Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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benahoaritus Machado, 1989, Trechus 327

benardi Breuning, 1931, Carabus 124

benardi Andrewes, 1924, Platynus [Anchomenus] 462



benedikti Bulirsch, 1995, Dyschirius 224

beneshi Kataev & Wrase, 1997, Harpalus 373

beneshi Sciaky, 1996, Pterostichus 488

benesi Deuve, 1996, Cychrus 203

benesi Morvan, 1998, Diacanthostylus 458

benesi Casale & Sciaky, 1999, Pseudotaphoxenus 540

benesi Deuve, 1993, Trechus 327

benesi Morvan, 1996, Xestagonum 467

bengalensis Aubé, 1838, Cybister 50

bengalensis Andrewes, 1929, Dyschirius 225

bengalensis Dejean, 1826, Scarites 233

benjamini Semenov, 1898, Carabus 189



benjamini Semenov, 1904, Cephalota [Cicindela] 102

bennigseni Sternberg, 1908, Chlaenius 355

benoiti Antoine, 1920, Agonum [Platynus] 451

benoiti Devecis, 1975, Carabus [Morphocarabus] 171

benoiti Jeannel, 1953, Stomis 517

bensai Gestro, 1892, Duvalius [Anophthalmus] 301

bensai Jeannel, 1927, Trechus 327

bensoni Hope, 1838, Macrocheilus 407

bensoni Westwood, 1845, Melanospilus 209

bentonis Bates, 1883, Negreum [Colpodes] 460

benxiensis Imura, 1991, Carabus 142



bepmalei Lapouge, 1904, Carabus 168

bepmalei Jeannel, 1921, Trechus 332



beraneki Reitter, 1887, Elaphrus 207

berardi Raynaud, 1967, Carabus [Chrysotribax] 140

berbera Antoine, 1951, Amara 567

berberus Géhin, 1885, Carabus 152

berberus Mateu, 1975, Laemostenus [Pristonychus] 535

berberus Antoine, 1925, Ophonus 389

berberus Antoine, 1963, Philorhizus 425



berdoa Bruneau de Miré & Legros, 1963, Hydrovatus 70

berezovskii Kurnakov, 1963, Parapenetretus [Penetretus] [DA] 284

berezowskii Shilenkov, 1984, Archastes 79

berezowskii Semenov, 1898, Carabus 142

berezowskii Tschitschérine, 1898, Pterostichus [Feronia] 501

bergamascus Jeannel, 1927, Trechus 336

bergesi Coiffait, 1958, Hydraphaenops [Aphoenops] 311

bergi Zaitzev, 1913, Agabus 35

bergmani Nilsson, 1995, Hydroporus 60

bergvalli Jeanne, 1996, Laemostenus 536

bericus Monguzzi, 1982, Speluncarius 516



beringensis Poppius, 1908, Pterostichus 492

beringi Netolitzky, 1939, Bembidion 247

bermamyticus Kurnakov, 1958, Pterostichus 501

bermejae Fernandez Cortes, 1995, Laemostenus 533

bernardi Bruneau de Miré, 1953, Coryza 221

bernardi J. Ochs, 1949, Duvalius 302

bernardi Jeannel, 1937, Pterostichus [Haptoderus] 490



bernensis Born, 1895, Carabus 180

bernhardinus Kraatz, 1878, Carabus 180

bernhardti Schuler, 1965, Amara [Percosia] 556

bernhaueri Ganglbauer, 1895, Anophthalmus 291

bernhauerorum Heinz, 1977, Carabus 188

bernii Vanni & Magrini, 1986, Duvalius 304

beroni Casale & V. B. Guéorguiev, 1994, Albanotrechus 290

beroni V. B. Guéorguiev, 1971, Duvalius 308



beroni Casale, 1988, Laemostenus 533

berrai F. Battoni, 1986, Lionychus 431

berrai F. Battoni, 1987, Orthomus [Pterostichus] 476

bertagnii Magrini, 1998, Duvalius 301



bertarinii Ganglbauer, 1891, Pterostichus [Pterostichus] 488

berthae Jeannel, 1910, Duvalius [Trechus] 301



berthouneaui Beuthin, 1904, Cylindera [Cicindela] 109

bertolinii Kraatz, 1878, Carabus [Orinocarabus] 180



bertolinii Seidlitz, 1887, Graphoderus 49

bertrandi Vazirani, 1969, Hyphoporus 73

bertrandi Legros, 1956, Stictotarsus [Deronectes] 69

beryllinus Preller, 1862, Poecilus 482

berytensis Reiche & Saulcy, 1855, Brachinus 213

berytensis Netolitzky, 1917, Ocys [Bembidion] 273

berytensis Reiche & Saulcy, 1855, Orthomus [Feronia] 477



besanconi Breuning & Ruspoli, 1970, Carabus 181

bescidensis Reitter, 1896, Carabus 179

bescidicus Reitter, 1903, Dromius 420

bescidicus Reitter, 1908, Trechus 336

beseanus Jeanne, 1970, Platyderus 522

beshkovi Coiffait, 1970, Duvalius 298

beskellum Morvan, 1996, Xestagonum 467



beskidensis Hlisnikowski, 1942, Duvaliopsis [Duvaliopsis] 298

besleticus Kurnakov, 1972, Carabus 132

besnicensis Pretner, 1949, Anophthalmus 291

bessarabicus Fischer von Waldheim, 1823, Carabus 197

besseri Fischer von Waldheim, 1820, Carabus 198

besseri Dejean, 1826, Cephalota [Cicindela] 102



besseri Fischer von Waldheim, 1828, Nebria 88

bessoni Cabidoche, 1962, Aphoenops 293

bessoni Cabidoche, 1967, Geotrechus 309

bessoni Cabidoche, 1964, Pterostichus 498



besucheti Schweiger, 1964, Carabus 131

besucheti Saha & Sengupta, 1979, Chlaenius 353

besucheti Brancucci, 1979, Geodessus 55

besucheti Baehr, 1998, Gunvorita 573

besucheti Straneo, 1988, Pterostichus [Haptotapinus] 496

besucheti Bucciarelli, 1958, Tapinopterus [Crisimus] 518

besucheti Pawowski, 1977, Trechus 327

besucheti Vigna Taglianti, 1972, Typhlocharis 241

besuchetianus Deuve, 1987, Trechus 327

betegara Andrewes, 1924, Bembidion 261

bettii Magrini, 1982, Duvalius 301

bettingeri Fagniez, 1927, Duvalius [Duvalites] 302

betuliae Csiki, 1940, Carabus 167


beusti Schaufuss, 1863, Trechus [Anophthalmus] 327

beuthini Schulz, 1899, Carabus [Tmesicarabus] 197

beuthini Fuente, 1912, Cicindela 105

beuthini Schulz, 1908, Cicindela 103

beuthini Gebien, 1899, Dicheirotrichus 402

beybienkoi Kryzhanovskij, 1974, Amara [Curtonotus] 559

beybienkoi Kryzhanovskij, 1979, Bembidion 244

beybienkoi Kryzhanovskij, 1973, Carabus 149

beydagensis Jeanne, 1996, Dyschirius 230

bhadarwahensis Deuve, 1982, Carabus 155

bhadarwahensis Lassalle, 1983, Laemostenus [Pristonychus] 537

bhadarwahensis Deuve, 1982, Trechus 327

bhaktiai Deuve, 1993, Carabus 162

bhamoensis Bates, 1892, Ophionea 444

bhamoensis Bates, 1889, Trigonotoma 520

bhotiya J. Schmidt, 1998, Aparupa 456

bhutanense Morvan, 1982, Xestagonum [Colpoides] 467

bhutanensis Morvan, 1980, Chaetobroscus 236

bhutanensis Mandl, 1978, Chlaenius 353

bhutanensis Mandl, 1992, Chlaenius 353

bhutanensis Mateu, 1977, Dromius 420

bhutanensis Morvan, 1982, Eobroscus 237

bhutanensis Wewalka, 1975, Platambus 44

bhutanensis Morvan, 1978, Pterostichus 509

bhutanensis Morvan, 1980, Pterostichus [Bothriopterus] [HN, repl. in vol. 2] 487

bhutanensis Morvan, 1979, Xestopus [Wittmerosphodrus] 530

bhutanicus Uéno, 1977, Trechus 327

bianchii Jeannel, 1928, Duvalius 301

biangulata Dupuis, 1912, Pentagonica 448

biarcuata Casey, 1924, Amara [Curtonotus] 561

biarticulata Motschulsky, 1844, Amara 548



biasiolii Gredler, 1876, Bembidion 267

bibanensis Lie, 1994, Carabus 196

biblis Britton, 1948, Tachyura [Tachys] 21, 279

bibula Antoine, 1944, Amara 549

bibulus Lompe, 1999, Trechus 327



bicarinata Drapiez, 1819, Yola [Dytiscus] 57

bicarinata Latreille, 1804, Yola [Dytiscus] 57

biceratops Cavazzuti, 1999, Cychrus 203

bicikensis Perrault, 1971, Duvalius 301



bicinctus Born, 1905, Carabus 165

bicinctus Hope, 1845, Somotrichus [Coptodera] 439

bickhardti Sainte‑Claire Deville, 1906, Amara 565

bicolor Solsky, 1874, Agatus [Glycia] 408

bicolor Dejean, 1828, Agonum [Anchomenus] 454



bicolor Geoffroy, 1785, Anchomenus [Buprestis] 455

bicolor Britton, 1948, Atrotus 439



bicolor Schilsky, 1888, Bembidion 249

bicolor Jaeger, 1998, Bradycellus 400

bicolor Fabricius, 1781, Calochroa [Cicindela] 99

bicolor Schulz, 1900, Cicindela 107

bicolor Zamotajlov & Sciaky, 1996, Diplous 283



bicolor Maindron, 1910, Dolichus 530

bicolor Fischer von Waldheim, 1829, Harpalus 386



bicolor J. Müller, 1933, Hydroporus 65

bicolor Reitter, 1890, Laemostenus [Antisphodrus] 533

bicolor Motschulsky, 1859, Parena 433

bicolor Aragona, 1830, Pterostichus [HN, repl. in vol. 2] 503



bicolor Dejean, 1831, Pterostichus [HN] 503

bicolor Fischer von Waldheim, 1829, Pterostichus [Argutor] [ND] 515



bicolor Chaudoir, 1843, Siagona 218

bicolor Rambur, 1837, Singilis 438



bicolor Boheman, 1858, Styphlomerus [Brachinus] 218

bicolor Tschitschérine, 1906, Trichotichnus [Amaroschesis] 394



bicoloratus Jedlika, 1937, Stereocerus [Pterostichus] 517

bicostatus Schaum, 1864, Deronectes [Hydroporus] 58

bicristata Sharp, 1882, Yola [Bidessus] 57

bicruciatus Germar, 1838, Stictonectes [Hydroporus] 69

bidentatus Gyllenhal, 1808, Oreodytes [Hyphydrus] 67

bidochei Jeanne, 1964, Pterostichus [Lianoe] 497

bidulus Deuve & Hodebert, 1991, Stevensius 320

biebersteini Ménétriés, 1832, Carabus 200

biellensis Born, 1923, Carabus 129

biellesinus Jeannel, 1937, Trechus 341

bielzi Birthler, 1886, Carabus 151

bielzi Seidlitz, 1867, Duvaliopsis [Anophthalmus] 298

bielzii Fuss, 1858, Pterostichus [Argutor] 506

bieneri A. Zimmermann, 1921, Graphoderus 49

biexcisus Straneo, 1955, Pterostichus 506

bifaria Mannerheim, 1853, Nebria 88

bifarius Kirby, 1837, Agabus [Colymbetes] 37



bifasciatum Schuler, 1959, Bembidion [Peryphus] 19, 256

bifasciatum Stephens, 1828, Bembidion [Notaphus] 254

bifasciatus Reitter, 1912, Callistus 347

bifasciatus Dejean, 1825, Calodromius [Dromius] 419



bifasciatus Régimbart, 1889, Canthydrus [add, syn of nitidulus Sharp] 34

bifasciatus Kollar, 1836, Ceratoderus [Pausus] 209

bifenestrata A. Morawitz, 1862, Lebia 430

bifidiphallus Nemoto, 1988, Pterostichus 497



bifoveatus W. J. MacLeay, 1871, Elaphropus [Bembidium] 273

bifoveicollis Reitter, 1896, Carabus 191

bifoveolatum C. R. Sahlberg, 1827, Agonum [Harpalus] 450

bifoveolatum Rambur, 1838, Bembidion 252

bifoveolatus Küster, 1846, Harpalus 371

bifoveolatus Park, Kwon & Lafer, 1996, Pterostichus 499



bifoveolatus A. Zimmermann, 1918, Stictonectes [Hydroporus] 69

bifrons Gyllenhal, 1810, Amara [Harpalus] [NP] 555

bifrons Andrewes, 1929, Dyschirius 229

bigemina Klug, 1834, Cylindera [Cicindela] 110



bigeminus C. G. Thomson, 1884, Notiophilus 96

bigerricus Jeannel, 1948, Geotrechus 309

bigerricus Jeannel, 1937, Pterostichus [Haptoderus] 489

bigerricus Jeannel, 1921, Trechus 326



bigerrio Lapouge, 1925, Carabus 139

bigoti Guignot, 1957, Bidessus 55

biguttatum Fabricius, 1779, Bembidion [Carabus] 263



biguttatum Motschulsky, 1850, Bembidion [Peryphus] 258

biguttatum Bates, 1892, Lachnoderma 438

biguttatus Olivier, 1795, Agabus [Dytiscus] 38

biguttatus Chaudoir, 1842, Brachinus 214

biguttatus Régimbart, 1895, Canthydrus 34

biguttatus Motschulsky, 1854, Chlaenius 348

biguttatus Clairville, 1806, Dromius [Lebia] 420

biguttatus Kirby, 1837, Laccophilus 76

biguttatus Chaudoir, 1874, Metaxys 472

biguttatus Fabricius, 1779, Notiophilus [Elaphrus] 96



bigutticeps Louwerens, 1953, Platynus 463

biguttulus C. G. Thomson, 1867, Agabus [Gaurodytes] 38



bihamatus Chaudoir, 1856, Chlaenius 354

bihamatus Chevrolat, 1861, Graptodytes [Hydroporus] 59

biharicus Maan, 1955, Abax 469

biharicus Maan, 1934, Calathus 527

biharicus Reitter, 1896, Carabus 196

biharicus Maan, 1933, Patrobus 285

biharicus Breit, 1912, Pterostichus 513

biharicus Meixner, 1912, Trechus 327

bihariensis Csiki, 1911, Duvalius [Anophthalmus] 300

biimpressus Fuss, 1858, Pterostichus [Argutor] [HN] 486

biimpressus Küster, 1853, Pterostichus [Omaseus] 510

biincisum Antoine, 1956, Bembidion [Princidium] 269

bilekensis Apfelbeck, 1904, Molops 473

bilimekii Sturm, 1847, Typhlotrechus [Anophthalmus] 344



bilineata Della Beffa, 1909, Cicindela 103

bilineatus Géhin, 1885, Carabus 140

bilineatus DeGeer, 1774, Graphoderus [Dytiscus] 49



bilineatus Sturm, 1835, Graptodytes [Hydroporus] 59

bilineatus Dejean, 1828, Percus [Feronia] 478

billbergi Mannerheim, 1827, Carabus 183

billieriensis Schaefer, 1969, Carabus [Autocarabus] 195

bilmenekus Lassalle, 1996, Carabus 147

biltoni Fery & Hosseinie, 1998, Deronectes 58



bilunata Heer, 1837, Cicindela 105

bilunatus Guérin‑Méneville, 1843, Chlaenius 354

bilunulatum Bielz, 1852, Bembidion 247

bimaculata Satô, 1972, Allopachria [Microdytes] 73

bimaculata L. Redtenbacher, 1844, Archicolliuris [Casnonia] 443

bimaculata Baudrimont, 1923, Eurynebria 79

bimaculata Bedel, 1907, Glycia 409



bimaculata Fischer von Waldheim, 1828, Lebia 429

bimaculatus Aubé, 1838, Cybister 50



bimaculatus Joukl, 1905, Diachromus 362

bimaculatus Dejean, 1825, Dromius 419

bimaculatus Paulino de Oliveira, 1876, Lionychus 431

bimaculatus W. S. MacLeay, 1825, Planetes 359

bimaculatus L. Dufour, 1852, Rhithrodytes [Hydroporus] 68

binaghii Burlini, 1939, Agonum 450

binaghii Baliani, 1943, Amara 567

binaghii Bucciarelli, 1978, Boldoriella 296

binaghii Vigna Taglianti, 1982, Duvalius 304

binaghii Casale, 1985, Typhloreicheia 223



binodulus Motschulsky, 1844, Chlaenius 352

binominis Reitter, 1886, Syntomus [Dromius] 432

binotata Dejean, 1825, Anchista [Plochionus] 408

binotata Fischer von Waldheim, 1820, Cymindis 417

binotatum Motschulsky, 1850, Bembidion 269

binotatum Stephens, 1828, Bembidion [Tachys] 264

binotatus Aubé, 1837, Agabus 38

binotatus Fabricius, 1787, Anisodactylus [Carabus] 360

binotatus Fischer von Waldheim, 1828, Badister 440

binotatus Zoubkoff, 1833, Brachinus 215

binotatus Klug, 1835, Cybister 51

binotatus Reitter, 1901, Microlestes [Blechrus] 422

binotatus Fischer von Waldheim, 1829, Pterostichus [Argutor] [ND] 515



binotatus Ragusa, 1886, Stenolophus 406

binotatus Putzeys, 1870, Trechus 330

binotulatus Motschulsky, 1865, Chlaenius 354

biocellatus O. F. Müller, 1776, Platambus [Dytiscus] 44

bioculata Andrewes, 1919, Dhanya 208

bioculata A. Morawitz, 1863, Lebidia 426

bioculata Putzeys, 1875, Tachyura [Tachys] 279


bioculatus Roubal, 1922, Callistus 347

bioculatus Chaudoir, 1856, Chlaenius 354

biokovensis Holdhaus, 1911, Duvalius [Trechus] 298

biokovensis J. Müller, 1916, Molops 472

biokovensis Csiki, 1908, Omphreus 445

bipartita Motschulsky, 1860, Amara 551

bipartitus Jacquelin du Val, 1857, Tachys 276

bipennifer Babington, 1835, Philorhizus [Dromius] 426

bipilifer Andrewes, 1927, Broscus 236



biplagiatum J. Müller, 1941, Zuphium 574

biplagiatus Boheman, 1858, Anthracus [Stenolophus] 399

biprolongata W. Horn, 1924, Cylindera [Cicindela] 110

bipunctata Kraatz, 1890, Cephalota [Cicindela] 101

bipunctata Kraatz, 1890, Cicindela 105

bipunctata Kraatz, 1890, Cylindera [Cicindela] 109

bipunctata Souverbie, 1855, Eurynebria [Nebria] 79

bipunctata Ragusa, 1887, Yola [Bidessus] 57

bipunctatum Duftschmid, 1812, Bembidion [Elaphrus] 257

bipunctatum Linné, 1761, Bembidion [Carabus] 269



bipunctatus Fabricius, 1787, Agabus [Dytiscus] 40

bipunctatus Gautier des Cottes, 1867, Calathus 527

bipunctatus Heer, 1838, Lionychus [Dromius] 432

bipunctatus Sturm, 1824, Platynus [Agonum] 463

bipustulatum L. Redtenbacher, 1849, Bembidion 264

bipustulatus Linné, 1767, Agabus [Dytiscus] 38



bipustulatus O. F. Müller, 1776, Agabus [Dyticus] 39

bipustulatus Quensel, 1806, Brachinus 214

bipustulatus Fabricius, 1775, Panagaeus [Carabus] 447

birmana Baliani, 1934, Amara 548



birmanica Bates, 1892, Galerita 359

birmanicus Régimbart, 1888, Hyphydrus 74



birmanicus Régimbart, 1891, Laccophilus 76

birmanicus Vazirani, 1970, Rhantus 46

birmanicus Bates, 1892, Trichotichnus 395



birmanus Andrewes, 1947, Harpalus 386

biroi Csiki, 1927, Carabus 192

biroi Csiki, 1905, Duvalius [Anophthalmus] 300

biroi Csiki, 1907, Pogonus 287

biroi Jedlika, 1952, Pseudotaphoxenus [Taphoxenus] 541

biroi Jedlika, 1952, Scybalicus 363

birsteini Ljovuschkin, 1963, Jeannelius 312

birthleri Hopffgarten, 1888, Agonum 451

birthleri Reitter, 1885, Carabus 173

birulai Poppius, 1913, Amara 558

birulai Poppius, 1910, Bembidion 265



bisarcensis Cabidoche, 1967, Hydraphaenops 311

bisbiguttatus J. R. Sahlberg, 1921, Hydroporus 63

bisbimaculata Chevrolat, 1860, Tachyura [Bembidium] 278



bisbipunctatus Puel, 1935, Harpalus 375

bischoffi Jedlika, 1946, Amara 557

bischoffi Chaudoir, 1848, Carabus 153

bischoffi Meschnigg, 1936, Duvalius 301



bischoffi Meschnigg, 1936, Neotrechus 315

bischoffi Meschnigg, 1934, Omphreus 445

bischoffi Meschnigg, 1933, Reicheadella [Reicheia] 222

bischoffi J. Müller, 1936, Tapinopterus 519

bischoffi J. Müller, 1936, Zabrus 571

bischoffianus Jedlika, 1936, Pterostichus 515

bisemilunatus Xie & Yu, 1991, Dischissus 447

biseriata Lutshnik, 1915, Nebria 88



biseriatus Chaudoir, 1846, Carabus 198

bisetiger Uéno, 1973, Awatrechus 296

bisetosus Deuve, 1995, Cychrus 203

bisetosus Belousov, 1998, Porocimmerites 318

bisetosus Straneo, 1938, Pterostichus [Feronia] 506

bisignatum Audinet‑Serville, 1821, Bembidion 264

bisignatum Ménétriés, 1832, Bembidion 268

bisignatus Aubé, 1838, Cybister 51

bisignifer A. Costa, 1882, Brachinus 214

bisioi Cavazzuti & Meli, 1999, Carabus 180



biskrensis Schulz, 1900, Cassolaia [Cicindela] 101

biskrensis Puel, 1938, Distichus [Scarites] 231

biskrensis Bedel, 1895, Dyschirius 225

bispicifer Andrewes, 1929, Dyschirius 230

bispinosus Deuve, 1989, Cychrus 203



bispinosus Gautier des Cottes, 1858, Molopidius [Pterostichus] 472

bispinula Hieke, 1997, Amara 563

bisquadripunctatus Klug, 1862, Distichus [Scarites] 231


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