Bacteria, Viruses and Protistans
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Bacteria, Viruses and Protistans (chap 18)
Bacteria – Most ancient forms of life (Table 18.1)
Great metabolic diversity (Table 18.2)
Photoautotrophic – synthesize own organic compounds <> using sunlight and carbon dioxide
Photoheterotrophic – use sunlight, but carbon comes from organic compounds
Chemoautotrophic – produce organic compounds using carbon dioxide and energy in inorganic substances
Chemoheterotrophic – cannot produce own organic compounds
Parasitic – draw nutrition from living hosts <>
Saprobic – obtain nutrition from products, wastes, or remains of other organisms
Bacterial sizes and shapes
Usually 1-10 micrometers
Three basic shapes
Coccus – spherical
Bacillus – rod-shaped
Spirillum – spiral-shaped
Structural features
Prokaryotic – no nucleus [or other membrane-bound organelles]
Metabolism occurs in cytoplasm or at plasma membrane
Proteins assembled on floating ribosomes
Nearly all possess cell wall
Gram-positive and gram-negative
Has jellylike capsule for attachment and to deter antibiotic activity
May have filamentous structures attached to cell wall
Flagellum – for movement
Pili – for attachment [incl. conjugation]
Bacterial reproduction
Binary fission – single “chromosome” (Fig. 18.5)
Plasmids carry only a few genes – replicated independently of main “chromosome”
Bacterial classification
Traditionally characterized by staining reactions, cell shape, metabolic patterns, and mode of nutrition
True classification based on evolutionary relationships becoming possible due to biochemistry studies
Major groups of bacteria
Archaebacteria
First living cells – exist in unusual habitats
Methanogens (“methane-makers”)
Inhabitat swamps, mud, sewage, and animal guts
Make ATP anaerobically by converting carbon dioxide and hydrogen to methane
Halophiles (“salt-lovers”)
Tolerate high salt environments [brackish ponds, salt lakes, volcanic vents, etc.]
Most are heterotrophic aerobes
Extreme thermophiles (“heat-lovers”)
Live in hot springs and volcanic vents
Use hydrogen sulfide for ATP formation
Eubacteria
Photoautotrophic eubacteria
Cyanobacteria [blue-green algae] are photosynthetic <>
Green and purple bacteria use hydrogen sulfide and hydrogen gas for photosynthesis
Chemoautotrophic eubacteria
Most important are nitrifying bacteria – nitrogen cycle
Use ammonia in generating ATP
Chemoheterotrophic eubacteria
Some are major decomposers in soil
Actinomycetes produce antibiotics, Lactobacillus used in dairy product conversions, etc.
Some E. coli causes serious diarrhea, Clostridium botulinum – botulism, Borrelia burgdorferi – lyme disease
Viruses
Defining characteristics
Noncellular infectious agent
Consists of nucleic acid core (DNA or RNA) surrounded by protein coat
“Reproduces” by causing host cell to make more viral particles
Vertebrate immune system can detect and fight viruses, but viruses continually mutate
Examples of viruses (Table 18.3)
Bacteriophages – infect bacterial cells
Animal [human] viruses – influenza, chickenpox, colds, HIV (AIDS)
Plant viruses – usually rely on insects to provide penetration of host cells
Infectious agents smaller than viruses
Prions – infectious protein particles (“Mad Cow” disease) that destroys nervous system
Viroids – naked pieces of RNA (no protein coat) that cause plant diseases
Viral multiplication cycles (Figs. 18.12 & 18.13)
Steps of viral replication
Virus “recognizes” and attaches to host cell
Nucleic acid core enters host cell
Viral genes direct host cell to replicate new viral components
Host cell ruptures releasing new viral particles
Replication can proceed by two pathways
Lytic pathway [see above] (Fig. 18.12)
Lysogenic pathway – latent period in host (Fig. 18.13)
Protistans – simplest eukaryotic organisms, unicellular and multicellular
Predatory and parasitic molds
Chytrids and water molds
Chytrids – decomposers or parasites in muddy or aquatic habitats
Water molds – attack weakened fish or land plants (e.g., Irish potato famine)
Slime molds (Fig. 18.15)
Heterotrophic, free-living, amoebalike protistans
Feeds by engulfing food particles; reproduces by spores
Two groups – cellular slime molds and plasmodial slime molds
Animal-like protistans – protozoans
Defining characteristics
All are predators or parasites
Mostly asexual reproduction by fission or budding
A few cause diseases in humans <>
Amoeboid protozoans (Fig. 18.17)
Pseudopodia – extensions of cell body
Amoeba proteus – lab animal; Entamoeba – causes dysentery
Foraminiferans – shelled forms; radiolarians – shells of silica; heliozoans – needle-like pseudopods
Ciliated protozoans
Numerous cilia – beat in synchrony
Paramecium – sexual reproduction involves conjugation (Fig. 18.18)
Flagellated protozoans (Fig. 18.19)
One or more whip-like entensions
Tichomonas vaginalis – spread by sexual contact; Giardia lamblia – mild diarrhea to death; Trypanosoma – African sleeping sickness
Sporozoans (Fig. 18.20)
Parasitic – sporozoite stage usually transmitted by insects, and encysted stage
Plasmodium – malaria (transmitted by mosquitoes); toxoplasmosis – can cause birth defects (transmitted from cats to humans through infected feces)
(Mostly) Single-celled photosynthetic protistans
Euglenoids
Mostly photosynthetic autotrophs, some are heterotrophic
Euglena – flexible pellicle, flagellum, eyespot
Chrysophytes
Golden-brown algae
Diatoms – shells of silica
Bottom of marine food chain
Commercially valuable as abrasives and filtering materials
Dinoflagellates
Two flagella located in grooves in cell wall
Some cause “red tides” (producing neutotoxins)
(Mostly) Multicelled photosynthetic protistans
Red algae (Fig. 18.23a)
Pigments trap sunlight in deep marine waters
Complex asexual and sexual life cycles
Some aid in reef building; others yield agar
Brown algae (Fig. 18.23b)
Includes kelps of intertidal zones
Plantlike – with leaflike blades on a stemlike stipe attached to a rootlike holdfast; some have floats
Produce algin – used as a thickening or suspension <> agent
Green algae (Fig. 18.24)
Grow nearly everywhere
Same pigments as land plants; store carbohydrates as starch
Some are symbionts with fungi <> and other organisms, others are colonial (Volvox ), many live singly (Chlamydomonas )
Plants and Fungi (chap 19)
Evolutionary trends among plants
Overview of the Plant Kingdom
Mostly multicelled photosynthetic <> autotrophs
Most have vascular tissues for transport of water and nutrients; and roots, stems, and leaves
Nonvascular plants [e.g., bryophytes] lack true roots or leaves
Evolution of roots, stems, and leaves
Roots – absorption of water and minerals
Shoots <> – exploiting sunlight and absorbing carbon dioxide
Lignin – hard substance in cell wall allowing extensive growth <>
Cuticle – covering stems and leaves to minimize water loss; evaporation <> controlled by stomata
From haploid to diploid dominance (Fig. 19.2)
Simple aquatic plants <> dominated by haploid phase
Complex land plants dominated by diploid sporophyte
Parts of sporophyte undergo meiosis to produce haploid spores
Spore develops into gametophyte, which produces gametes <>
Evolution of pollen and seeds
Spores of some algae and simple vascular plants are all alike [homosporous]
In gymnosperms and angiosperms <>, spores are of two types [heterosporous]
Pollen grain = male gametophyte
Female gametophyte remains within plant, producing seeds
Bryophytes – mosses, liverworts, and hornworts [mosses are the most common] (Fig. 19.4)
Lack xylem and phloem <>; rhizoids attach gametophyte to soil
Gametophyte
Developments from spore – haploid, leafy, independent plant
Contains archegonium <> and antheridium <>
Sporophyte
Developments from zygote [fertilized egg] – diploid and dependent on gametophyte
Composed of foot, stalk, and capsule [sporangium] that contains spores
Existing seedless vascular plants – sporophyte is dominant; xylem and phloem are present
Whisk ferns
“Twiggy” looking plant – no roots or leaves [photosynthesis occurs in stem tissue]
Underground rhizome; sporangia on branches
Lycophytes
Once tree-sized, but now small club mosses on forest floor
True roots, stems, and small leaves – cone-like strobili contain sporangia
Horsetails or scouring-rushes
Once treelike, now common “bamboo-like” roadside plants around here
Roots, rhizomes <>, aerial stems <>, but no true leaves [stem is photosynthetic]
Ferns
Roots, rhizomes, and big leaves [fronds – young ones called fiddleheads]
Sporangia are clustered into sori on the underneath of fronds
Gymnosperms – plants with “naked” seeds
Life cycle (Fig. 27.10)
Microspore (in “male” cone) develops into pollen grain – wind pollination
Megaspore develops into ovule [immature seed] – exposed or in “female” cone
Conifers
Cone-bearing trees with needlelike or scalelike leaves [mostly evergreen]
Male pollen cones and female seed cones
Fertilization results in a zygote that develops into an embryo within the seed
Lesser known gymnosperms
Cycads
Small palmlike trees – 100 species of the tropics and subtropics
Large pollen and seeds cones on separate plants
Ginkgos [or Maidenhair tree]
Only one species survived the Mesozoic – living fossil
Leaf fan-shaped, seed exposed
Gnetophytes [e.g., Mormon tea]
Found in tropical and desert areas
Angiosperms – flowering plants (Figs. 19.11 and 19.12)
Produce flowers with special tissues to protect ovules and seeds
Most species coevolved with pollinators
Dominated the land for 100 million years
Monocots and dicots
Monocots – grasses [inclu. cereal grains], lilies, etc. <>
Dicots – trees [except conifers], shrubs, and herbaceous plants <>
Life cycle
The diploid sporophyte has roots and shoots, and retains and nourishes the gametophyte
Flowers attract animal <> pollinators
Sepals and petals
Stamen – anther and filament
Pistil – stigma, style, and ovary [containing ovules]
Embryos nourished by endosperm within seeds, which are packaged inside fruits
Fungi
“Characteristics”
Three major groups [zygomycetes <>, sac fungi, and club fungi] and one “catch-all” category [imperfect fungi]
Nutritional modes
Heterotrophs that utilize organic matter
Saprobes – get nutrients from nonliving matter [valuable decomposers]
Parasites – thrive on tissues in living host
Digest the surrounding food and absorbs nutrients
Life cycle
Reproduce both asexually and sexually, producing spores
Food-absorbing part is a mesh of branching filaments [the mycelium, composed of individual hyphae]
Classification of fungi
Zygomycetes (Fig. 19.15)
Mostly bread molds
Reproduces asexually by small, dustlike spores and sexually by zygospores
Sac fungi (Fig. 19.16)
Edible morels and truffles, plus Penicillium [antibiotics] and Aspergillus [soy sauce]; also single-celled yeasts [bread and alcohol]
Larger ones produce structures with sacs that contain ascospores; yeasts mostly reproduce asexually, by budding
a. Club fungi (Fig. 19.14)
Mushrooms, puffballs, shelf fungi, rusts, and smuts
Produce basidiospores on club-shaped cells; mushroom has stalk and cap [with gills]
Imperfect fungi
All fungi lacking a sexual stage – gets moved out when sexual spore are discovered
Symbiotic associations between fungi and plants
Lichens
Mutualistic associations between fungi and cyanobacteria or green algae
Algae is protected from drying out
Fungi feeds on sugars produced by the algae
Can live in inhospitable places such as bare rock and tree trunks, but are sensitive to air pollution
Three body forms – crustose, foliose, and fruticose
Mycorrhizae
Symbiotic relationship in which fungi hyphae surround roots of woody plants
Increases the absorption power of the roots [water and minerals]
Animals: The Invertebrates (chap 20)
General characteristics of animals
Multicellular, heterotrophic, aerobic, reproduce sexually, motile at some point in life cycle
Life cycles include period of embryonic development; germ tissue layers (ectoderm, mesoderm, endoderm) give rise to adult organs
Diversity in body plans
Vertebrates – have a backbone; invertebrates – lack a backbone
Body symmetry
Radial – round
Bilateral – left and right sides
Show anterior (head), posterior (tail), dorsal (back), and ventral (belly) orientations
Cephalization – have a definite head, usually with feeding and sensory features
Type of gut
Saclike – with one opening
Complete – with two openings (mouth and anus)
Body cavities
Coelom – space between gut and body wall; lined with peritoneum
False coelom – such as in roundworms; not lined with peritoneum
No coelom – packed solidly with tissue
Segmentation – composed of repeating body units (may be grouped and modified for specialized tasks)
Possible origin of animals
Compartmentalization of a ciliate (like Paramecium )
Arose from colonial organisms like Volvox
Sponges – 8,000 species
Asymmetric body with no true tissues or organs
Flattened cells cover the exterior
Collar cells line interior chambers – move water and trap suspended food particles
Jelly-like matrix (with wandering amoeboid-like cells) between the two
Reproduce asexually (by fragmentation) and sexually (zygote produces swimming ciliated larva)
Cnidarians – tissues emerge (11,000 species)
Tentacled, radial animals
Inclu. jellyfishes, sea anemones, hydrozoans
Name comes from ability to sting by discharging nematocysts
Body plans
Medusa – jellyfish body plan; polyp – tubelike and usually attached (may be solitary or part of a colony)
Digestive cavity is saclike (only a mouth)
Outer epidermis, inner gastrodermis, and jellylike mesoglea between
Nerve net that coordinates sensory and motor activities
Reproduce asexually, and sexually (with swimming ciliated larva)
Flatworms, roundworms, rotifers – simple organ systems
Flatworms – 15,000 species (Table 28.2)
Body plan
Saclike gut (but none in tapeworms)
Bilateral symmetry, and cephalization
No coelom
Hermaphroditic – both sexes in one body
Planarians
Free-living, freshwater aquatic
Possess a pharynx tube that extends to feed
Flame cells – regulate body fluid volume
Asexual reproduction by fission of body
Flukes
Internal parasites of liver, lung, or blood
Complex life cycle – requires primary host for sexual reproduction, and intermediate host (usually a snail)
Tapeworms (Fig. 20.14)
Internal parasites of vertebrate intestines
Body – scolex (head) and proglottids (segments)
Roundworms – 20,000 species
Most small and free-living, but some parasitic (ex., hookworms in human organs)
Bilateral symmetry, complete digestive tract in a pseudocoelom, cuticle covers and protects body
Rotifers – 1,800 species
Microscopic, multicelled aquatic animals
Complex, with a complete set of organs
E. Mollusks – 110,000 species
1. Body usually incl. head, foot, shell, mantle, gills, and radula
2. Gastropods – snails and slugs
a. "Belly footed" animals – most organs in spiraled shell
b. Torsion – 180 turn that places some organs toward the head
3. Bivalves – clams, scallops, oysters, mussels, etc.
a. Two shells for protection – formed by mantle
b. No head; foot usually specialized for burrowing
c. Water and suspended food particles are drawn in by action of cilia on the gills
4. Cephalopods – squids, cuttlefishes, octopuses, nautiluses
a. Largest of the invertebrates
b. Body modified for active predatory life-style – tentacles, beaklike jaws, and jet propulsion
c. Closed circulatory system , nervous system well developed, eyes form images, learning and memory possible
F. Annelids – segmented worms (15,000 species)
1. Annelid groups
a. Earthworms – valuable tillers of soil
b. Leeches – aquatic "blood suckers"
c. Polychaetes – marine worms (with tentacles and "gills")
2. Annelid adaptations [of the earthworm]
a. Extensive segmentation with internal partitions; fluid filled to provide a hydrostatic skeleton
b. Paired nephridia in every segment for body fluid and liquid waste control
c. Digestive system is complete; circulation is closed
G. Arthropods – 1,000,000+ species
1. Adaptations
a. Hardened exoskeleton of protein and chitin (plus calcium in some)
1) Flexible and lightweight; good barrier to water loss
2) But, must be shed periodically
b. Fewer body segments – grouped to form head, thorax, and abdomen
c. Jointed appendages for feeding, sensing, and locomotion
d. "Breathe" using tubes (trachaeas) connected to holes (spiracles) in the abdomen segments
e. Multi-faceted compound eyes allow for wide angle of vision and motion perception, but not focusing
f. Metamorphosis – larval stages concentrate on feeding and growth; adults specialize in dispersal and reproduction
1) Complete metamorphosis – egg, larva, pupa, adult
2) Incomplete metamorphosis – egg, "nymph," adult
3) Gradual metamorphosis – egg –> adult
Spiders and relatives – 25,000 species
Chelicerates – spiders, scorpions, ticks, mites, horseshoe crabs, sea spiders (Fig. 20.22)
Spiders – eight-legged predators that trap insects in webs
Mites – some free-living, others are pests of plants and animals
Ticks – blood-suckers and disease carriers
Body plan – chelicerae (piercing), pedipalps (grasping), open circulatory system, book lungs
Crustaceans – 35,000 species
Shrimps [and krills], lobsters, crayfishes, crabs, barnacles, copepods, pillbugs
Important components of food webs and some serve as human food
Body plan
Cephalothorax and abdomen; gills in aquatic ones; ventral nerve cord (Fig. 20.24)
Appendages – two pairs of antennae, a pair of mandibles and maxillae, and five pairs of legs
Millipedes and centipedes – long, segmented body with many legs
Millipedes
Cylindrical body with two pairs of legs on each body segment
Slow-moving vegetarians
Centipedes
Flattened body with one pair of legs on each body segment
Fast-moving carnivore of small invertebrates
Insects – 900,000+ species
Body plan
Three regions – head (sensory and feeding), thorax (locomotion – six legs, two pairs of wings), and abdomen
Malpighian tubules process metabolic waste and aid in water retention
Most successful of all groups
Echinoderms – spiny skinned animals (6,000 species) (Fig. 20.30)
Sea stars (starfish), brittle stars, sea urchins, sand dollars, sea cucumbers, sea lilies, feather stars
Larvae are bilateral; adults are radially symmetrical
Five-rayed body plan; spines and skin gills
Tube feet (locomotion and capture of prey) connected to water vascular system
Sea stars feed on bivalve mollusks – can evert stomach for exterior digestion
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