Catalogue of Palaearctic Coleoptera Volume 1, 2003 I. Lobl & A. Smetana



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bissenica Bielz, 1887, Nebria 87



bistriata Putzeys, 1866, Amara [Curtonotus] 561

bistriatus Bergsträsser, 1778, Rhantus [Dytiscus] 46

bistriatus Duftschmid, 1812, Tachys [Elaphrus] 275

bistriatus Jeannel, 1928, Trechus 328



bistriga Reitter, 1894, Acupalpus 398

bistriolatus Duftschmid, 1805, Haliplus [Dytiscus] 33

bistroemi Wewalka, 1997, Microdytes 75

bisulcatum Chaudoir, 1844, Bembidion [Leia] 248

bisulcatus Curtis, 1840, Bidessus [Hygrotus] 55

bisulcatus Clark, 1863, Hyphydrus 74

bisulcatus Nicolai, 1822, Porotachys [Trechus] 274



biswasi Vazirani, 1965, Platambus 43

bithynicus Korge, 1971, Apotomus 237

bithynicus Schauberger, 1925, Morion 443



bitschnaui Born, 1926, Carabus 184

bitschnaui Reitter, 1908, Patrobus 285

bittneri Apfelbeck, 1904, Zabrus 572

bituberculatus Tschitschérine, 1899, Pterostichus [Feronia] 493

bivirgulata Fairmaire, 1889, Cylindera [Cicindela] 108

bivittata Fischer von Waldheim, 1823, Cymindis 417

bivittatus Andrewes, 1919, Amblystomus 360

bivittatus Laporte, 1835, Hydaticus 53

bivittis Motschulsky, 1859, Neohydrocoptus [Hydrocoptus] 34



bivittulus Reitter, 1893, Anthracus 399

bivulnerus Aubé, 1838, Cybister 51

bjelasicensis Winkler, 1933, Neotrechus 315



bjelasnicensis Apfelbeck, 1902, Carabus 173

bjelasnicensis Winkler, 1926, Duvalius 300

bjelasnicensis Apfelbeck, 1904, Leistus 83

blacki Edwards, 1913, Notiophilus 96

blairi Breuning, 1934, Carabus 177



blairi Omer‑Cooper, 1931, Gyrinus 27

blaisei Bedel, 1910, Mimocolliuris [Ophionea] 444

blakistoni Newman, 1858, Carabus 164

blancheti A. Gaudin, 1947, Hydraphaenops 311



blancki Peyerimhoff, 1927, Acilius 49

blanda Andrewes, 1924, Tachyura [Tachys] 278



blandicolle Netolitzky, 1910, Bembidion 264

blandulum Netolitzky, 1910, Bembidion 248

blandulus L. Miller, 1859, Pterostichus 489

blandus I. Frivaldszky von Frivald, 1865, Carabus 129



blandus Germar, 1836, Hygrotus [Hydroporus] 71

blaptoides Kollar, 1836, Carabus [Damaster] 148

blaptoides Creutzer, 1799, Zabrus [Carabus] 570

blatta A. I. Jakovlev, 1897, Agabus 39



bleusei Oberthür, 1885, Carabus 139

bleusei Beuthin, 1904, Cylindera [Cicindela] 109

blevennek Morvan, 1998, Xestagonum 467

bluehweissi A. Hoffmann, 1913, Neotrechus [Anophthalmus] 315

blumenthali Casale, 1982, Pseudotaphoxenus [Taphoxenus] 540

blumenthali Heinz, 1965, Pterostichus 489

blumenthalianus Mandl, 1963, Carabus 181

blumenthaliellus Deuve, 1988, Carabus 189

blumenthaliensis Heinz & Korge, 1967, Carabus 194

bocagei Paulino de Oliveira, 1876, Zuphium 574

bochkrizet Morvan, 1999, Xestagonum 467



bodeana Jedlika, 1953, Amara 548

bodemeyeri Reitter, 1907, Agonum 451

bodemeyeri Baliani, 1934, Amara 549

bodemeyeri K. Daniel & J. Daniel, 1902, Bembidion 255

bodemeyeri Apfelbeck, 1904, Brachinus 213

bodemeyeri H. Kolbe, 1929, Carabus [Macrothorax] 161

bodemeyeri Lapouge, 1925, Carabus 186

bodemeyeri Ganglbauer, 1900, Hydroporus 60

bodemeyeri Ganglbauer, 1900, Laemostenus 533

bodemeyeri Roubal, 1916, Notiophilus 96

bodemeyeri A. Fleischer, 1915, Tachyura [Tachys] 278

bodemeyeri Ganglbauer, 1900, Tapinopterus [Pterostichus] 519

bodemeyeri Reitter, 1913, Trechus 327

bodemeyeri Ganglbauer, 1915, Zabrus 570

bodenheimeri Netolitzky, 1935, Bembidion 251

bodenheimeri Bulirsch, 1997, Orientoreicheia 222

bodoanus Reitter, 1913, Duvalius [Trechus] 301



bodonensis Depoli, 1939, Abax 470

bodpaica Ledoux & Roux, 1998, Nebria 89

boeberi M. F. Adams, 1817, Carabus 137

boetica Ehlers, 1883, Typhlocharis 241

boeticus Rambur, 1837, Brachinus 19, 212

boeticus A. Deyrolle, 1852, Carabus 161



boeticus Jeannel, 1927, Trechus 330

boffai Casale, 1987, Typhloreicheia 223



bogatshevi Lutshnik, 1934, Amara 549

bogatshevi Zamotajlov, 1989, Carabus 160

bogatshevi Zamotajlov, 1988, Deltomerus 282

bogatshevi Kryzhanovskij, 1983, Pterostichus 498

bogatshevi Belousov, 1987, Trechus 327

bogdanovi Zamotajlov, 1991, Brachinus 215

bogdanovi Lutshnik, 1936, Calomera [Cicindela] 100

bogdanowi Ballion, 1878, Carabus 191

bogdoinus Breuning, 1932, Carabus 178

bogemanni Gyllenhal, 1813, Colymbetes [Dytiscus] 45

bogemannii Gyllenhal, 1813, Sericoda [Harpalus] 466

boghinorum Belousov & Kabak, 1992, Trechus 327

bogossicus Zamotajlov, 1992, Deltomerus 282

bohaci P. Moravec, 1987, Trechus 327

bohatschi Reitter, 1881, Carabus 196

bohdan Farka, 1999, Leistus 80

bohemani Ménétriés, 1832, Carabus 192

bohemani C. G. Thomson, 1856, Laccornis [Hydroporus] 75

bohemica Fassati, 1944, Amara 559

bohemicus Haury, 1881, Carabus 138


bohemicus Roubal, 1917, Ophonus 388

bohemorum Sciaky, 1994, Leistus 80

bohemorum Pawowski, 1973, Trechus 327

bohiniensis J. Müller, 1926, Abax 469

bohiniensis Ganglbauer, 1903, Anophthalmus [Trechus] 291

bohiniensis J. Müller, 1926, Oreonebria [Nebria] 96



boisduvali Gory, 1833, Plochionus 409

boisgiraudi L. Dufour, 1843, Pterostichus [Feronia] 498

boissyi Colas, 1951, Amara 563

boissyi Bonadona, 1955, Duvalius 306

boiteli Alluaud, 1930, Acinopus 368

boiteli Alluaud, 1932, Nebria 90

boiteli Alluaud, 1935, Omophron 208

boja Schrank, 1798, Cylindera [Cicindela] 109

bojani Daffner, 1998, Anophthalmus 291



bokharensis Csiki, 1927, Carabus 201

bokori Csiki, 1929, Amara 559



bokori Csiki, 1927, Carabus 172

bokori Csiki, 1910, Duvalius [Anophthalmus] 306

bokori Csiki, 1930, Poecilus [Pterostichus] 480

bokorianus Csiki, 1910, Duvalius [Trechus] 307

boldi Jedlika, 1966, Bembidion 268

boldi Jedlika, 1966, Cymindis 417

boldorii Jeannel, 1928, Allegrettia 290



boldorii Jeannel, 1930, Boldoriella [Speotrechus] 296

boldorii Porta, 1923, Cylindera [Cicindela] 109

boldorii Jeannel, 1926, Duvalius 301

boldorii Dodero, 1924, Laemostenus [Antisphodrus] 535

boldorii Schatzmayr, 1943, Zabrus 570

bolei Pretner, 1963, Duvalius [DA] 301

bolivari Andrewes, 1927, Amerizus [Bembidion] 241



bolivari Breuning, 1926, Carabus [Mesocarabus] 167

bolivari Zariquiey, 1920, Duvalius [Trechus] 301

bolivari Jeannel, 1913, Iberotrechus [Trechus] 312

bolivari Jeannel, 1937, Laemostenus [Ceuthosphodrus] 535

bolivari Jeanne, 1966, Nebria 90

bolivari Mateu, 1952, Trymosternus 410

bolognai Casale & Vigna Taglianti, 1984, Atranopsis [Tauroderus] 521



boluensis Jedlika, 1962, Harpalus 382

boluensis Schweiger, 1966, Speluncarius 516

bombarda Illiger, 1800, Aptinus [Carabus] 212

bombarda Duftschmid, 1812, Brachinus 214

bombycina Andrewes, 1925, Tachyura [Tachys] 277



bombycinus Leprieur, 1876, Deronectes [Hydroporus] 58

bonadonai Colas, 1948, Carabus 140

bonadonai Colas, 1948, Duvalius 304

bonadonius Giordan & Raffaldi, 1982, Duvalius 301

bonasus Jeannel, 1926, Aphoenops 294

bonelli Dejean, 1826, Nebria 85

bonelliana Lutshnik, 1921, Nebria 85

bonellii Dejean, 1826, Carabus 184

bonellii Putzeys, 1846, Dyschirius 228

bonellii M. F. Adams, 1817, Nebria [Carabus] 85

bonellii Guérin‑Méneville, 1833, Neocollyris [Collyris] 116

boneti Dupré, 1991, Trechus 336

bonfilsi Audinet‑Serville, 1821, Plochionus [Lebia] 409

bonina Nakane & Kurosawa, 1959, Cylindera [Cicindela] 110

boninensis Kasahara & Satô, 1990, Morion 443

boninensis Kasahara, 1991, Platynus [Colpodes] 464

boninensis Nakane, 1979, Tachyura [Tachys] 278

bonnairii Fairmaire, 1872, Hydroporus 62

bonneti Fourès, 1948, Aphoenops 294



bonneti Colas, 1936, Carabus 140

bonnetianus Colas, 1937, Carabus 140



bonomii Holdhaus, 1942, Amara 563

bonplandi Ménétriés, 1836, Carabus 158

bontempsi Lassalle, 1992, Carabus 160

bonvoisini Reiche & Saulcy, 1855, Poecilus [Feronia] 481

bonvouloiri Chaudoir, 1863, Carabus 163

bonvouloiri Sharp, 1882, Hydrovatus 70

bonvouloiri Pandellé, 1867, Trechus 327

bonvouloiri Vuillefroy, 1866, Typsiharpalus [Harpalus] 397



bonvouloirii Schaum, 1860, Cardiomera 457

bonzanoi Casale & Vigna Taglianti, 1990, Duvalius 302

boomensis Kabak, 1994, Carabus 178

boops J. R. Sahlberg, 1900, Anthracus [Acupalpus] 399

borchmanni Mandl, 1954, Lophyra [Cicindela] 112

borcka Jedlika, 1963, Pterostichus 504

bordei Peyerimhoff, 1909, Trechus 327

bordonii Magrini & Vanni, 1993, Anillus 238



borealensis Raynaud, 1973, Carabus 130

borealis Andrewes, 1942, Abacetus 347



borealis Sharp, 1882, Agabus 36

borealis Motschulsky, 1865, Agonum [Batenus] 452

borealis Chaudoir, 1843, Amara [Leirus] 558

borealis Motschulsky, 1844, Amara 556

borealis Motschulsky, 1850, Calathus 527

borealis Born, 1908, Carabus 160

borealis Géhin, 1885, Carabus 197

borealis Kwon & Lee, 1984, Carabus [Megodontus] 165

borealis Poppius, 1906, Carabus 135

borealis Motschulsky, 1860, Dytiscus 52

borealis Gerhardt, 1877, Haliplus 31

borealis Harris, 1869, Notiophilus 96



borealis Gyllenhal, 1826, Oreodytes [Hyphydrus] 67

borealis Motschulsky, 1844, Patrobus 22, 285

borealis Paykull, 1790, Pelophila [Carabus] 98



borealis Ménétriés, 1851, Pterostichus [Platysma] 493

borealis Motschulsky, 1844, Pterostichus [Steropus] 506

borealis Zetterstedt, 1828, Pterostichus [Harpalus] 487

borealis Andrewes, 1925, Tachyura [Tachys] [DA] 277

borealis Uéno, 1961, Trechiama 322

boreellus J. R. Sahlberg, 1871, Agabus [Gaurodytes] 38

boreellus Casey, 1914, Dicheirotrichus [Trichocellus] 403

boreellus J. R. Sahlberg, 1871, Pterostichus [Feronia] 507

borensis Lapouge, 1918, Carabus 191

borensis Roubal, 1918, Trechus 341

boreodzungarica Kabak, 1990, Amara 562



boreonense Avon, 1994, Bembidion [Peryphus] 251

boreus Csiki, 1927, Carabus 160

boreus Gordon, 1981, Hydroporus 65

boreus Csiki, 1930, Pterostichus 493

borgiae Dejean, 1826, Chlaenius 351

borisi Breit, 1936, Zabrus 570



boristhenicus Hochhuth, 1871, Nebrioporus [Hydroporus] 66

borni Csiki, 1906, Carabus 179

borni H. Kolbe, 1917, Carabus 138

borni Krausse, 1908, Carabus 161

borni Lapouge, 1925, Carabus 138

borni Meyer‑Darcis, 1902, Carabus [Coptolabrus] 144

bornianus G. Hauser, 1922, Carabus 128



borniellus Csiki, 1927, Carabus 138

borodini Heinz, 1996, Carabus 175

bortesii Casale & Vigna Taglianti, 1984, Duvalius 298

borulcini Jedlika, 1964, Pseudotaphoxenus [Taphoxenus] 540

borussicus Csiki, 1927, Carabus 135

borysthenicus Motschulsky, 1850, Dromius 421



borysthenicus Krynicki, 1832, Harpalus 371

boscensis A. Nicolas, 1923, Carabus [Chrysocarabus] 139

boschi Schauberger, 1934, Carterus 364

boschi Jeannel, 1929, Duvalius 302

boschi Winkler, 1949, Oreonebria 95



boschniaki Faldermann, 1836, Carabus 160

bosiljevici Drovenik & Pavievi, 1985, Carabus 163

bosnica Ganglbauer, 1889, Nebria 85



bosnicola J. Müller, 1937, Leistus 82

bosnicus Ganglbauer, 1891, Calathus 524

bosnicus Apfelbeck, 1890, Carabus 163

bosnicus Apfelbeck, 1912, Deltomerus 282



bosnicus Apfelbeck, 1909, Hydroporus 62

bosnicus Reitter, 1889, Laemostenus [Antisphodrus] 534



bosnicus Schweiger, 1970, Leistus 83

bosnicus Ganglbauer, 1889, Molops 472

bosnicus Apfelbeck, 1904, Pterostichus 487

bosnicus Ganglbauer, 1891, Trechus 327



bosogensis Obydov, 1996, Carabus 123

bosphoranus Fischer von Waldheim, 1823, Carabus 199



bosphoranus Motschulsky, 1844, Carabus 185

bosphoranus Reiche, 1862, Harpalus 381

boszdaghensis Breuning, 1966, Carabus 129

botchan Imura & Mizusawa, 1995, Carabus 176

botezati Netolitzky, 1922, Bembidion 266

bothoreli Morvan, 1999, Xestagonum 467



boudeti Beuthin, 1892, Carabus 141

boudikae Morvan, 1982, Trechus 328

boui Coiffait, 1969, Aphoenops 294

bouiganensis Fourès, 1954, Aphoenops 295

bouilloni Coiffait, 1955, Aphoenops 294

bouilloni Breuning & Ruspoli, 1976, Carabus 194

bouilloni Coiffait, 1958, Geotrechus 310

bouisseti Barthe, 1936, Carabus 139

boulbeni Morvan, 1998, Andrewesius 456

boulbeni Ledoux & Roux, 1998, Archastes 79

boulbeni Deuve, 1997, Cychrus 203

boulbeni Morvan, 1998, Diacanthostylus 458

boulbeni Deuve, 1998, Trechus 328

boulbenianus Deuve, 1996, Carabus 124

bourdeaui Coiffait, 1976, Aphoenops 294

bourderyi Maindron, 1906, Nebria 90

bourgoini Deuve, 1998, Carabus 187

bourgoini Jeannel, 1946, Hydraphaenops [Aphoenops] 311

bourgoini Jeannel, 1946, Hydraphaenops [Aphaenops] 311

boursini Fagniez, 1922, Duvalius [Duvalites] 305

bousquetellus Deuve, 1998, Carabus 124

bousqueti Deuve, 1993, Carabus 142

bousqueti Deuve, 1991, Cychropsis 201

bousqueti Ledoux, 1993, Nebria 88

bousqueti Mateu, 1990, Physodera 438

bousqueti N. Ito, 1997, Stenolophus 405

bousqueti Deuve, 1988, Trechus 328



bousqueti N. Ito, 1998, Trichotichnus 395

bousquetianus Deuve, 1994, Carabus 127

bouvieri Tschitschérine, 1897, Trichotichnus 395

bovelinii Heer, 1837, Cychrus 202

bowanus Sciaky, 1997, Pterostichus 500

bowringi Chaudoir, 1863, Carabus 165

bowringii Clark, 1864, Hydaticus 53

bowringii Westwood, 1850, Paussus 210

boyeri Blanchard, 1842, Calomera [Cicindela] 99

boysi Chaudoir, 1855, Scarites 233

boysi Arrow, 1901, Yamatosa [Rhysodes] 78

boysii Tatum, 1851, Carabus 155

boysii Chaudoir, 1850, Lebia 428



boysii Chaudoir, 1860, Neocollyris [Collyris] 117

boysii Westwood, 1845, Paussus 210

boysii Chaudoir, 1850, Platytarus [Cymindis] 410

bozoi Mlejnek & J. Moravec, 1995, Anophthalmus 291



brabeus Schaufuss, 1862, Carabus 167

braccata Bates, 1891, Dioryche [Platymetopus] 369

braccata Andrewes, 1934, Pristosia [Calathus] 530

bracculatum Bates, 1889, Bembidion 260



brachycephala W. Horn, 1893, Neocollyris [Collyris] 116

brachyderum Chaudoir, 1850, Agonum [Anchomenus] 451

brachymorphus Chaudoir, 1859, Pterostichus [Feronia] 488

brachypedilus A. Morawitz, 1886, Carabus 182

brachypteron Fassati, 1945, Bembidion 263

brachypterus Tschitschérine, 1898, Harpalus 373

brachypus Steven, 1809, Microderes [Carabus] 387

brachys Andrewes, 1925, Polyderis [Tachys] 274

brachythorax Lindroth, 1963, Bembidion 265

bracteonoides Reitter, 1908, Bembidion 266

bradycelliformis Reitter, 1900, Dicheirotrichus 403

bradycelliformis Csiki, 1906, Trechus 328



bradycelloides Reitter, 1900, Harpalus 380

bradycelloides Apfelbeck, 1904, Trechus 328

bradycelloides Reitter, 1903, Trechus 331

bradycephalus Jeannel, 1928, Duvalius 307

bradyta Hieke, 1988, Amara 563

bradytoides Reitter, 1889, Amara [Harpalodema] 562



bramardi Guignot, 1942, Rhantus 47

bran Morvan, 1997, Lepcha 459



branchiatus Babington, 1832, Agabus [Colymbetes] 38

brancoi Rocchi, 1981, Hydroporus 60

brancuccii Zamotajlov, 1992, Apatrobus 281

brancuccii Fery & Hosseinie, 1998, Deronectes 58



brancuccii Fery, 1987, Hydroporus 60

brancuccii Casale, 1982, Laemostenus [Pristonychus] 537

brancuccii Farka, 1995, Leistus 80

brancuccii Sciaky, 1995, Myas [Trigonognatha] 475

brancuccii Naviaux, 1992, Neocollyris 116

brancuccii Mateu, 1984, Paradromius 425

brancuccii Deuve, Lassalle & Quéinnec, 1985, Pristosia 530

brancuccii Straneo, 1982, Pterostichus 510

brancuccii J. Schmidt, 1997, Xestagonum 467

brandisi Ganglbauer, 1897 (as 1896), Duvalius [Trechus] 302



brandisi Reitter, 1908, Poecilus 482

brandmayri Sciaky, 1991, Philorhizus 425

brandti Harold, 1880, Agabus 39

brandti Faldermann, 1835, Carabus 137

branickii Taczanowski, 1888, Carabus 144

brannani Schaufuss, 1871, Carabus 161

brannanii Schaufuss, 1869, Deronectes [Hydroporus] 58



braudoi Poppius, 1906, Pterostichus 493

brauni Heinz, 1993, Carabus 155

bravoorum Mateu, 1957, Philorhizus 425

brdensis Bernau, 1910, Carabus 196

breiti Baliani, 1940, Amara 554

breiti Netolitzky, 1918, Bembidion 255



breiti Le Moult, 1913, Carabus 140

breiti Ganglbauer, 1900, Duvalius [Trechus] 302

breiti J. Müller, 1922, Dyschirius 226

breiti Mandl, 1942, Pheggomisetes 318

breitianus Knirsch, 1912, Duvalius [Trechus] 298

breizhinensi Morvan, 1976, Trechus 329

brembana Binaghi, 1937, Boldoriella [Speotrechus] 296

brembanus Focarile, 1949, Trechus 328

bremii Stierlin, 1881, Carabus 137

bremii Germar, 1831, Oreonebria [Nebria] 95

bremondi Guignot, 1934, Graptodytes 59

brendelli Morvan, 1999, Aparupa 456

brendelli Morvan, 1992, Deltocolpodes 457

brescianus Jeannel, 1926, Duvalius 306

bresi Verdier, Quézel & Rioux, 1951, Syntomus 432

bresk Morvan, 1998, Bruskmoal 457



breuili Jeannel, 1926, Geotrechus 310

breuili Jeannel, 1921, Platyderus 522

breuili Jeannel, 1919, Pterostichus [Troglorites] 514

breuili Jeannel, 1926, Thalassophilus 346

breuili Jeannel, 1913, Trechus 328

breuilianus Jeannel, 1916, Paraphaenops [Trechus] 318



breuningi Mandl, 1954, Calosoma 119

breuningi Csiki, 1927, Carabus 167

breuningi Jedlika, 1931, Chlaenius 349

breuningi Schweiger, 1964, Cychrus 202

breuningi Nègre, 1959, Speluncarius 516

breuningi Morvan, 1972, Trechus 328

breuningianus Le Moult, 1930, Carabus 124

breuningiellus Deuve, 1988, Carabus 197


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