus 141

yangpachensis Hieke, 1997, Amara 564

yangxianensis Deuve, 1999, Carabus 129

yanjinganus Imura & Mizusawa, 1998, Carabus 150

yanmenensis Deuve, 1996, Carabus 188



yanoi Jedlika, 1951, Bembidion 244

yanoi Kult, 1951, Clivina 220

yanoi Nakane, 1963, Lymnastis 274

yanoi Kult, 1949, Reicheiodes [Dyschirius] 231

yanyuanicus Cavazzuti, 1996, Carabus 129

yao Imura, 1999, Carabus 128

yaophilus Deuve, 1990, Carabus 127

yaraligozi S. Battoni, 1982, Carabus 148

yasudai Uéno, 1972, Caecidium 272

yasudai Ishikawa, 1971, Carabus [Procrustes] 164

yasudai Uéno, 1973, Trechus 343

yasuii Habu, 1974, Negreum [Platynus] 460



yasujense Morvan, 1973, Bembidion [Nepha] 253

yasumatsui Habu, 1953, Perigona 448

yasumatsui Habu, 1955, Synuchus [Calathus] 546

yatsenkokhmelevskyi Iablokoff‑Khnzorian, 1960, Duvalius [Trechus] 306

yatsuana Nakane, 1960, Nippononebria 95

yatsuensis Straneo, 1955, Pterostichus 494

yatsuensis Morita, 1997, Trichotichnus 397

yayeyamensis Satô, 1971, Orectochilus 29

yedoensis Kanô, 1933, Cylindera [Cicindela] 111


yemenensis Britton, 1948, Perileptus 289

yemenita Basilewsky, 1968, Abacetus 346

yemenita Mateu, 1986, Merizomena [Glycia] 409

yemenita Mateu, 1986, Syntomus 433

yengensis Morvan, 1982, Trechus 343

yeongsiki Kwon & Lee, 1984, Carabus [Tomocarabus] 149

yessoensis Sharp, 1891, Rhantus 47

yezoana Nakane, 1957, Cicindela 106

yezoanum Nakane, 1961, Agonum [Europhilus] 453

yezoensis Bates, 1883, Carabus 136

yezoensis Habu, 1941, Diplous 284

yezoensis Bates, 1883, Dyschirius 227

yezoensis Habu, 1947, Licinus 442

yichunensis Deuve & Li, 1999, Carabus 144

yinae Uéno, 1996, Deuveotrechus 298

yinchuanensis Huang, 1993, Harpalus 386



yingkouensis Li, 1992, Hydroglyphus [Guignotus] 56

yingpanjienus Cavazzuti, 1999, Carabus 129



yodo Nakane, 1955, Callytron [Cicindela] 101

yokoae Deuve, 1988, Carabus 159

yokohamae Bates, 1883, Cillenus [Cillenum] 272

yokohamae Nakane & Straneo, 1979, Pterostichus 502



yokoyamai Nakane, 1963, Cymindis 417

yokoyamai Uéno, 1958, Trechiama 324

yonaguniensis Habu, 1977, Stenolophus 405

yonghongi Li, 1992, Carabus [Tomocarabus] 149

yonghongi Li, 1992, Carabus [Leptocarabus] 174

yonghongianus Deuve, 1992, Carabus 174

yongshan Kleinfeld, 1999, Carabus 125

yooni Deuve, 1990, Carabus 122

yoritomus Bates, 1873, Pterostichus 498

yoro Ashida, 1998, Trechiama 324

yoroensis Ishikawa & Kubota, 1994, Carabus 177

yoshiakii Uéno, 1978, Trechiama 324

yoshidai Uéno, 1969, Awatrechus 296

yoshidai Kasahara, 1985, Pterostichus 502

yoshidai Uéno, 1983, Trechiama 324

yoshihiroi Morita, 1997, Trichotichnus 397

yoshikawai Morita, 1996, Bembidion [Ocydromus] 266

yoshikawai Uéno, 1960, Kusumia 314

yoshikawai Ishida, 1959, Pterostichus 498

yoshikoae Uéno, 1970, Rakantrechus 319

yoshikoae Uéno, 1994, Trechiama 324



yoshimurai Kamiya, 1932, Hydroglyphus [Bidessus] 56

yoshiyukii Imura & Mizusawa, 1999, Carabus 177



yosiiana Uéno, 1960, Kusumia 314

yosiianus Uéno, 1999, Ishikawatrechus 312

yosiianus Uéno, 1954, Trechus 325

youngi Fery & Hosseinie, 1998, Deronectes 59

ypsilon Dejean, 1830, Clivina 220

ypsilon Reiche, 1863, Graptodytes [Hydroporus] 59

yuae Ledoux & Roux, 1995, Archastes 79

yuae Deuve, 1989, Carabus 127

yuae Deuve, 1989, Cychrus 204

yuae Deuve, 1992, Deuveotrechus [Kozlovites] 298

yuae Deuve, 1997, Sinozolus 346

yuanbaoensis Deuve, 1994, Carabus 127

yuanshanensis Kleinfeld, 1996, Carabus 127

yuasai Nakane, 1955, Callytron [Cicindela] 101

yubariensis Ishikawa, 1966, Carabus [Procrustes] 164

yubeicus Deuve & Tian, 1999, Carabus 124

yudanus Nakane, 1977, Carabus 149

yuellus Deuve, 1992, Carabus 127



yuianus Deuve, 1991, Carabus 141

yukihikoi Habu, 1962, Anchomenus [Agonum] 456

yukihikoi Habu, 1961, Trichotichnus 397

yukikoae Uéno, 1985, Trechiama 324

yukonum Fall, 1926, Bembidion 272

yulongshanensis Sciaky, 1997, Pterostichus 500

yulongshanus Jaeger, 1996, Bradycellus 402

yulongxuensis Deuve, 1990, Carabus 188

yulongxuicus Deuve, 1990, Cychrus 206

yunanensis Born, 1905, Carabus 128

yungeri Breuning & Ruspoli, 1970, Carabus 181

yunglaensis Deuve, 1997, Carabus 176

yunkaicus Deuve, 1992, Carabus 127

yunlingensis Deuve, 1991, Carabus 128

yunlingicus Deuve, 1994, Carabus 128

yunnana Baliani, 1934, Amara 548

yunnana Fairmaire, 1887, Heptodonta 115

yunnana Jedlika, 1933, Lebia 429

yunnana Bänninger, 1928, Nebria 91

yunnana Naviaux, 1999, Neocollyris 117

yunnana Kirschenhofer, 1994, Parena 434

yunnana Csiki, 1931, Pristosia [jun. HN?] 531

yunnana Jedlika, 1931, Pristosia [Laemostenopsis] [jun. HN?] 532

yunnanense Breuning, 1927, Calosoma 120

yunnanensis Jedlika, 1937, Pristosia 531

yunnanensis Jedlika, 1934, Pterostichus 501

yunnanicola Deuve, 1989, Carabus 128


yunnanicus Deuve, 1998, Carabus 127

yunnanicus Mateu, 1961, Microlestes 424

yunnanulus Mandl, 1992, Chlaenius 353

yunnanum Andrewes, 1923, Bembidion 256

yunnanus Uéno, 1999, Agonotrechus 289

yunnanus Csiki, 1931, Andrewesius [Colpodes] 456

yunnanus Jedlika, 1963, Brachinus 215

yunnanus Jedlika, 1931, Bradycellus [Tachycellus] 402



yunnanus Maindron, 1909, Callistomimus [Pristomachaerus] 348

yunnanus Born, 1910, Carabus [Coptolabrus] 142

yunnanus Fairmaire, 1886, Carabus 129

yunnanus Fairmaire, 1886, Chinapenetretus [Patrobus] 281

yunnanus Andrewes, 1920, Chlaenius 349

yunnanus Jedlika, 1941, Chydaeus 362

yunnanus N. Ito & Wrase, 2000, Coleolissus 369

yunnanus Fairmaire, 1887, Cychrus 206

yunnanus Jedlika, 1932, Diplous 284

yunnanus Schauberger, 1930, Harpalus 385

yunnanus Bänninger, 1925, Leistus 81

yunnanus Straneo, 1943, Myas [Trigonognatha] 476

yunnanus Casale & Sciaky, 1999, Pseudotaphoxenus 543



yunnanus Fairmaire, 1887, Pterostichus [Platysma] 487

yunnanus Jedlika, 1935, Stenolophus [Egadroma] 405

yunnanus Fairmaire, 1887, Trichotichnus [Amathitis] 395

yunwuensis Kleinfeld, 1996, Carabus 125

yupeiyui Casale, 1988, Pseudotaphoxenus 543

yushana Uéno, 1989, Epaphiopsis 309

yushanensis Habu, 1973, Apatrobus [Patrobus] 281

yushanensis N. Ito, 1985, Bradycellus 402

yushanensis Habu, 1979, Trichotichnus 397

yushuensis Deuve, 1991, Carabus 150

yvanii Dejean, 1828, Pterostichus [Feronia] 504

zaballosi Jeanne & Ruiz‑Tapiador, 1996, Poecilus 484

zaboli Jedlika, 1968, Lebia [DA] 429

zabriformis Schauberger, 1936, Trichotichnus 395

zabroides Dejean, 1828, Amara 566

zabroides Putzeys, 1873, Calathus 526

zabroides Dejean, 1829, Harpalus 384

zacharenkoi Gramma & Prisny, 1973, Haliplus 31

zadoiensis Lassalle, 1999, Acalathus 529

zaerensis Antoine, 1922, Acupalpus 399

zaerensis Antoine, 1928, Trechus 343

zagrosense Morvan, 1972, Bembidion [Emphanes] 249



zagrosense Morvan, 1973, Bembidion [Peryphus] 255

zagrosense Morvan, 1973, Bembidion [Peryphanes] 259

zagrosense Morvan, 1973, Bembidion [Testediolum] 269

zagrosense Morvan, 1973, Bembidion [Peryphus] 269

zagrosensis Morvan, 1973, Amara [Curtonotus] 561

zagrosensis Morvan, 1975, Platyderus 523

zahlbrucknerii Dejean, 1828, Pterostichus 503

zaianus Antoine, 1941, Carabus 152

zaianus Antoine, 1963, Trymosternus 410

zaikai Obydov, 1999, Carabus 170

zaimisi Jeannel, 1929, Duvalius 306



zaisani Jedlika, 1964, Amara 550

zaisanicum Müller‑Motzfeld & Kryshanovskij, 1983, Bembidion 246

zaitzevi Lutshnik, 1938, Bembidion 264

zaitzevi Nakane, 1990, Dytiscus 52



zaitzevi Jakobson, 1908, Hydroporus 60

zaitzewi Poppius, 1909, Agabus 37

zajtzewi Eichler, 1924, Xanthomelina [Apristus] 426



zakharschewskii Motschulsky, 1845, Carabus 137

zamotailovi O. Berlov, 1997, Cychrus 19, 202

zamotajlovi Belousov, 1998, Cimmerites 297

zamotajlovi Farka & Sciaky, 1998, Leistus 81

zamotajlovi Belousov, 1991, Pterostichus 501

zamotajlovi Belousov, 1990, Trechus 343

zanderi S. Battoni & Blumenthal, 1973, Carabus 193

zangezuricus Iablokoff‑Khnzorian, 1966, Trechus 335

zangherii Jeannel, 1927, Trechus 343



zanteanus Breuning, 1964, Carabus 187

zantunensis Cavazzuti & Ratti, 1998, Carabus 127

zanzibaricus Chaudoir, 1878, Dyschirius 227

zaonus Habu, 1954, Stomis 518



zaraensis Breuning & Ruspoli, 1970, Carabus 198

zarandicus P. Moravec, 1986, Trechus 343

zargalani Jedlika, 1968, Cymindis 416

zargoides Wollaston, 1863, Cymindis [Tarus] 418



zariquieyi Born, 1924, Cychrus 19, 203

zariquieyi Español, 1940, Duvalius 301

zariquieyi Bolívar y Pieltain, 1916, Microtyphlus [Scotodipnus] 240

zariquieyi Holdhaus, 1924, Reicheia 222

zarudniana Tschitschérine, 1903, Cephalota [Cicindela] 103



zarudnianum Semenov & Redikorzev, 1928, Calosoma 119

zarudnianus Semenov & Znojko, 1929, Odotoncarus [Carterus] 366

zarudnyi Tschitschérine, 1901, Abacetus 347

zarudnyi Semenov & Znojko, 1932, Carabus 147

zarudnyi Semenov, 1928, Leistus 84

zaslavskii Jeannel, 1962, Trechus 343

zavattarii Ghidini, 1934, Allegrettia 290


zawadzkii Kraatz, 1854, Carabus 174

zayula Andrewes, 1936, Nebria 93

zebiaensis Horvatovich, 1975, Nebria 86

zebianus Schauberger, 1927, Pterostichus 488

zebii Schaum, 1857, Tapinopterus [Feronia] 519

zechneticum Müller‑Motzfeld, 1989, Bembidion 259

zeelandica L. Redtenbacher, 1867, Diplocheila [Rembus] 439



zeelandicus L. Redtenbacher, 1867, Calathus 525

zelengorica Horvatovich, 1974, Nebria 85

zenbai Uéno, 1990, Kurasawatrechus 313

zengingoenueli Schweiger, 1965, Anillidius [Nannotrechus] 290

zepcensis Reitter, 1902, Carabus 163

zeravschanicus Glasunov, 1893, Orectochilus 29

zetterstedti C. G. Thomson, 1856, Agabus 37

zetterstedti Gyllenhal, 1827, Trachypachus [Blethisa] (In errata, Vol.2, p. 21: should be under Trachypachidae, not Broscinae) 33

zeytinlikensis Charet, 1996, Carabus 201

zezeae Csiki, 1931, Andrewesius [Colpodes] 456



zezeae Csiki, 1930, Tapinopterus [Pterostichus] 520

zhabyk Belousov & Kabak, 1994, Trechus 343

zhangi Deuve, 1992, Carabus 123

zhangi Deuve, 1989, Trechus 343

zhanglaensis Deuve, 1991, Carabus 189

zhanglaicola Deuve, 1996, Carabus 189

zhanglanus Zamotajlov & Sciaky, 1996, Parapenetretus 284

zhaojueanus Imura, 1999, Carabus 128

zhdankoi Kataev, 1990, Harpalus 384

zhdankoi Belousov & Kabak, 1992, Trechus 343

zheduoshanensis Cavazzuti, 1998, Carabus 190

zheduoshanus Uéno, 1998, Queinnectrechus 318

zhejiangensis Kirschenhofer, 1997, Pterostichus 516

zherichini Shilenkov, 1990, Carabus 174



zhicharevi Lutshnik, 1928, Amara 567

zhicharevi Silfverberg, 1977, Harpalus 383

zhicharevi Lutshnik, 1915, Jeannelius [Trechus] 312

zhicharevi Lutshnik, 1933, Poecilus [Platysma] 481

zhiltzovae Lafer, 1996, Sikhotealinia 25

zhongdianica Hieke, 1997, Amara 558

zhongdianus Farka, 1999, Leistus 81

zhongnanshan Kleinfeld & Schütze, 1997, Carabus 125

zhongtiaoshanus Imura & Yamaya, 1994, Carabus 144

zhoui Imura, Su & Osawa, 1998, Cychrus 206

zhubajie Imura, 1993, Carabus 191



zhugquensis Marcilhac, 1993, Pterostichus 500

ziegenhorni G. Hauser, 1921, Carabus [Coptolabrus] 141

ziegleri Géhin, 1876, Carabus 135

ziegleri Kraatz, 1883, Carabus 196

ziegleri Duftschmid, 1812, Pterostichus [Carabus] 508



ziegleri Panzer, 1809, Stenolophus [Carabus] 406

zierisi P. Moravec, 1987, Trechus 326

ziganensis Jeanne, 1976, Trechus 343



zigzag A. Costa, 1882, Ophonus 391

zilahiensis Csiki, 1906, Carabus 173

zimini Znojko, 1928, Dyschirius 228

zimmermanni Scholz, 1920, Agabus 38

zimmermanni Heer, 1837, Amara 555

zimmermanni Putzeys, 1875, Amara 548

zimmermanni Mandl, 1971, Calomera [Cicindela] 101

zimmermanni Gschwendtner, 1934, Copelatus 48



zimmermanni Gschwendtner, 1921, Haliplus 32

zimmermanni J. Müller, 1926, Hydroporus 65



zimmermanni Wallis, 1933, Rhantus 47

zinaidae Obydov, 1997, Carabus 174



zinaidae Lutshnik, 1915, Poecilus [Platysma] 482

zinovievi Belousov & Kabak, 1996, Trechus 343

zintshenkoi Belousov & Kabak, 1999, Trechus 343

zivkovi Knirsch, 1925, Duvalius [Trechus] 308

zlotini Mikhailov, 1996, Bembidion 258

znojkoi Semenov & Breuning, 1931, Carabus 145

znojkoi Kryzhanovskij, 1964, Pseudotaphoxenus [Taphoxenus] 543

znojkoiellus Deuve, 1988, Carabus 147

zogangensis Deuve, 1998, Deuveotrechus 298

zoiai Casale & Vigna Taglianti, 1999, Laemostenus 535

zoiai Sciaky, 1996, Pterostichus 488

zoiai Sciaky, 1989, Typhloreicheia 223

zoige Farka, 1999, Leistus 81

zoigeicus Deuve, 1990, Cychrus 206

zoigensis Deuve, 1990, Carabus 191

zoigensis Deuve, 1989, Trechus 343

zoldanus Depoli, 1939, Abax 470

zolotarevi Zamotajlov, 1988, Carabus 132



zolotarevianus Lutshnik, 1915, Poecilus [Platysma] 484

zolotarewi Reitter, 1910, Bembidion 256

zolotarewi Reitter, 1911, Pterostichus 505

zolotichini Zamotajlov, 1999, Deltomerodes 282

zolotikhini Belousov, 1990, Trechus 343

zonatus Panzer, 1796, Chlaenius [Carabus] 351

zonatus Hoppe, 1795, Graphoderus [Dytiscus] 49



zonatus Panzer, 1796, Graphoderus [Dytiscus] 49

zorkae Novotný & Voíšek, 1988, Carabus 200

zorkae Vysoký, 1983, Pterostichus 495

zorzini Vigna Taglianti, 1997, Guizhaphaenops 311

zoufali Maan, 1927, Abax 470

zoufali Reitter, 1913, Reicheadella 222

zouhari Jedlika, 1961, Elaphropus [Tachys] 273

zouhari Jedlika, 1961, Harpalus 384

zugmayeriae Reitter, 1888, Carabus [Neoplectes] 132

zulficari Schatzmayr & Koch, 1934, Porotachys [Tachys] [DA] 275

zupcensis Pavievi, 1990, Adriaphaenops [Aphaenopsis] 289

zvani Daffner, 1996, Anophthalmus 292

zvarici Vereschagina & Kabak, 1996, Pseudotaphoxenus 543

zvarici Bulirsch, 1990, Reicheiodes [Dyschirius] 230



zwickii Heer, 1837, Carabus 139

zwickii Fischer von Waldheim, 1820, Cephalota [Cicindela] 102

zygosensis Maan, 1939, Tapinopterus 520

zyrjanovskianus Shilenkov & O. Berlov, 1996, Carabus 170

zyzzovi O. Berlov & E. Berlov, 1996, Pterostichus 487



zyzzyus Lassalle, 1992, Carabus 201


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